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<document id="A28D7C42A78E0540F1D4B1B9A7E3B13C" ID-DOI="10.5281/zenodo.157036" ID-GBIF-Dataset="8845a192-a81c-4e81-a280-e19c508ce7dd" ID-ISSN="1175­5326" ID-Zenodo-Dep="157036" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459633983606" checkinUser="plazi" docAuthor="Shear, William A." docDate="2003" docId="F7710943FF913D1898781631FE28FC66" docLanguage="en" docName="zt00321.pdf" docOrigin="Zootaxa 321" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Causeyella causeyae Shear, 2003, n. sp." docType="treatment" docVersion="10" lastPageNumber="31" masterDocId="0B48713BFF8C3D0699701225FFF0FFBB" masterDocTitle="The milliped family Trichopetalidae, Part 1: Introduction and Genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="30" updateTime="1698204205547" updateUser="plazi">
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<mods:title id="26594B1EE86265040C5AA5B6BF4BACFB">The milliped family Trichopetalidae, Part 1: Introduction and Genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea)</mods:title>
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<mods:namePart id="B0CC41FDBDA5346CD095097B53D5236B">Shear, William A.</mods:namePart>
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<mods:date id="8591ABD1A7AC7683ABC4DB5C6583E4F4">2003</mods:date>
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<treatment id="F7710943FF913D1898781631FE28FC66" ID-DOI="http://doi.org/10.5281/zenodo.6273596" ID-GBIF-Taxon="119332279" ID-Zenodo-Dep="6273596" LSID="urn:lsid:plazi:treatment:F7710943FF913D1898781631FE28FC66" httpUri="http://treatment.plazi.org/id/F7710943FF913D1898781631FE28FC66" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="37C2EBDEFF913D1B98781631FE62FBEA" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="7F67B855FF913D1B98781631FDA1FB95" blockId="29.[264,593,1044,1105]" box="[264,593,1044,1070]" pageId="29" pageNumber="30">
<heading id="242F0F39FF913D1B98781631FDA1FB95" bold="true" box="[264,593,1044,1070]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="4DAC6447FF913D1B98781631FDA1FB95" bold="true" box="[264,593,1044,1070]" pageId="29" pageNumber="30">
<emphasis id="4DAC6447FF913D1B98781631FDF7FB95" bold="true" box="[264,519,1044,1070]" italics="true" pageId="29" pageNumber="30">
<taxonomicName id="B8D8C3D6FF913D1B98781631FDF1FB95" box="[264,513,1044,1070]" class="Diplopoda" family="Trichopetalidae" genus="Causeyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="causeyae" status="sp. nov.">Causeyella causeyae</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="569FD93CFF913D1B9B7F1631FDA1FB95" box="[527,593,1044,1070]" pageId="29" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="7F67B855FF913D1B98781612FE62FBEA" blockId="29.[264,593,1044,1105]" box="[264,402,1079,1105]" pageId="29" pageNumber="30">
<figureCitation id="E7E3A4D0FF913D1B98781612FE62FBEA" box="[264,402,1079,1105]" captionStart="FIGURES 22 26" captionStartId="27.[264,382,1555,1579]" captionTargetBox="[278,1305,334,1536]" captionTargetId="figure@27.[273,1313,329,1544]" captionTargetPageId="27" captionText="FIGURES 22 26. Causeyella dendropus Loomis, 22, gonopods, anterior view. C. causeyae, n. sp., 23, 24. 23, gonopods, anterior view. 24, male ninth legs, anterior view. C. youngsteadtorum, n. sp., 25, 26. 25, gonopods, anterior view. 26, male ninth legs, anterior view. Scale lines = 0.1 mm (100 µm) for Figs. 22, 23, 25, 0.5 mm (500 µm) for Figs. 24, 26." httpUri="https://zenodo.org/record/157043/files/figure.png" pageId="29" pageNumber="30">Figs. 23, 24</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="37C2EBDEFF913D1B987816A2FB1AF9E2" pageId="29" pageNumber="30" type="description">
<paragraph id="7F67B855FF913D1B987816A2FB1AF9E2" blockId="29.[264,1325,1159,1905]" pageId="29" pageNumber="30">
<emphasis id="4DAC6447FF913D1B987816A2FEA2FB1B" box="[264,338,1159,1184]" italics="true" pageId="29" pageNumber="30">
<typeStatus id="A06306F7FF913D1B987816A2FEB8FB1B" box="[264,328,1159,1184]" pageId="29" pageNumber="30">Types</typeStatus>
:
</emphasis>
Male
<typeStatus id="A06306F7FF913D1B98EF16A2FDF9FB1A" box="[415,521,1159,1185]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, male
<typeStatus id="A06306F7FF913D1B9B2416A2FD49FB1A" box="[596,697,1159,1185]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
and
<specimenCount id="69DE73DCFF913D1B9B8216A2FCA7FB1A" box="[754,855,1159,1185]" pageId="29" pageNumber="30" type="female">2 female</specimenCount>
<typeStatus id="A06306F7FF913D1B9A2E16A2FC3EFB1A" box="[862,974,1159,1185]" pageId="29" pageNumber="30" type="paratype">paratypes</typeStatus>
from ARKANSAS,
<emphasis id="4DAC6447FF913D1B9DB216A2FE86FB73" italics="true" pageId="29" pageNumber="30">Independence Co</emphasis>
., Foushee Cave,
<quantity id="B82015B0FF913D1B9B33168AFD69FB72" box="[579,665,1199,1225]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="29" pageNumber="30" unit="mi" value="6.0">6 miles</quantity>
west of Locust Grove, collected by unknown person or persons, May, 1978 (FSCA). The following are
<typeStatus id="A06306F7FF913D1B9A2616F2FC3BFB4A" box="[854,971,1239,1265]" pageId="29" pageNumber="30" type="paratype">paratypes</typeStatus>
: ARKANSAS:
<emphasis id="4DAC6447FF913D1B9DF816F2FEDAFAA3" italics="true" pageId="29" pageNumber="30">Independence Co</emphasis>
., Cushman Cave, 0.5 miles from Cushman,
<date id="0B669E95FF913D1B9A1116DAFC0BFAA2" box="[865,1019,1279,1305]" pageId="29" pageNumber="30" value="1974-07-08">8 July 1974</date>
, G. Harp, four males,
<specimenCount id="69DE73DCFF913D1B9C6C16DAFE97FAFA" pageId="29" pageNumber="30" type="female">3 females</specimenCount>
; Dodd Cave,
<date id="0B669E95FF913D1B9B771702FD19FAFA" box="[519,745,1319,1345]" pageId="29" pageNumber="30" value="2001-12-14">14 December 2001</date>
, G. Greening, D. Fenolio, male (SBP).
<emphasis id="4DAC6447FF913D1B9DC51702FAD3FAFB" box="[1205,1315,1319,1344]" italics="true" pageId="29" pageNumber="30">Izard Co.</emphasis>
, Clay Cave,
<quantity id="B82015B0FF913D1B98E2176AFE37FAD2" box="[402,455,1359,1385]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="29" pageNumber="30" unit="mi" value="7.0">7 mi</quantity>
SW Melbourne,
<date id="0B669E95FF913D1B9BE1176AFCA8FAD2" box="[657,856,1359,1385]" pageId="29" pageNumber="30" value="1974-10-11">11 October 1974</date>
, G. Harp, numerous males and females; Donovan Cave,
<quantity id="B82015B0FF913D1B98B91752FDF0FA2A" box="[457,512,1399,1425]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="29" pageNumber="30" unit="mi" value="2.0">2 mi</quantity>
SW Calico Rock,
<date id="0B669E95FF913D1B9B921752FC5AFA2A" box="[738,938,1399,1425]" pageId="29" pageNumber="30" value="1964-01-24">24 January 1964</date>
, D. Martin, male; Goat (Prison) Cave,
<date id="0B669E95FF913D1B982717BAFDB6FA02" box="[343,582,1439,1465]" pageId="29" pageNumber="30" value="2002-11-22">22 November 2002</date>
, B. Wagner, D. Kempwerth, male (SBP).
<emphasis id="4DAC6447FF913D1B9D2C17BAFB21FA03" box="[1116,1233,1439,1464]" italics="true" pageId="29" pageNumber="30">Stone Co.</emphasis>
, Hells Creek Cave, 2.5 mi N Mountain View,
<date id="0B669E95FF913D1B9BAD17E2FC55FA5A" box="[733,933,1479,1505]" pageId="29" pageNumber="30" value="1974-01-26">26 January 1974</date>
, V. R. McDaniel,
<specimenCount id="69DE73DCFF913D1B9D0D17E2FB2CFA5A" box="[1149,1244,1479,1505]" pageId="29" pageNumber="30" type="male">2 males</specimenCount>
; Blanchard Springs Cave,
<date id="0B669E95FF913D1B988D17CAFD13F9B2" box="[509,739,1519,1545]" pageId="29" pageNumber="30" value="1967-09-27">27 September 1967</date>
, T. C. Barr, juveniles probably this species; Rowland Cave,
<date id="0B669E95FF913D1B98FC1432FDDAF98A" box="[396,554,1559,1585]" pageId="29" pageNumber="30" value="2001-06-12">12 June 2001</date>
, G. Greening et al., male (SBP); Alexander Cave,
<date id="0B669E95FF913D1B9D031432FAD7F98A" box="[1139,1319,1559,1585]" pageId="29" pageNumber="30" value="2001-03-24">24 March 2001</date>
, M. Slay, L. Moritz; Gustafson Cave,
<date id="0B669E95FF913D1B9BCA141AFC83F9E2" box="[698,883,1599,1625]" pageId="29" pageNumber="30" value="2002-10-06">6 October 2002</date>
, G. Graening et al., male (SBP).
</paragraph>
</subSubSection>
<subSubSection id="37C2EBDEFF913D1B98481442FB60F912" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="7F67B855FF913D1B98481442FB60F912" blockId="29.[264,1325,1159,1905]" pageId="29" pageNumber="30">
<emphasis id="4DAC6447FF913D1B98481442FE49F93B" box="[312,441,1639,1664]" italics="true" pageId="29" pageNumber="30">Diagnosis:</emphasis>
A large (13.5 mm long) species of
<taxonomicName id="B8D8C3D6FF913D1B9A151442FC17F93B" box="[869,999,1639,1664]" class="Diplopoda" family="Trichopetalidae" genus="Causeyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="4DAC6447FF913D1B9A151442FC17F93B" box="[869,999,1639,1664]" italics="true" pageId="29" pageNumber="30">Causeyella</emphasis>
</taxonomicName>
with lamellate branches of the gonopod colpocoxites; frontal coxal setae somewhat thickened and blunt.
</paragraph>
</subSubSection>
<subSubSection id="37C2EBDEFF913D1B98481492FD28F96A" box="[312,728,1719,1745]" pageId="29" pageNumber="30" type="etymology">
<paragraph id="7F67B855FF913D1B98481492FD28F96A" blockId="29.[264,1325,1159,1905]" box="[312,728,1719,1745]" pageId="29" pageNumber="30">
<emphasis id="4DAC6447FF913D1B98481492FE4FF96B" box="[312,447,1719,1744]" italics="true" pageId="29" pageNumber="30">Etymology:</emphasis>
For Nell Bevel Causey.
</paragraph>
</subSubSection>
<subSubSection id="37C2EBDEFF913D18984814FAFADBFDCD" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
<paragraph id="7F67B855FF913D18984814FAFDDEFDF5" blockId="29.[264,1325,1159,1905]" lastBlockId="30.[264,1323,284,990]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<emphasis id="4DAC6447FF913D1B984814FAFE1CF943" box="[312,492,1759,1784]" italics="true" pageId="29" pageNumber="30">
Male
<typeStatus id="A06306F7FF913D1B980D14FAFE15F943" box="[381,485,1759,1784]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
:
</emphasis>
Length, 13.5 mm, width 0.9 mm. Unpigmented, eyeless. Segmental shoulders well developed, paranota­like; segmental setae 6570% of body width. Legpairs 37 noticeably enlarged compared to legpairs 12 and posterior; no podomeres modified. Gonopods (
<figureCitation id="E7E3A4D0FF913D1B98AB1572FDC3F8CA" box="[475,563,1879,1905]" captionStart="FIGURES 22 26" captionStartId="27.[264,382,1555,1579]" captionTargetBox="[278,1305,334,1536]" captionTargetId="figure@27.[273,1313,329,1544]" captionTargetPageId="27" captionText="FIGURES 22 26. Causeyella dendropus Loomis, 22, gonopods, anterior view. C. causeyae, n. sp., 23, 24. 23, gonopods, anterior view. 24, male ninth legs, anterior view. C. youngsteadtorum, n. sp., 25, 26. 25, gonopods, anterior view. 26, male ninth legs, anterior view. Scale lines = 0.1 mm (100 µm) for Figs. 22, 23, 25, 0.5 mm (500 µm) for Figs. 24, 26." httpUri="https://zenodo.org/record/157043/files/figure.png" pageId="29" pageNumber="30">Fig. 23</figureCitation>
) with well developed sternal lobes. Anterior surfaces of coxae entirely covered with prominent scaly trichomes becoming larger distally; about 15 setae on each coxa, not separated into groups; mesal coxites in anterior view slightly spatulate, ectal coxites only about half their length, broad in lateral view. Lamellate branch of colpocoxite very large, curled at tip, roughened with rows of blunt trichomes, extending anterior of mesal coxites. Ninth legpair (
<figureCitation id="E7E3A4D0FF923D189BF51399FD2BFE6D" box="[645,731,444,470]" captionStart="FIGURES 22 26" captionStartId="27.[264,382,1555,1579]" captionTargetBox="[278,1305,334,1536]" captionTargetId="figure@27.[273,1313,329,1544]" captionTargetPageId="27" captionText="FIGURES 22 26. Causeyella dendropus Loomis, 22, gonopods, anterior view. C. causeyae, n. sp., 23, 24. 23, gonopods, anterior view. 24, male ninth legs, anterior view. C. youngsteadtorum, n. sp., 25, 26. 25, gonopods, anterior view. 26, male ninth legs, anterior view. Scale lines = 0.1 mm (100 µm) for Figs. 22, 23, 25, 0.5 mm (500 µm) for Figs. 24, 26." httpUri="https://zenodo.org/record/157043/files/figure.png" pageId="30" pageNumber="31">Fig. 24</figureCitation>
) with coxae laterally flattened, mesally excavated, bearing vestigial glands, articulation of prefemora offset laterally; prefemora longer than coxae, two or three vaguely indicated articles distally, claw short, twisted. Legpairs 10, 11 with glands, unmodified.
</paragraph>
<paragraph id="7F67B855FF923D1898481079FADBFDCD" blockId="30.[264,1323,284,990]" box="[312,1323,604,630]" pageId="30" pageNumber="31">
<emphasis id="4DAC6447FF923D1898481079FDF8FDCE" box="[312,520,604,629]" italics="true" pageId="30" pageNumber="31">
Female
<typeStatus id="A06306F7FF923D1898E61079FDF1FDCE" box="[406,513,604,629]" pageId="30" pageNumber="31" type="paratype">paratype</typeStatus>
:
</emphasis>
Length, 13.5 mm, width 1.05 mm. Nonsexual characters as in male.
</paragraph>
</subSubSection>
<subSubSection id="37C2EBDEFF923D18984810A1FE28FC66" pageId="30" pageNumber="31" type="distribution">
<paragraph id="7F67B855FF923D18984810A1FEB5FC36" blockId="30.[264,1323,284,990]" pageId="30" pageNumber="31">
<emphasis id="4DAC6447FF923D18984810A1FE21FD26" box="[312,465,644,669]" italics="true" pageId="30" pageNumber="31">Distribution:</emphasis>
Independence, Stone, and Izard counties, ARKANSAS, as listed under “
<typeStatus id="A06306F7FF923D1898621089FEADFD7D" box="[274,349,684,710]" pageId="30" pageNumber="31">Types</typeStatus>
” above. Literature records: ARKANSAS:
<emphasis id="4DAC6447FF923D189A261089FC3BFD7E" box="[854,971,684,709]" italics="true" pageId="30" pageNumber="31">Izard Co.,</emphasis>
Bergren Cave, Needles Cave;
<emphasis id="4DAC6447FF923D18987810F1FE89FD56" box="[264,377,724,749]" italics="true" pageId="30" pageNumber="31">Stone Co.</emphasis>
, Hell Creek Cave (as “
<taxonomicName id="B8D8C3D6FF923D189BE110F1FCF3FD56" box="[657,771,724,749]" class="Diplopoda" family="Trichopetalidae" genus="Scoterpes" kingdom="Animalia" order="Chordeumatida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="4DAC6447FF923D189BE110F1FCF3FD56" box="[657,771,724,749]" italics="true" pageId="30" pageNumber="31">Scoterpes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="569FD93CFF923D189A7C10F1FCBDFD55" box="[780,845,724,750]" pageId="30" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
” in
<bibRefCitation id="1B49C5A4FF923D189AF110F1FB37FD55" author="McDaniel" box="[897,1223,724,750]" pageId="30" pageNumber="36" refString="McDaniel, L., Rick, V. &amp; Smith, K. L. (1976) Cave fauna of Arkansas: Selected invertebrate taxa. Arkansas Academy of Science Proceedings, 30, 57 - 60." type="journal article" year="1976">McDaniel and Smith, 1976</bibRefCitation>
). These records were not verified; the wherabouts of the specimens are unknown. It appears that this species occurs on both sides of the White River. Records of “
<emphasis id="4DAC6447FF923D189D7A1101FB7CFC86" box="[1034,1164,804,829]" italics="true" pageId="30" pageNumber="31">
<taxonomicName id="B8D8C3D6FF923D189D7A1101FB72FC86" box="[1034,1154,804,829]" class="Diplopoda" family="Trichopetalidae" genus="Scoterpes" kingdom="Animalia" order="Chordeumatida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">Scoterpes</taxonomicName>
</emphasis>
from Oregon Co., Missouri (see under
<taxonomicName id="B8D8C3D6FF923D189B4C1169FD2CFCDE" box="[572,732,844,869]" class="Diplopoda" family="Trichopetalidae" genus="Scoterpes" kingdom="Animalia" order="Chordeumatida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="dendropus">
<emphasis id="4DAC6447FF923D189B4C1169FD2CFCDE" box="[572,732,844,869]" italics="true" pageId="30" pageNumber="31">C. dendropus</emphasis>
</taxonomicName>
above) may be
<emphasis id="4DAC6447FF923D189AD21169FBC9FCDE" box="[930,1081,844,869]" italics="true" pageId="30" pageNumber="31">
<taxonomicName id="B8D8C3D6FF923D189AD21169FBC5FCDE" box="[930,1077,844,869]" class="Diplopoda" family="Trichopetalidae" genus="Causeyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="causeyae">C. causeyae</taxonomicName>
,
</emphasis>
but are probably in error.
</paragraph>
<paragraph id="7F67B855FF923D18984811B9FE28FC66" blockId="30.[264,1323,284,990]" pageId="30" pageNumber="31">
Causey labelled examples of this species as “
<taxonomicName id="B8D8C3D6FF923D189A3311B9FC48FC0D" box="[835,952,924,950]" class="Diplopoda" family="Trichopetalidae" genus="Scoterpes" kingdom="Animalia" order="Chordeumatida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">Scoterpes</taxonomicName>
martini
<emphasis id="4DAC6447FF923D189D6311B9FBD3FC0E" box="[1043,1059,924,949]" italics="true" pageId="30" pageNumber="31"></emphasis>
or “
<taxonomicName id="B8D8C3D6FF923D189D2411B9FE35FC65" class="Diplopoda" family="Trichopetalidae" genus="Scoterpes" kingdom="Animalia" order="Chordeumatida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="dendropus" subSpecies="martini">Scoterpes dendropus ssp. martini</taxonomicName>
.
<emphasis id="4DAC6447FF923D1898B811E1FE28FC66" box="[456,472,964,989]" italics="true" pageId="30" pageNumber="31"></emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>