322 lines
35 KiB
XML
322 lines
35 KiB
XML
<document id="C56C8EF2C7589EA882A4294FE4E350DA" ID-DOI="10.5281/zenodo.157036" ID-GBIF-Dataset="8845a192-a81c-4e81-a280-e19c508ce7dd" ID-ISSN="11755326" ID-Zenodo-Dep="157036" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459633983606" checkinUser="plazi" docAuthor="Shear, William A." docDate="2003" docId="F7710943FF993D11987817D1FD35FBC5" docLanguage="en" docName="zt00321.pdf" docOrigin="Zootaxa 321" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Trigenotyla blacki Shear, 2003, n. sp." docType="treatment" docVersion="10" lastPageNumber="24" masterDocId="0B48713BFF8C3D0699701225FFF0FFBB" masterDocTitle="The milliped family Trichopetalidae, Part 1: Introduction and Genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="22" updateTime="1698204205547" updateUser="plazi">
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<mods:mods id="D620CF2DA1B499EECB8585A1601C6089" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="50BDBC321164CF1DE2E2FE43E62406EB">
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<mods:title id="9FAEDEB63E9695D8F90AEF810A5E26EE">The milliped family Trichopetalidae, Part 1: Introduction and Genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea)</mods:title>
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<mods:name id="4526A2714C17A591C72B61FB9CC165B4" type="personal">
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<mods:roleTerm id="3AB59C846818DAEC2DED1D96671BEBD2">Author</mods:roleTerm>
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<mods:namePart id="5497B9388E9A39DB237C015803ABF00E">Shear, William A.</mods:namePart>
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<mods:typeOfResource id="073885B492974D623B2A16CAE0B24CBB">text</mods:typeOfResource>
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<mods:title id="F5C45C996AFCD0C42EED4CED9AAEF8C5">Zootaxa</mods:title>
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<mods:part id="7A1C5A2116F5F159B24758CCF4AB05FE">
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<mods:date id="110D471D5618536E62BF1FE8F611EBA8">2003</mods:date>
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<mods:number id="C3A6C0B9B9509D0E8C941188F7F19340">321</mods:number>
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<mods:classification id="C1BA00726CBB59744B060BCEDFB9878A">journal article</mods:classification>
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<mods:identifier id="6CFFD19BA03C95EA444F270F60B311EA" type="DOI">10.5281/zenodo.157036</mods:identifier>
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<treatment id="F7710943FF993D11987817D1FD35FBC5" ID-DOI="http://doi.org/10.5281/zenodo.6273588" ID-GBIF-Taxon="119332275" ID-Zenodo-Dep="6273588" LSID="urn:lsid:plazi:treatment:F7710943FF993D11987817D1FD35FBC5" httpUri="http://treatment.plazi.org/id/F7710943FF993D11987817D1FD35FBC5" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
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<subSubSection id="37C2EBDEFF993D13987817D1FE61F98A" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="7F67B855FF993D13987817D1FDC5F9B5" blockId="21.[264,565,1524,1585]" box="[264,565,1524,1550]" pageId="21" pageNumber="22">
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<heading id="242F0F39FF993D13987817D1FDC5F9B5" bold="true" box="[264,565,1524,1550]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
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<emphasis id="4DAC6447FF993D13987817D1FDC5F9B5" bold="true" box="[264,565,1524,1550]" pageId="21" pageNumber="22">
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<emphasis id="4DAC6447FF993D13987817D1FE1AF9B5" bold="true" box="[264,490,1524,1550]" italics="true" pageId="21" pageNumber="22">
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<taxonomicName id="B8D8C3D6FF993D13987817D1FE14F9B5" box="[264,484,1524,1550]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="blacki" status="sp. nov.">Trigenotyla blacki</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="569FD93CFF993D13988217D1FDC5F9B5" box="[498,565,1524,1550]" pageId="21" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7F67B855FF993D1398781432FE61F98A" blockId="21.[264,565,1524,1585]" box="[264,401,1559,1585]" pageId="21" pageNumber="22">
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<figureCitation id="E7E3A4D0FF993D1398781432FE61F98A" box="[264,401,1559,1585]" captionStart="FIGURES 12 – 15" captionStartId="20.[264,382,1559,1583]" captionTargetBox="[336,1254,680,1540]" captionTargetId="figure@20.[331,1254,675,1548]" captionTargetPageId="20" captionText="FIGURES 12 – 15. Trigenotyla parca Causey, 12, gonopods, anterior view. Trigenotyla blacki n. sp., 13 – 15. 13, gonopods, anterior view. 14, male ninth legs, anterior view. 15, male right sixth leg, posterior view. Scale lines for Figs. 12 – 14 = 0.1 mm (100 µm), for Fig. 15, 0.5 mm (500 µm)." httpUri="https://zenodo.org/record/157040/files/figure.png" pageId="21" pageNumber="22">Figs. 13–15</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="37C2EBDEFF993D1398781442FDA0F8CA" pageId="21" pageNumber="22" type="description">
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<paragraph id="7F67B855FF993D1398781442FDA0F8CA" blockId="21.[264,1323,1639,1905]" pageId="21" pageNumber="22">
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<emphasis id="4DAC6447FF993D1398781442FEA2F93B" box="[264,338,1639,1664]" italics="true" pageId="21" pageNumber="22">
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<typeStatus id="A06306F7FF993D1398781442FEB8F93B" box="[264,328,1639,1664]" pageId="21" pageNumber="22">Types</typeStatus>
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:
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</emphasis>
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Male
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<typeStatus id="A06306F7FF993D1398D11442FDFBF93A" box="[417,523,1639,1665]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
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, one male and two female
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<typeStatus id="A06306F7FF993D139A3B1442FC4BF93A" box="[843,955,1639,1665]" pageId="21" pageNumber="22" type="paratype">paratypes</typeStatus>
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from StansburyJanuary Cave,
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<quantity id="B82015B0FF993D13987814AAFECFF912" box="[264,319,1679,1705]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="21" pageNumber="22" unit="mi" value="4.0">4 mi</quantity>
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. N of Colcord,
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<emphasis id="4DAC6447FF993D13988E14AAFD52F913" box="[510,674,1679,1704]" italics="true" pageId="21" pageNumber="22">Delaware Co.</emphasis>
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, Oklahoma, collected
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<date id="0B669E95FF993D139AC014AAFB86F912" box="[944,1142,1679,1705]" pageId="21" pageNumber="22" value="1971-01-13">13 January 1971</date>
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by J. H. Black (FSCA). The following specimens are
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<typeStatus id="A06306F7FF993D139B861492FC9BF96A" box="[758,875,1719,1745]" pageId="21" pageNumber="22" type="paratype">paratypes</typeStatus>
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: OKLAHOMA:
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<emphasis id="4DAC6447FF993D139D4F1492FB13F96B" box="[1087,1251,1719,1744]" italics="true" pageId="21" pageNumber="22">Delaware Co</emphasis>
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.: Jail House Cave,
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<quantity id="B82015B0FF993D1398D614FAFE2FF942" box="[422,479,1759,1785]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="21" pageNumber="22" unit="mi" value="3.0">3 mi</quantity>
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. N of Choleta,
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<date id="0B669E95FF993D139BE214FAFCB8F942" box="[658,840,1759,1785]" pageId="21" pageNumber="22" value="1971-01-01">1 January 1971</date>
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, J. H. Black, juvenile; Bell’s Bluff Cave,
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<quantity id="B82015B0FF993D1398781522FEB3F89A" box="[264,323,1799,1825]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="21" pageNumber="22" unit="mi" value="5.0">5 mi</quantity>
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. S of Jay,
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<date id="0B669E95FF993D1398B11522FDAFF89A" box="[449,607,1799,1825]" pageId="21" pageNumber="22" value="1971-07-14">14 July 1971</date>
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, J. H. Black, male; Twin Cave,
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<quantity id="B82015B0FF993D139A921522FBEDF89A" box="[994,1053,1799,1825]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="21" pageNumber="22" unit="mi" value="5.0">5 mi</quantity>
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. W,
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<quantity id="B82015B0FF993D139D231522FB7EF89A" box="[1107,1166,1799,1825]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="21" pageNumber="22" unit="mi" value="3.0">3 mi</quantity>
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. S of Jay,
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<date id="0B669E95FF993D139C7D1522FE3DF8F2" pageId="21" pageNumber="22" value="1970-11-29">29 November 1970</date>
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, J. H. Black, female.
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<emphasis id="4DAC6447FF993D139B9C150AFC94F8F3" box="[748,868,1839,1864]" italics="true" pageId="21" pageNumber="22">Adair Co.</emphasis>
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: Cave #AD14,
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<date id="0B669E95FF993D139D44150AFBA0F8F2" box="[1076,1104,1839,1865]" pageId="21" pageNumber="22" value="2003-01-11">11</date>
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January 2003, G. Graening, D. Fenolio, male.
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</paragraph>
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</subSubSection>
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<subSubSection id="37C2EBDEFF9A3D1098481339FB36FEE5" pageId="22" pageNumber="23" type="diagnosis">
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<paragraph id="7F67B855FF9A3D1098481339FB36FEE5" blockId="22.[264,1325,284,870]" pageId="22" pageNumber="23">
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<emphasis id="4DAC6447FF9A3D1098481339FE49FE8E" box="[312,441,284,309]" italics="true" pageId="22" pageNumber="23">Diagnosis:</emphasis>
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The gonopod has ensiform setae like
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<emphasis id="4DAC6447FF9A3D109AFA1339FC04FE8E" box="[906,1012,284,309]" italics="true" pageId="22" pageNumber="23">
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<taxonomicName id="B8D8C3D6FF9A3D109AFA1339FC1FFE8E" box="[906,1007,284,309]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="parca">T. parca</taxonomicName>
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,
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</emphasis>
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but lacks an ectal coxite.
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<taxonomicName id="B8D8C3D6FF9A3D1098781361FE2FFEE6" box="[264,479,324,349]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="blacki">
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<emphasis id="4DAC6447FF9A3D1098781361FE2FFEE6" box="[264,479,324,349]" italics="true" pageId="22" pageNumber="23">Trigenotyla blacki</emphasis>
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</taxonomicName>
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has fewer ocelli, which are depigmented and in only two rows.
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</paragraph>
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</subSubSection>
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<subSubSection id="37C2EBDEFF9A3D1098481349FD1BFE15" pageId="22" pageNumber="23" type="etymology">
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<paragraph id="7F67B855FF9A3D1098481349FD1BFE15" blockId="22.[264,1325,284,870]" pageId="22" pageNumber="23">
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<emphasis id="4DAC6447FF9A3D1098481349FE4FFE3E" box="[312,447,364,389]" italics="true" pageId="22" pageNumber="23">Etymology:</emphasis>
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for the collector, Jeffrey H. Black, recognizing his contributions (
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<bibRefCitation id="1B49C5A4FF9A3D109DAD1349FEB4FE15" author="Black" pageId="22" pageNumber="35" refString="Black, J. H. (1971) The cave life of Oklahoma, a preliminary study (excluding Chiroptera). Oklahoma Underground, 4, 1 - 49." type="journal article" year="1971">Black, 1971</bibRefCitation>
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) to the biospeleology of Oklahoma.
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</paragraph>
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</subSubSection>
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<subSubSection id="37C2EBDEFF9A3D1098481399FAD5FC85" pageId="22" pageNumber="23" type="description">
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<paragraph id="7F67B855FF9A3D1098481399FC89FDCD" blockId="22.[264,1325,284,870]" pageId="22" pageNumber="23">
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<emphasis id="4DAC6447FF9A3D1098481399FE1BFE6E" box="[312,491,444,469]" italics="true" pageId="22" pageNumber="23">
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Male
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<typeStatus id="A06306F7FF9A3D10980B1399FE13FE6E" box="[379,483,444,469]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
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:
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</emphasis>
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Length, 6.5 mm, width 0.9 mm. Color pale yellowish, unmarked. Segmental shoulders moderate; segmental setae to 80% body width. Eight to eleven ocelli, not pigmented, irregular in size and wellseparated, in two rows (typically of 6–7 and 3 ocelli). Pregonopodal legs somewhat more crassate than postgonopodal legs, legpairs 6 (
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<figureCitation id="E7E3A4D0FF9A3D109DB51011FAEBFDF5" box="[1221,1307,564,590]" captionStart="FIGURES 12 – 15" captionStartId="20.[264,382,1559,1583]" captionTargetBox="[336,1254,680,1540]" captionTargetId="figure@20.[331,1254,675,1548]" captionTargetPageId="20" captionText="FIGURES 12 – 15. Trigenotyla parca Causey, 12, gonopods, anterior view. Trigenotyla blacki n. sp., 13 – 15. 13, gonopods, anterior view. 14, male ninth legs, anterior view. 15, male right sixth leg, posterior view. Scale lines for Figs. 12 – 14 = 0.1 mm (100 µm), for Fig. 15, 0.5 mm (500 µm)." httpUri="https://zenodo.org/record/157040/files/figure.png" pageId="22" pageNumber="23">Fig. 15</figureCitation>
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), 7 largest; prefemora of 6 curved, with basal swelling.
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</paragraph>
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<paragraph id="7F67B855FF9A3D10984810A1FC5FFCAD" blockId="22.[264,1325,284,870]" pageId="22" pageNumber="23">
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Gonopods (
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<figureCitation id="E7E3A4D0FF9A3D1098B010A1FDE5FD25" box="[448,533,644,670]" captionStart="FIGURES 12 – 15" captionStartId="20.[264,382,1559,1583]" captionTargetBox="[336,1254,680,1540]" captionTargetId="figure@20.[331,1254,675,1548]" captionTargetPageId="20" captionText="FIGURES 12 – 15. Trigenotyla parca Causey, 12, gonopods, anterior view. Trigenotyla blacki n. sp., 13 – 15. 13, gonopods, anterior view. 14, male ninth legs, anterior view. 15, male right sixth leg, posterior view. Scale lines for Figs. 12 – 14 = 0.1 mm (100 µm), for Fig. 15, 0.5 mm (500 µm)." httpUri="https://zenodo.org/record/157040/files/figure.png" pageId="22" pageNumber="23">Fig. 13</figureCitation>
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) much as described for
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<emphasis id="4DAC6447FF9A3D109A5D10A1FC64FD26" box="[813,916,644,669]" italics="true" pageId="22" pageNumber="23">
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<taxonomicName id="B8D8C3D6FF9A3D109A5D10A1FC61FD26" box="[813,913,644,669]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="parca">T. parca</taxonomicName>
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,
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</emphasis>
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but setae somewhat less ensiform, more clearly socketed, lateral group not raised on ectal shoulder; ectal coxites absent, mesal coxites more robust, fimbriate branch prominent. Legpair 9 (
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<figureCitation id="E7E3A4D0FF9A3D109D2310F1FB5DFD55" box="[1107,1197,724,750]" captionStart="FIGURES 12 – 15" captionStartId="20.[264,382,1559,1583]" captionTargetBox="[336,1254,680,1540]" captionTargetId="figure@20.[331,1254,675,1548]" captionTargetPageId="20" captionText="FIGURES 12 – 15. Trigenotyla parca Causey, 12, gonopods, anterior view. Trigenotyla blacki n. sp., 13 – 15. 13, gonopods, anterior view. 14, male ninth legs, anterior view. 15, male right sixth leg, posterior view. Scale lines for Figs. 12 – 14 = 0.1 mm (100 µm), for Fig. 15, 0.5 mm (500 µm)." httpUri="https://zenodo.org/record/157040/files/figure.png" pageId="22" pageNumber="23">Fig. 14</figureCitation>
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) as for
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<emphasis id="4DAC6447FF9A3D109C6510F1FEA5FCAE" italics="true" pageId="22" pageNumber="23">
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<taxonomicName id="B8D8C3D6FF9A3D109C6510F1FEBEFCAE" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="parca">T. parca</taxonomicName>
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;
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</emphasis>
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legpairs 10, 11 with glands, otherwise unmodified.
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</paragraph>
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<paragraph id="7F67B855FF9A3D1098481101FAD5FC85" blockId="22.[264,1325,284,870]" box="[312,1317,804,830]" pageId="22" pageNumber="23">
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<emphasis id="4DAC6447FF9A3D1098481101FDF9FC86" box="[312,521,804,829]" italics="true" pageId="22" pageNumber="23">
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Female
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<typeStatus id="A06306F7FF9A3D1098E71101FDF3FC86" box="[407,515,804,829]" pageId="22" pageNumber="23" type="paratype">paratype</typeStatus>
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:
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</emphasis>
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Length, 6.4 mm, width 0.85 mm. Nonsexual characters as in male.
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</paragraph>
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</subSubSection>
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<subSubSection id="37C2EBDEFF9A3D1098481169FE61FA2D" pageId="22" pageNumber="23" type="distribution">
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<paragraph id="7F67B855FF9A3D1098481169FB21FCDD" blockId="22.[264,1325,284,870]" box="[312,1233,844,870]" pageId="22" pageNumber="23">
|
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<emphasis id="4DAC6447FF9A3D1098481169FE21FCDE" box="[312,465,844,869]" italics="true" pageId="22" pageNumber="23">Distribution:</emphasis>
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Known only from caves in Delaware and Adair Cos., Oklahoma.
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</paragraph>
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<paragraph id="7F67B855FF9A3D10984811B9FD93FBBD" blockId="22.[264,1323,924,1270]" pageId="22" pageNumber="23">
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||
<emphasis id="4DAC6447FF9A3D10984811B9FD97FC0E" box="[312,615,924,949]" italics="true" pageId="22" pageNumber="23">
|
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Notes:
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<taxonomicName id="B8D8C3D6FF9A3D1098FB11B9FD93FC0E" box="[395,611,924,949]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="blacki">Trigenotyla blacki</taxonomicName>
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,
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</emphasis>
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unlike the preceeding species, shows troglobitic adaptations in its reduced ocelli and pale pigmentation. As such it is presently the only troglobitic milliped known from Oklahoma.
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</paragraph>
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<paragraph id="7F67B855FF9A3D1098481631FE10FB4D" blockId="22.[264,1323,924,1270]" pageId="22" pageNumber="23">
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Causey recognized this species as new, and labelled it “reducta,” a name which was never published.
|
||
<bibRefCitation id="1B49C5A4FF9A3D1098AB1619FD86FBED" author="Black" box="[475,630,1084,1110]" pageId="22" pageNumber="35" refString="Black, J. H. (1971) The cave life of Oklahoma, a preliminary study (excluding Chiroptera). Oklahoma Underground, 4, 1 - 49." type="journal article" year="1971">Black (1971)</bibRefCitation>
|
||
lists “two undescribed species” of
|
||
<taxonomicName id="B8D8C3D6FF9A3D109D671619FB6EFBEE" box="[1047,1182,1084,1109]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4DAC6447FF9A3D109D671619FB6EFBEE" box="[1047,1182,1084,1109]" italics="true" pageId="22" pageNumber="23">Trigenotyla</emphasis>
|
||
</taxonomicName>
|
||
from Oklahoma caves, as determined by Causey. However, the present study reveals that actually three species are involved (see below). The Delaware Co. records refer to
|
||
<emphasis id="4DAC6447FF9A3D109DE316A9FB13FB1E" box="[1171,1251,1164,1189]" italics="true" pageId="22" pageNumber="23">
|
||
<taxonomicName id="B8D8C3D6FF9A3D109DE316A9FB2EFB1E" box="[1171,1246,1164,1189]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="blacki">blacki</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
those from
|
||
<taxonomicName id="B8D8C3D6FF9A3D1098391691FE48FB75" box="[329,440,1204,1230]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="34" phylum="Arthropoda" rank="species" species="seminole">Seminole</taxonomicName>
|
||
Co. to
|
||
<emphasis id="4DAC6447FF9A3D109B7E1691FD68FB76" box="[526,664,1204,1229]" italics="true" pageId="22" pageNumber="23">
|
||
<taxonomicName id="B8D8C3D6FF9A3D109B7E1691FD61FB76" box="[526,657,1204,1229]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="seminole" status="sp. nov.">T. seminole</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="569FD93CFF9A3D109BD21691FCC2FB75" box="[674,818,1204,1230]" pageId="22" pageNumber="23" rank="species">new species</taxonomicNameLabel>
|
||
, and those from Johnston and Murray Cos. to
|
||
<taxonomicName id="B8D8C3D6FF9A3D10985616F9FE10FB4D" authority="Causey." authorityName="Causey." box="[294,480,1244,1270]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="vaga">
|
||
<emphasis id="4DAC6447FF9A3D10985616F9FE8CFB4E" box="[294,380,1244,1269]" italics="true" pageId="22" pageNumber="23">T. vaga</emphasis>
|
||
Causey.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="7F67B855FF9A3D1098781771FE61FA2D" blockId="22.[264,674,1364,1430]" pageId="22" pageNumber="23">
|
||
<taxonomicName id="B8D8C3D6FF9A3D1098781771FDF5FAD5" box="[264,517,1364,1390]" class="Diplopoda" family="Trichopetalidae" genus="Trigneotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="seminole" status="sp. nov.">
|
||
<emphasis id="4DAC6447FF9A3D1098781771FDF5FAD5" bold="true" box="[264,517,1364,1390]" italics="true" pageId="22" pageNumber="23">Trigneotyla seminole</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="569FD93CFF9A3D109B651771FD52FAD5" box="[533,674,1364,1390]" pageId="22" pageNumber="23" rank="species">new species</taxonomicNameLabel>
|
||
<figureCitation id="E7E3A4D0FF9A3D1098781759FE61FA2D" box="[264,401,1404,1430]" captionStart="FIGURES 16 – 19" captionStartId="23.[264,382,1789,1813]" captionTargetBox="[298,1285,1222,1768]" captionTargetId="figure@23.[297,1290,1215,1775]" captionTargetPageId="23" captionText="FIGURES 16 – 19. Trigenotyla seminole, n. sp. 16, male right sixth leg, posterior view. 17, gonopods, anterior view. 18, male ninth legs, posterior view. 19, male right eleventh legpair, anterior view. Scale lines for Figs. 17 and 18 = 0.1 mm (100 µm), for Figs. 16 and 19, 0, 5 mm (500 µm)." httpUri="https://zenodo.org/record/157041/files/figure.png" pageId="22" pageNumber="23">Figs. 16–19</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="37C2EBDEFF9A3D10987817E9FCA6F93D" pageId="22" pageNumber="23" type="description">
|
||
<paragraph id="7F67B855FF9A3D10987817E9FCA6F93D" blockId="22.[264,1323,1484,1870]" pageId="22" pageNumber="23">
|
||
<emphasis id="4DAC6447FF9A3D10987817E9FEA2FA5E" box="[264,338,1484,1509]" italics="true" pageId="22" pageNumber="23">
|
||
<typeStatus id="A06306F7FF9A3D10987817E9FEB8FA5E" box="[264,328,1484,1509]" pageId="22" pageNumber="23">Types</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Male
|
||
<typeStatus id="A06306F7FF9A3D1098EE17E9FDF8FA5D" box="[414,520,1484,1510]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
, two male
|
||
<typeStatus id="A06306F7FF9A3D109BF617E9FD06FA5D" box="[646,758,1484,1510]" pageId="22" pageNumber="23" type="paratype">paratypes</typeStatus>
|
||
and one female
|
||
<typeStatus id="A06306F7FF9A3D109AC717E9FBECFA5D" box="[951,1052,1484,1510]" pageId="22" pageNumber="23" type="paratype">paratype</typeStatus>
|
||
from Whiskey Cave,
|
||
<quantity id="B82015B0FF9A3D109C6C17E9FEDBF9B5" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="22" pageNumber="23" unit="mi" value="9.0">9 mi</quantity>
|
||
. SE Bowlegs,
|
||
<taxonomicName id="B8D8C3D6FF9A3D10989117D1FDA0F9B5" box="[481,592,1524,1550]" class="Diplopoda" family="Trichopetalidae" genus="Trigneotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="34" phylum="Arthropoda" rank="species" species="seminole">Seminole</taxonomicName>
|
||
Co., Oklahoma, collected
|
||
<date id="0B669E95FF9A3D109AEA17D1FB96F9B5" box="[922,1126,1524,1550]" pageId="22" pageNumber="23" value="1974-01-14">14 January 1974</date>
|
||
, by J. H. Black (FSCA). The following specimens (all collected in
|
||
<taxonomicName id="B8D8C3D6FF9A3D109A021439FC10F98D" box="[882,992,1564,1590]" class="Diplopoda" family="Trichopetalidae" genus="Trigneotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="34" phylum="Arthropoda" rank="species" species="seminole">Seminole</taxonomicName>
|
||
Co. by J. H. Black; FSCA) are
|
||
<typeStatus id="A06306F7FF9A3D1098451461FE5AF9E5" box="[309,426,1604,1630]" pageId="22" pageNumber="23" type="paratype">paratypes</typeStatus>
|
||
: Doolin Cave,
|
||
<quantity id="B82015B0FF9A3D109B181461FD6FF9E5" box="[616,671,1604,1630]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="22" pageNumber="23" unit="mi" value="8.0">8 mi</quantity>
|
||
NW of Sasakwa,
|
||
<date id="0B669E95FF9A3D109A0C1461FBC5F9E5" box="[892,1077,1604,1630]" pageId="22" pageNumber="23" value="1974-03-14">14 March 1974</date>
|
||
, male; Cold Springs Cave,
|
||
<quantity id="B82015B0FF9A3D1098231449FE78F93D" box="[339,392,1644,1670]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="22" pageNumber="23" unit="mi" value="9.0">9 mi</quantity>
|
||
SE Bowlegs,
|
||
<date id="0B669E95FF9A3D109B5E1449FD02F93D" box="[558,754,1644,1670]" pageId="22" pageNumber="23" value="1974-01-14">14 January 1974</date>
|
||
, female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="37C2EBDEFF9A3D10984814B1FB4AF96D" pageId="22" pageNumber="23" type="diagnosis">
|
||
<paragraph id="7F67B855FF9A3D10984814B1FB4AF96D" blockId="22.[264,1323,1484,1870]" pageId="22" pageNumber="23">
|
||
<emphasis id="4DAC6447FF9A3D10984814B1FE49F916" box="[312,441,1684,1709]" italics="true" pageId="22" pageNumber="23">Diagnosis:</emphasis>
|
||
Resembles
|
||
<taxonomicName id="B8D8C3D6FF9A3D109B3514B1FD08F915" authority="Causey" authorityName="Causey" box="[581,760,1684,1710]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="vaga">
|
||
<emphasis id="4DAC6447FF9A3D109B3514B1FD6BF916" box="[581,667,1684,1709]" italics="true" pageId="22" pageNumber="23">T. vaga</emphasis>
|
||
Causey
|
||
</taxonomicName>
|
||
in having gonopod ectal coxites broad in lateral view, differs in the shape of those coxites, smaller size and lack of pigmentation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="37C2EBDEFF9A3D10984814C1FB41F945" box="[312,1201,1764,1790]" pageId="22" pageNumber="23" type="etymology">
|
||
<paragraph id="7F67B855FF9A3D10984814C1FB41F945" blockId="22.[264,1323,1484,1870]" box="[312,1201,1764,1790]" pageId="22" pageNumber="23">
|
||
<emphasis id="4DAC6447FF9A3D10984814C1FE4EF946" box="[312,446,1764,1789]" italics="true" pageId="22" pageNumber="23">Etymology:</emphasis>
|
||
The species name is a noun in apposition, for
|
||
<taxonomicName id="B8D8C3D6FF9A3D109AAF14C1FBBEF945" box="[991,1102,1764,1790]" class="Diplopoda" family="Trichopetalidae" genus="Trigneotyla" kingdom="Animalia" order="Chordeumatida" pageId="22" pageNumber="34" phylum="Arthropoda" rank="species" species="seminole">Seminole</taxonomicName>
|
||
County.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="37C2EBDEFF9A3D1198481529FAE6FD25" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
|
||
<paragraph id="7F67B855FF9A3D1198481529FCE6FEE5" blockId="22.[264,1323,1484,1870]" lastBlockId="23.[264,1323,284,1150]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
|
||
<emphasis id="4DAC6447FF9A3D1098481529FE1AF89E" box="[312,490,1804,1829]" italics="true" pageId="22" pageNumber="23">
|
||
Male
|
||
<typeStatus id="A06306F7FF9A3D10980B1529FE13F89E" box="[379,483,1804,1829]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Length, 5.0 mm, width 0.75 mm. Unpigmented. Segmental shoulders low; segmental setae (most rubbed off in
|
||
<typeStatus id="A06306F7FF9A3D109B9C1511FCEEF8F5" box="[748,798,1844,1870]" pageId="22" pageNumber="23">type</typeStatus>
|
||
series) about equal to body width. Fourteen to 17 round, wellpigmented ocelli in four rows forming triangular eyepatch. Pregonopodal legs enlarged, pair 6 (
|
||
<figureCitation id="E7E3A4D0FF9B3D119B401361FD75FEE5" box="[560,645,324,350]" captionStart="FIGURES 16 – 19" captionStartId="23.[264,382,1789,1813]" captionTargetBox="[298,1285,1222,1768]" captionTargetId="figure@23.[297,1290,1215,1775]" captionTargetPageId="23" captionText="FIGURES 16 – 19. Trigenotyla seminole, n. sp. 16, male right sixth leg, posterior view. 17, gonopods, anterior view. 18, male ninth legs, posterior view. 19, male right eleventh legpair, anterior view. Scale lines for Figs. 17 and 18 = 0.1 mm (100 µm), for Figs. 16 and 19, 0, 5 mm (500 µm)." httpUri="https://zenodo.org/record/157041/files/figure.png" pageId="23" pageNumber="24">Fig. 16</figureCitation>
|
||
) the largest.
|
||
</paragraph>
|
||
<paragraph id="7F67B855FF9B3D1198481349FE24FDCD" blockId="23.[264,1323,284,1150]" pageId="23" pageNumber="24">
|
||
Gonopods (
|
||
<figureCitation id="E7E3A4D0FF9B3D1198B01349FDE5FE3D" box="[448,533,364,390]" captionStart="FIGURES 16 – 19" captionStartId="23.[264,382,1789,1813]" captionTargetBox="[298,1285,1222,1768]" captionTargetId="figure@23.[297,1290,1215,1775]" captionTargetPageId="23" captionText="FIGURES 16 – 19. Trigenotyla seminole, n. sp. 16, male right sixth leg, posterior view. 17, gonopods, anterior view. 18, male ninth legs, posterior view. 19, male right eleventh legpair, anterior view. Scale lines for Figs. 17 and 18 = 0.1 mm (100 µm), for Figs. 16 and 19, 0, 5 mm (500 µm)." httpUri="https://zenodo.org/record/157041/files/figure.png" pageId="23" pageNumber="24">Fig. 17</figureCitation>
|
||
) with sternal lobes low; coxae not fused in midline; coxal setae not ensiform, in two groups, lateral group of 5 or 6 tightly clustered at distal ectal shoulder of coxa, mesal group of three near gonopod midline. Mesal coxites thin, evenly tapered, curved at tips, posterior edges not serrate; ectal coxites laterally flattened, in lateral view with 3 teeth. Ninth legpair (
|
||
<figureCitation id="E7E3A4D0FF9B3D119B271029FD5CFD9D" box="[599,684,524,550]" captionStart="FIGURES 16 – 19" captionStartId="23.[264,382,1789,1813]" captionTargetBox="[298,1285,1222,1768]" captionTargetId="figure@23.[297,1290,1215,1775]" captionTargetPageId="23" captionText="FIGURES 16 – 19. Trigenotyla seminole, n. sp. 16, male right sixth leg, posterior view. 17, gonopods, anterior view. 18, male ninth legs, posterior view. 19, male right eleventh legpair, anterior view. Scale lines for Figs. 17 and 18 = 0.1 mm (100 µm), for Figs. 16 and 19, 0, 5 mm (500 µm)." httpUri="https://zenodo.org/record/157041/files/figure.png" pageId="23" pageNumber="24">Fig. 18</figureCitation>
|
||
) typical of genus, but with 2–3 vestigial articles distal to prefemur; coxal glands prominent; legpairs 10, 11 (
|
||
<figureCitation id="E7E3A4D0FF9B3D119AE51011FC1CFDF5" box="[917,1004,564,590]" captionStart="FIGURES 16 – 19" captionStartId="23.[264,382,1789,1813]" captionTargetBox="[298,1285,1222,1768]" captionTargetId="figure@23.[297,1290,1215,1775]" captionTargetPageId="23" captionText="FIGURES 16 – 19. Trigenotyla seminole, n. sp. 16, male right sixth leg, posterior view. 17, gonopods, anterior view. 18, male ninth legs, posterior view. 19, male right eleventh legpair, anterior view. Scale lines for Figs. 17 and 18 = 0.1 mm (100 µm), for Figs. 16 and 19, 0, 5 mm (500 µm)." httpUri="https://zenodo.org/record/157041/files/figure.png" pageId="23" pageNumber="24">Fig. 19</figureCitation>
|
||
) with coxal glands, otherwise unmodified.
|
||
</paragraph>
|
||
<paragraph id="7F67B855FF9B3D11984810A1FAE6FD25" blockId="23.[264,1323,284,1150]" box="[312,1302,644,670]" pageId="23" pageNumber="24">
|
||
<emphasis id="4DAC6447FF9B3D11984810A1FDF9FD26" box="[312,521,644,669]" italics="true" pageId="23" pageNumber="24">
|
||
Female
|
||
<typeStatus id="A06306F7FF9B3D1198E710A1FDF3FD26" box="[407,515,644,669]" pageId="23" pageNumber="24" type="paratype">paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Length 5.0 mm, width, 0.8 mm. Nonsexual characters as in male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="37C2EBDEFF9B3D1198481089FD35FBC5" pageId="23" pageNumber="24" type="distribution">
|
||
<paragraph id="7F67B855FF9B3D1198481089FC48FCAE" blockId="23.[264,1323,284,1150]" pageId="23" pageNumber="24">
|
||
<emphasis id="4DAC6447FF9B3D1198481089FE21FD7E" box="[312,465,684,709]" italics="true" pageId="23" pageNumber="24">Distribution:</emphasis>
|
||
Known only from caves in
|
||
<taxonomicName id="B8D8C3D6FF9B3D119A551089FC64FD7D" box="[805,916,684,710]" class="Diplopoda" family="Trichopetalidae" genus="Trigneotyla" kingdom="Animalia" order="Chordeumatida" pageId="23" pageNumber="34" phylum="Arthropoda" rank="species" species="seminole">Seminole</taxonomicName>
|
||
Co., Oklahoma. All localites are north of the Canadian River, and it is hypothesized that this barrier separates the present species from the distribution of the closely related
|
||
<emphasis id="4DAC6447FF9B3D119A2A10D9FC48FCAE" box="[858,952,764,789]" italics="true" pageId="23" pageNumber="24">
|
||
<taxonomicName id="B8D8C3D6FF9B3D119A2A10D9FC44FCAE" box="[858,948,764,789]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="vaga">T. vaga</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="7F67B855FF9B3D1198481101FC51FC35" blockId="23.[264,1323,284,1150]" pageId="23" pageNumber="24">
|
||
<emphasis id="4DAC6447FF9B3D1198481101FD74FC86" box="[312,644,804,829]" italics="true" pageId="23" pageNumber="24">
|
||
Notes:
|
||
<taxonomicName id="B8D8C3D6FF9B3D1198FC1101FD74FC86" box="[396,644,804,829]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="seminole">Trigenotyla seminole</taxonomicName>
|
||
</emphasis>
|
||
populations are unpigmented or very lightly so, but have pigmented, wellformed, and numerous ocelli. Though the species has never been found on the surface, I suspect it is troglophilic, not troglobitic.
|
||
</paragraph>
|
||
<paragraph id="7F67B855FF9B3D11984811B9FD35FBC5" blockId="23.[264,1323,284,1150]" pageId="23" pageNumber="24">
|
||
Causey had labelled all Oklahoma
|
||
<taxonomicName id="B8D8C3D6FF9B3D119BAD11B9FC93FC0E" box="[733,867,924,949]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4DAC6447FF9B3D119BAD11B9FC93FC0E" box="[733,867,924,949]" italics="true" pageId="23" pageNumber="24">Trigenotyla</emphasis>
|
||
</taxonomicName>
|
||
(aside from the Delaware Co. collections described above as
|
||
<emphasis id="4DAC6447FF9B3D119B4211E1FD6FFC66" box="[562,671,964,989]" italics="true" pageId="23" pageNumber="24">
|
||
<taxonomicName id="B8D8C3D6FF9B3D119B4211E1FD65FC66" box="[562,661,964,989]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="blacki">T. blacki</taxonomicName>
|
||
)
|
||
</emphasis>
|
||
as “profunda,” an unpublished name. This judgement is reflected in Black’s (1971) treatment of Oklahoma
|
||
<emphasis id="4DAC6447FF9B3D119A0711C9FBF6FBBE" box="[887,1030,1004,1029]" italics="true" pageId="23" pageNumber="24">
|
||
<taxonomicName id="B8D8C3D6FF9B3D119A0711C9FBF2FBBE" box="[887,1026,1004,1029]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Trigenotyla</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
However,
|
||
<taxonomicName id="B8D8C3D6FF9B3D119DFB11C9FB0AFBBD" box="[1163,1274,1004,1030]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="23" pageNumber="34" phylum="Arthropoda" rank="species" species="seminole">Seminole</taxonomicName>
|
||
Co. collections north of the Canadian River are clearly distinct from those south of the river in being smaller, less pigmented, and have differences in the gonopods and ninth legs. Thus they are named here as a new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |