243 lines
26 KiB
XML
243 lines
26 KiB
XML
<document id="5BD402613471D194876903E4B5403F0A" ID-DOI="10.5281/zenodo.157036" ID-GBIF-Dataset="8845a192-a81c-4e81-a280-e19c508ce7dd" ID-ISSN="11755326" ID-Zenodo-Dep="157036" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459633983606" checkinUser="plazi" docAuthor="Shear, William A." docDate="2003" docId="F7710943FF9D3D1498781024FEA1FBC5" docLanguage="en" docName="zt00321.pdf" docOrigin="Zootaxa 321" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Trigenotyla Causey" docType="treatment" docVersion="10" lastPageNumber="19" masterDocId="0B48713BFF8C3D0699701225FFF0FFBB" masterDocTitle="The milliped family Trichopetalidae, Part 1: Introduction and Genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="18" updateTime="1698204205547" updateUser="plazi">
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<mods:title id="44589F408549618AE57B7CA55F802B0A">The milliped family Trichopetalidae, Part 1: Introduction and Genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea)</mods:title>
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<mods:namePart id="BB9B01CB4890C3C1240AA4E91F11F17A">Shear, William A.</mods:namePart>
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<mods:title id="5D547DB4EC2BB0726EB1A870856EE952">Zootaxa</mods:title>
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<mods:part id="E09A65A8B7DF770C58CA4F38EFBC22A7">
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<mods:date id="C716DADDDB78DDF1A819CB60CBA92E2E">2003</mods:date>
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<mods:number id="D6B1992A9BDC0507FB7F0A4A2BDD3BD7">321</mods:number>
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<mods:identifier id="F82D999DCD933B411F1546960C0B3889" type="DOI">10.5281/zenodo.157036</mods:identifier>
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<treatment id="F7710943FF9D3D1498781024FEA1FBC5" ID-DOI="http://doi.org/10.5281/zenodo.6273582" ID-GBIF-Taxon="119332271" ID-Zenodo-Dep="6273582" LSID="urn:lsid:plazi:treatment:F7710943FF9D3D1498781024FEA1FBC5" httpUri="http://treatment.plazi.org/id/F7710943FF9D3D1498781024FEA1FBC5" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
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<subSubSection id="37C2EBDEFF9D3D1798781024FDBDFDA0" box="[264,589,513,539]" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="7F67B855FF9D3D1798781024FDBDFDA0" blockId="17.[264,589,513,539]" box="[264,589,513,539]" pageId="17" pageNumber="18">
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<heading id="242F0F39FF9D3D1798781024FDBDFDA0" bold="true" box="[264,589,513,539]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
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<emphasis id="4DAC6447FF9D3D1798781024FDBDFDA0" bold="true" box="[264,589,513,539]" pageId="17" pageNumber="18">
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Genus
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<taxonomicName id="B8D8C3D6FF9D3D17982F1024FDBDFDA0" ID-CoL="7Q56B" authority="Causey" authorityName="Causey" box="[351,589,513,539]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="4DAC6447FF9D3D17982F1024FE1AFDA0" bold="true" box="[351,490,513,539]" italics="true" pageId="17" pageNumber="18">Trigenotyla</emphasis>
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Causey
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="37C2EBDEFF9D3D1798781067FB7FFDE3" box="[264,1167,577,601]" pageId="17" pageNumber="18" type="reference_group">
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<paragraph id="7F67B855FF9D3D1798781067FB7FFDE3" blockId="17.[264,1167,577,601]" box="[264,1167,577,601]" pageId="17" pageNumber="18">
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<treatmentCitationGroup id="5FC89F7BFF9D3D1798781067FB7FFDE3" box="[264,1167,577,601]" pageId="17" pageNumber="18">
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<taxonomicName id="B8D8C3D6FF9D3D1798781067FDB5FDE3" ID-CoL="7Q56B" authority="Causey 1951:118" authorityName="Causey" authorityPageNumber="118" authorityYear="1951" box="[264,581,577,601]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="4DAC6447FF9D3D1798781067FE73FDE2" box="[264,387,578,601]" italics="true" pageId="17" pageNumber="18">Trigenotyla</emphasis>
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Causey 1951:118
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</taxonomicName>
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; 1959:76.
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<treatmentCitation id="FE799E44FF9D3D179BCC1064FC9DFDE3" author="Shear" box="[700,877,577,600]" page="280" pageId="17" pageNumber="18" year="1972">
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<bibRefCitation id="1B49C5A4FF9D3D179BCC1064FCCEFDE3" author="Shear" box="[700,830,577,600]" pageId="17" pageNumber="36" refString="Shear, W. A. (1972) Studies in the milliped Order Chordeumida (Diplopoda): a revision of the Family Cleidogonidae and a reclassification of the Order Chordeumida in the New World. Bulletin of the Museum of Comparative Zoology, 144, 151 - 352." type="journal article" year="1972">Shear, 1972</bibRefCitation>
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:280
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</treatmentCitation>
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. Hoffman, 1999:238 (list).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="37C2EBDEFF9D3D179878105AFAEAFD58" pageId="17" pageNumber="18" type="materials_examined">
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<paragraph id="7F67B855FF9D3D179878105AFCD5FD22" blockId="17.[264,805,639,665]" box="[264,805,639,665]" pageId="17" pageNumber="18">
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<emphasis id="4DAC6447FF9D3D179878105AFE55FD23" box="[264,421,639,664]" italics="true" pageId="17" pageNumber="18">
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<typeStatus id="A06306F7FF9D3D179878105AFECFFD23" box="[264,319,639,664]" pageId="17" pageNumber="18">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="B8D8C3D6FF9D3D1798DC105AFCD1FD22" authority="Causey 1951" authorityName="Causey" authorityYear="1951" box="[428,801,639,665]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="parca">
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<emphasis id="4DAC6447FF9D3D1798DC105AFD8EFD23" box="[428,638,639,664]" italics="true" pageId="17" pageNumber="18">Trigenotyla parca</emphasis>
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Causey 1951
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="7F67B855FF9D3D17987810EFFAEAFD58" blockId="17.[264,1324,713,1899]" box="[264,1306,713,739]" pageId="17" pageNumber="18">
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<emphasis id="4DAC6447FF9D3D17987810EFFE24FD58" box="[264,468,714,739]" italics="true" pageId="17" pageNumber="18">Included species:</emphasis>
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The
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<typeStatus id="A06306F7FF9D3D179B6110ECFDB6FD58" box="[529,582,713,739]" pageId="17" pageNumber="18">type</typeStatus>
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,
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<taxonomicName id="B8D8C3D6FF9D3D179B2210EFFCB9FD58" authority="Causey 1959" authorityName="Causey" authorityYear="1959" box="[594,841,713,739]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vaga">
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<emphasis id="4DAC6447FF9D3D179B2210EFFD55FD58" box="[594,677,714,739]" italics="true" pageId="17" pageNumber="18">T. vaga</emphasis>
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<bibRefCitation id="1B49C5A4FF9D3D179BDF10ECFCB9FD58" author="Causey" box="[687,841,713,739]" pageId="17" pageNumber="35" refString="Causey, N. B. (1959) A second species of the milliped genus Trigenotyla (Chordeumidea: Conotylidae: Trichopetalinae: Scoterpini). Proceedings of the Biological Society of Washington, 72, 75 - 78." type="journal article" year="1959">Causey 1959</bibRefCitation>
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</taxonomicName>
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,
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<emphasis id="4DAC6447FF9D3D179A2510EFFC32FD58" box="[853,962,714,739]" italics="true" pageId="17" pageNumber="18">
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<taxonomicName id="B8D8C3D6FF9D3D179A2510EFFC4DFD58" box="[853,957,714,739]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="blacki" status="sp. nov.">T. blacki</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="569FD93CFF9D3D179AB910ECFBF9FD58" box="[969,1033,713,739]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
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, and
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<emphasis id="4DAC6447FF9D3D179D3710EFFB23FD58" box="[1095,1235,714,739]" italics="true" pageId="17" pageNumber="18">
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<taxonomicName id="B8D8C3D6FF9D3D179D3710EFFB3FFD58" box="[1095,1231,714,739]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="seminole" status="sp. nov.">T. seminole</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="569FD93CFF9D3D179DAB10ECFAEAFD58" box="[1243,1306,713,739]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="37C2EBDEFF9D3D17984810D7FCC3FC38" pageId="17" pageNumber="18" type="diagnosis">
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<paragraph id="7F67B855FF9D3D17984810D7FCC3FC38" blockId="17.[264,1324,713,1899]" pageId="17" pageNumber="18">
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<emphasis id="4DAC6447FF9D3D17984810D7FE49FCB0" box="[312,441,754,779]" italics="true" pageId="17" pageNumber="18">Diagnosis:</emphasis>
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Closely related to
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<emphasis id="4DAC6447FF9D3D179BD310D7FCEEFCB0" box="[675,798,754,779]" italics="true" pageId="17" pageNumber="18">Causeylla,</emphasis>
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but differing in the ninth legs of males, in which the coxae extend distal to the articulation with the telopodite. The anterior surfaces of the gonopod coxae lack trichomes. All species with 30 trunk segments; Causey (1951) originally stated that
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<taxonomicName id="B8D8C3D6FF9D3D179B70114CFD91FC39" box="[512,609,873,898]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="parca">
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<emphasis id="4DAC6447FF9D3D179B70114CFD91FC39" box="[512,609,873,898]" italics="true" pageId="17" pageNumber="18">T. parca</emphasis>
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</taxonomicName>
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had 28 segments.
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</paragraph>
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</subSubSection>
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<subSubSection id="37C2EBDEFF9D3D14984811B4FEA1FBC5" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="distribution">
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<paragraph id="7F67B855FF9D3D17984811B4FB2DFC10" blockId="17.[264,1324,713,1899]" box="[312,1245,913,939]" pageId="17" pageNumber="18">
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<emphasis id="4DAC6447FF9D3D17984811B4FE21FC11" box="[312,465,913,938]" italics="true" pageId="17" pageNumber="18">Distribution:</emphasis>
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Northwestern Arkansas, northeastern and southcentral Oklahoma.
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</paragraph>
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<paragraph id="7F67B855FF9D3D179848119CFD15FB78" blockId="17.[264,1324,713,1899]" pageId="17" pageNumber="18">
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<emphasis id="4DAC6447FF9D3D179848119CFDE7FC69" box="[312,535,953,978]" italics="true" pageId="17" pageNumber="18">
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Notes:
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<taxonomicName id="B8D8C3D6FF9D3D1798E0119CFDE7FC69" box="[400,535,953,978]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Trigenotyla</taxonomicName>
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</emphasis>
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species are often, but not always, collected in caves; for many areas, caves are more heavily collected than forest litter habitats, and this may contribute to the preponderance of cave records for this genus. Only
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<taxonomicName id="B8D8C3D6FF9D3D179AA8162CFBCDFB99" box="[984,1085,1033,1058]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="blacki">
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<emphasis id="4DAC6447FF9D3D179AA8162CFBCDFB99" box="[984,1085,1033,1058]" italics="true" pageId="17" pageNumber="18">T. blacki</emphasis>
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</taxonomicName>
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seems restricted to caves, and only this species shows any troglomorphic adapatations (reduced eyes). It is also very likely that surface populations of Oklahoma
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<taxonomicName id="B8D8C3D6FF9D3D179AE5167CFBEBFBC9" box="[917,1051,1113,1138]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="4DAC6447FF9D3D179AE5167CFBEBFBC9" box="[917,1051,1113,1138]" italics="true" pageId="17" pageNumber="18">Trigenotyla</emphasis>
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</taxonomicName>
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species are active and available for collection in the winter rather than the hot, dry summers that prevail in the region; little collecting is done in winter.
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</paragraph>
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<paragraph id="7F67B855FF9D3D17984816F4FDFEFA30" blockId="17.[264,1324,713,1899]" pageId="17" pageNumber="18">
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The distribution of
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<taxonomicName id="B8D8C3D6FF9D3D179B5616F4FD5DFB51" box="[550,685,1233,1258]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="4DAC6447FF9D3D179B5616F4FD5DFB51" box="[550,685,1233,1258]" italics="true" pageId="17" pageNumber="18">Trigenotyla</emphasis>
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</taxonomicName>
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as now understood can be divided into two distinct regions.
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<taxonomicName id="B8D8C3D6FF9D3D17980516DCFDB6FAA9" box="[373,582,1273,1298]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="parca">
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<emphasis id="4DAC6447FF9D3D17980516DCFDB6FAA9" box="[373,582,1273,1298]" italics="true" pageId="17" pageNumber="18">Trigenotyla parca</emphasis>
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</taxonomicName>
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is known from caves and epigean sites in northwestern Arkansas, while
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<taxonomicName id="B8D8C3D6FF9D3D1798F61704FE1DFA81" box="[390,493,1313,1338]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="blacki">
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<emphasis id="4DAC6447FF9D3D1798F61704FE1DFA81" box="[390,493,1313,1338]" italics="true" pageId="17" pageNumber="18">T. blacki</emphasis>
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</taxonomicName>
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has been found in caves across the border in northeastern Oklahoma. The other region is in southcentral Oklahoma, in
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<taxonomicName id="B8D8C3D6FF9D3D179A13176CFC25FAD8" box="[867,981,1353,1379]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="34" phylum="Arthropoda" rank="species" species="seminole">Seminole</taxonomicName>
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, Johnston, Murray, LeFlore, and Latimer Counties.
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</paragraph>
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<paragraph id="7F67B855FF9D3D17984817BCFE74F8A1" blockId="17.[264,1324,713,1899]" pageId="17" pageNumber="18">
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Latimer County is the
|
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<typeStatus id="A06306F7FF9D3D179B3617BCFD89FA08" box="[582,633,1433,1459]" pageId="17" pageNumber="18">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="B8D8C3D6FF9D3D179A7417BCFCCDFA09" box="[772,829,1433,1458]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vaga">
|
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<emphasis id="4DAC6447FF9D3D179A7417BCFCCDFA09" box="[772,829,1433,1458]" italics="true" pageId="17" pageNumber="18">vaga</emphasis>
|
||
</taxonomicName>
|
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and, with LeFlore County, is a good distance east of the other records. Causey labelled the more western collections as “profunda,” suggesting that she believed them to be a new species. Unfortunately, the
|
||
<typeStatus id="A06306F7FF9D3D179DA717CCFAFAF9B8" box="[1239,1290,1513,1539]" pageId="17" pageNumber="18">type</typeStatus>
|
||
of
|
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<taxonomicName id="B8D8C3D6FF9D3D1798781434FEADF991" box="[264,349,1553,1578]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vaga">
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<emphasis id="4DAC6447FF9D3D1798781434FEADF991" box="[264,349,1553,1578]" italics="true" pageId="17" pageNumber="18">T. vaga</emphasis>
|
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</taxonomicName>
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is the only known male specimen, and the gonopods, originally mounted on a slide (AMNH), have been lost (L. Sorkin, pers. comm. 2002). I found that
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<taxonomicName id="B8D8C3D6FF9D3D179D4C141CFB33F9E9" box="[1084,1219,1593,1618]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="4DAC6447FF9D3D179D4C141CFB33F9E9" box="[1084,1219,1593,1618]" italics="true" pageId="17" pageNumber="18">Trigenotyla</emphasis>
|
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</taxonomicName>
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south of the Canadian River in Murray and Johnston Counties closely conformed to the illustrations of
|
||
<taxonomicName id="B8D8C3D6FF9D3D17981D14ACFE56F919" box="[365,422,1673,1698]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vaga">
|
||
<emphasis id="4DAC6447FF9D3D17981D14ACFE56F919" box="[365,422,1673,1698]" italics="true" pageId="17" pageNumber="18">vaga</emphasis>
|
||
</taxonomicName>
|
||
provided by
|
||
<bibRefCitation id="1B49C5A4FF9D3D179B3814ACFD08F918" author="Causey" box="[584,760,1673,1699]" pageId="17" pageNumber="35" refString="Causey, N. B. (1959) A second species of the milliped genus Trigenotyla (Chordeumidea: Conotylidae: Trichopetalinae: Scoterpini). Proceedings of the Biological Society of Washington, 72, 75 - 78." type="journal article" year="1959">Causey (1959)</bibRefCitation>
|
||
, while there were differences in the specimens from
|
||
<taxonomicName id="B8D8C3D6FF9D3D17983D1494FE4CF970" box="[333,444,1713,1739]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="34" phylum="Arthropoda" rank="species" species="seminole">Seminole</taxonomicName>
|
||
County north of the Canadian. Therefore, I am naming the
|
||
<taxonomicName id="B8D8C3D6FF9D3D179DCC1494FADBF970" box="[1212,1323,1713,1739]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="34" phylum="Arthropoda" rank="species" species="seminole">Seminole</taxonomicName>
|
||
county specimens as a new species,
|
||
<emphasis id="4DAC6447FF9D3D179BDF14FCFCC9F949" box="[687,825,1753,1778]" italics="true" pageId="17" pageNumber="18">
|
||
<taxonomicName id="B8D8C3D6FF9D3D179BDF14FCFCC2F949" box="[687,818,1753,1778]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="seminole">T. seminole</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and referring specimens south of the river to
|
||
<emphasis id="4DAC6447FF9D3D1798561524FE74F8A1" box="[294,388,1793,1818]" italics="true" pageId="17" pageNumber="18">
|
||
<taxonomicName id="B8D8C3D6FF9D3D1798561524FE70F8A1" box="[294,384,1793,1818]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="vaga">T. vaga</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="7F67B855FF9D3D149848150CFC04FEE5" blockId="17.[264,1324,713,1899]" lastBlockId="18.[264,1323,284,1150]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
|
||
I predict that
|
||
<taxonomicName id="B8D8C3D6FF9D3D1798AC150CFD92F8F9" box="[476,610,1833,1858]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4DAC6447FF9D3D1798AC150CFD92F8F9" box="[476,610,1833,1858]" italics="true" pageId="17" pageNumber="18">Trigenotyla</emphasis>
|
||
</taxonomicName>
|
||
species will be found generally in suitable habitats (caves, riverine forests) throughout eastern Oklahoma and western Arkansas when collectors are in the right place at the right time. It also seems likely that the genus will turn up in southwestern Missouri, southeastern Kansas, and northeastern Texas.
|
||
</paragraph>
|
||
<paragraph id="7F67B855FF9E3D1498481349FD4AFD55" blockId="18.[264,1323,284,1150]" pageId="18" pageNumber="19">
|
||
<taxonomicName id="B8D8C3D6FF9E3D1498481349FE4FFE3E" box="[312,447,364,389]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4DAC6447FF9E3D1498481349FE4FFE3E" box="[312,447,364,389]" italics="true" pageId="18" pageNumber="19">Trigenotyla</emphasis>
|
||
</taxonomicName>
|
||
gonopods are similar in general appearance to those of
|
||
<taxonomicName id="B8D8C3D6FF9E3D149D361349FB38FE3E" box="[1094,1224,364,389]" class="Diplopoda" family="Trichopetalidae" genus="Causeyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4DAC6447FF9E3D149D361349FB38FE3E" box="[1094,1224,364,389]" italics="true" pageId="18" pageNumber="19">Causeyella</emphasis>
|
||
</taxonomicName>
|
||
species, but in some respects seem more apomorphic. The gonopod setae tend to be ensiform, distinctly so in
|
||
<emphasis id="4DAC6447FF9E3D1498ED1399FDF3FE6E" box="[413,515,444,469]" italics="true" pageId="18" pageNumber="19">
|
||
<taxonomicName id="B8D8C3D6FF9E3D1498ED1399FE0CFE6E" box="[413,508,444,469]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="parca">T. parca</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and are divided into two groups, with the typical 3 basal setae. The mesal coxite is broad in two species (
|
||
<taxonomicName id="B8D8C3D6FF9E3D149BB513C1FCBAFE46" box="[709,842,484,509]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="seminole">
|
||
<emphasis id="4DAC6447FF9E3D149BB513C1FCBAFE46" box="[709,842,484,509]" italics="true" pageId="18" pageNumber="19">T. seminole</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="B8D8C3D6FF9E3D149AF513C1FC2AFE46" box="[901,986,484,509]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="vaga">
|
||
<emphasis id="4DAC6447FF9E3D149AF513C1FC2AFE46" box="[901,986,484,509]" italics="true" pageId="18" pageNumber="19">T. vaga</emphasis>
|
||
</taxonomicName>
|
||
), thin and acute in
|
||
<emphasis id="4DAC6447FF9E3D149DB213C1FAD8FE46" box="[1218,1320,484,509]" italics="true" pageId="18" pageNumber="19">
|
||
<taxonomicName id="B8D8C3D6FF9E3D149DB213C1FAD1FE46" box="[1218,1313,484,509]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="parca">T. parca</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and absent in
|
||
<emphasis id="4DAC6447FF9E3D1498D91029FDE4FD9E" box="[425,532,524,549]" italics="true" pageId="18" pageNumber="19">
|
||
<taxonomicName id="B8D8C3D6FF9E3D1498D91029FDE1FD9E" box="[425,529,524,549]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="blacki">T. blacki</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The ectal coxite in the former two species is very much like that of species of
|
||
<taxonomicName id="B8D8C3D6FF9E3D1498F71011FDF9FDF6" box="[391,521,564,589]" class="Diplopoda" family="Trichopetalidae" genus="Causeyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4DAC6447FF9E3D1498F71011FDF9FDF6" box="[391,521,564,589]" italics="true" pageId="18" pageNumber="19">Causeyella</emphasis>
|
||
</taxonomicName>
|
||
and suggests that the latter two are apomorphic in the reduction of the structure. Trichomes are absent from the surfaces of the gonopod coxae, though under high magnification the coxae can be seen to be minutely roughened with tiny, conical projections. It is not clear at present if this is apomorphic (loss of trichomes) or plesiomorphic (partially developed trichomes).
|
||
</paragraph>
|
||
<paragraph id="7F67B855FF9E3D14983010D9FEA1FBC5" blockId="18.[264,1323,284,1150]" pageId="18" pageNumber="19">
|
||
The ninth legs of males are unique among trichopetalids in that the Lshaped coxa is extended distad to the articulation of the telopodite, a modification hinted at in some
|
||
<taxonomicName id="B8D8C3D6FF9E3D149D801101FEA9FCDE" class="Diplopoda" family="Trichopetalidae" genus="Causeyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4DAC6447FF9E3D149D801101FEA9FCDE" italics="true" pageId="18" pageNumber="19">Causeyella</emphasis>
|
||
</taxonomicName>
|
||
species. Causey (1951) called the basal podomere a coxoprefemur, but if it is homologous to the same podomeres in other trichopetalids, it is the coxa alone, and the large distal joint includes the prefemur and other distal podomeres. In one species (
|
||
<taxonomicName id="B8D8C3D6FF9E3D149DA711B9FEB2FC66" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="seminole">
|
||
<emphasis id="4DAC6447FF9E3D149DA711B9FEB2FC66" italics="true" pageId="18" pageNumber="19">T. seminole</emphasis>
|
||
</taxonomicName>
|
||
) there are tiny vestigial podomeres at the end of the prefemur. The form of the prefemur and the presence of these vestigial podomeres suggests that the telopodite article is only the prefemur, and unlike the situation in other genera, does not result from the fusion of other podomeres. The ninth coxa also has an unusually distinct, but likely vestigial, gland.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |