267 lines
33 KiB
XML
267 lines
33 KiB
XML
<document id="6774F69A8BFE15E084C2D6F6DE71687B" ID-CLB-Dataset="34512" ID-DOI="http://dx.doi.org/10.3897/zookeys.603.9038" ID-GBIF-Dataset="4b7c1390-937e-4d41-82c6-948f828ee497" ID-PMC="PMC4978001" ID-Pensoft-Pub="1313-2970-603-71" ID-PubMed="27551197" ID-ZooBank="86AB7257735E499CB64A3C87D4D62390" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-603-71" ModsDocOrigin="ZooKeys 603" ModsDocTitle="Two new species of Pachylaelaps Berlese, 1888 from the Iberian Peninsula, with a key to European species (Acari, Gamasida, Pachylaelapidae)" checkinTime="1467898551323" checkinUser="pensoft" docAuthor="Masan, Peter, Oezbek, Hasan Hueseyin & Fenďa, Peter" docDate="2016" docId="F6CCEAD53F0362F822E8D321024BF288" docLanguage="en" docName="ZooKeys 603: 71-95" docOrigin="ZooKeys 603" docSource="http://dx.doi.org/10.3897/zookeys.603.9038" docTitle="Pachylaelaps (Pachylaelaps) pyrenaicus Masan, Oezbek & Fenďa, 2016, sp. n." docType="treatment" docUuid="4E352060-75DC-4D92-A1FE-8BCDC82CD6B7" docUuidSource="ZooBank" docVersion="7" lastPageNumber="78" masterDocId="FFF4FFC0C312FFF1FFA30A49FFEFFF90" masterDocTitle="Two new species of Pachylaelaps Berlese, 1888 from the Iberian Peninsula, with a key to European species (Acari, Gamasida, Pachylaelapidae)" masterLastPageNumber="95" masterPageNumber="71" pageNumber="73" updateTime="1701376385657" updateUser="ExternalLinkService">
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<mods:title id="462A219AD5177015EB3B2684BCD25FFF">Two new species of Pachylaelaps Berlese, 1888 from the Iberian Peninsula, with a key to European species (Acari, Gamasida, Pachylaelapidae)</mods:title>
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<mods:namePart id="25902CF813E793E7B5C39D6365C83657">Masan, Peter</mods:namePart>
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<mods:namePart id="D14EC35F0AFF07B2A75D9950523F7CF1">Oezbek, Hasan Hueseyin</mods:namePart>
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<mods:namePart id="757C169DF3ABCB4345491691859FCFF5">Fenďa, Peter</mods:namePart>
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<mods:title id="D205486D3577F34B20B92091B949003C">ZooKeys</mods:title>
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<mods:date id="5215FC2AE5E2D2F2134A676E6A2C0BE3">2016</mods:date>
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<mods:classification id="F03BED242F1234848DD643D265DCF297">journal article</mods:classification>
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<mods:identifier id="6529133A80469F712F2EACF510BD275B" type="DOI">http://dx.doi.org/10.3897/zookeys.603.9038</mods:identifier>
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<treatment id="F6CCEAD53F0362F822E8D321024BF288" ID-GBIF-Taxon="127884163" LSID="urn:lsid:zoobank.org:act:4E352060-75DC-4D92-A1FE-8BCDC82CD6B7" httpUri="http://treatment.plazi.org/id/F6CCEAD53F0362F822E8D321024BF288" lastPageId="7" lastPageNumber="78" pageId="2" pageNumber="73">
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<subSubSection id="93B14E57B4268367ED2C12DFFDF0B0F8" pageId="2" pageNumber="73" type="multiple">
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<paragraph id="8D44F84DE1664C7606621A7E2329DF15" pageId="2" pageNumber="73">Taxon classification Animalia Mesostigmata Pachylaelapidae</paragraph>
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</subSubSection>
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<subSubSection id="1D6E9FACD4962F0BB11E65D98FFBCDD2" pageId="2" pageNumber="73" type="nomenclature">
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<paragraph id="FCC861F98A836050FD2F89B1F76C03C9" pageId="2" pageNumber="73">
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<taxonomicName id="59F3B6A15853ECB51A9B107D3EA91EDE" ID-CoL="8Y7LW" LSID="http://zoobank.org/4E352060-75DC-4D92-A1FE-8BCDC82CD6B7" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps (Pachylaelaps) pyrenaicus" order="Mesostigmata" pageId="2" pageNumber="73" phylum="Arthropoda" rank="species" species="pyrenaicus" subGenus="Pachylaelaps">Pachylaelaps (Pachylaelaps) pyrenaicus</taxonomicName>
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<taxonomicNameLabel id="BF6A384E4F88C9B24A330519FE3030A2" pageId="2" pageNumber="73">sp. n.</taxonomicNameLabel>
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Figures 1-2, 3-7, 8-12, 13-19
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</paragraph>
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</subSubSection>
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<subSubSection id="B5BBC60351B0C8B5710B566A965F6039" pageId="2" pageNumber="73" type="materials_examined">
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<paragraph id="E8EBA0AB6C04F6E0D8E78F1CB5D39032" pageId="2" pageNumber="73">Specimens examined.</paragraph>
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<paragraph id="1CF328F9B736BBFB4E23D7F39D079BA2" pageId="2" pageNumber="73">
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Holotype female: North Spain, Central Pyrenees Mts., Cinca Valley, Bielsa Cadaster, Salinas Village (near-by San Marcial Settlement), pine forest (
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<taxonomicName id="9133E97952F8658F61BEFCE7ED50A9EC" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus" order="Pinales" pageId="2" pageNumber="73" phylum="Tracheophyta" rank="genus">Pinus</taxonomicName>
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spp.) with admixed beech (
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<taxonomicName id="014A34689450DCF4B501D52B0725B8E5" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Fagus sylvatica" order="Fagales" pageId="2" pageNumber="73" phylum="Tracheophyta" rank="species" species="sylvatica">Fagus sylvatica</taxonomicName>
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), soil detritus with deep layer of raw humus between rock boulders, altitude 1050 m,
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<geoCoordinate id="0B778D84710BA2FA360328354736E32E" direction="north" orientation="latitude" precision="1" value="42.59783">42°35'52,2"N</geoCoordinate>
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,
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<geoCoordinate id="8DABC00E242587A56FFE42EF3740181F" direction="east" orientation="longitude" precision="1" value="0.23888889">00°14'20,0"E</geoCoordinate>
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, 16 June 2007, coll. P.
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<normalizedToken id="0EE7F41EFE36C56B8E3845143A5B4B75" originalValue="Fenďa">Fenďa</normalizedToken>
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. Paratypes: four females and one male, with the same data as the holotype. The holotype and four paratypes are deposited at the Institute of Zoology, Slovak Academy of Sciences, Bratislava; one female paratype is deposited at the Acarology Laboratory of Erzincan University, Turkey.
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</paragraph>
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</subSubSection>
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<subSubSection id="D9FBB7BE8B62331A39DE064EDB36D091" pageId="2" pageNumber="73" type="diagnosis">
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<paragraph id="276ACB4624AEFD34F17EE22F7A7A2062" pageId="2" pageNumber="73">Diagnosis.</paragraph>
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<paragraph id="5D72CA9EB9B0EDD0BA8C9C82EDC8D8FB" pageId="2" pageNumber="73">
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Slit-like glandular poroids gdZ1 and gdS4 with conspicuously adjacent position. Soft integument with decreased number of 11 pairs of setae in female and eight setal pairs in male. Prestigmatic section of peritreme long, with anterior tip reaching dorsal surface close to setae z1. Dorsal setae long (the longest setae more than 100
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<normalizedToken id="47C4C1801FA7A7E63ED0F5553FCAB2D0" originalValue="μm">μm</normalizedToken>
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in length), and seta j5 with tip reaching base of following seta z5. Cheliceral digits unidentate. Male palptibia with two well developed petal-like projections. In female, ventrodistal femur with small spine-like process associated with a seta. Terminal part of male tarsus II with only one spur-like distal seta (pl1). Sperm induction system with tubular components: tubes irregularly formed, folded, curved or with small bumps on distal sections, progressively widened basally; basal part widely abutting to anterior margin of coxa IV.
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</paragraph>
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</subSubSection>
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<subSubSection id="DE750B016852A3C5FA351FAEC0F94C20" lastPageId="6" lastPageNumber="77" pageId="2" pageNumber="73" type="description">
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<paragraph id="F3560E1532B076EAF136BDDBD3065858" pageId="2" pageNumber="73">Description.</paragraph>
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<paragraph id="1A9E6FFCF10B0495142B31E7813E0579" lastPageId="3" lastPageNumber="74" pageId="2" pageNumber="73">
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Female. Dorsal idiosoma (Figure 1). Dorsal shield 870-915
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<normalizedToken id="61D9FAF02B4FC02FC1049DAD57733C36" originalValue="μm">μm</normalizedToken>
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long and 560-610
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<normalizedToken id="8382F19D49455BFD26AFB3F222941145" originalValue="μm">μm</normalizedToken>
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wide, suboval (length/width 1.48-1.63), weakly and unevenly reticulated on surface, and bearing 30 pairs of smooth and needle-shaped dorsal setae. Setae z1 conspicuously shortened, setae J5 strongly reduced in length, vestigial microsetae; other setae relatively longer, subequal and uniform. Length and spacing of some selected dorsal shield setae as follows: j1 53-67
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<normalizedToken id="0286B31BC382757E7B98768F633A9C2D" originalValue="μm">μm</normalizedToken>
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, j5 73-83
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<normalizedToken id="EE89A303045B1575F007FDC675447FF5" originalValue="μm">μm</normalizedToken>
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, j5-j5 128-144
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<normalizedToken id="0D4F3517EEEF8644936F00DB0FD596BA" originalValue="μm">μm</normalizedToken>
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, j5-z5 72-81
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<normalizedToken id="126B0C32D5D8DC80A9A04225F00CFCE4" originalValue="μm">μm</normalizedToken>
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, J1-J2 87-111
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<normalizedToken id="D8F83289957C91D37CC098766576D4DF" originalValue="μm">μm</normalizedToken>
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, J2 97-109
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<normalizedToken id="E0DCBAB949836F42C06F57098F9CD772" originalValue="μm">μm</normalizedToken>
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, J2-J2 216-242
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<normalizedToken id="05004833EEB6B5588EF46A382DC880F9" originalValue="μm">μm</normalizedToken>
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, J2-J3 142-172
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<normalizedToken id="4DE1D355E08F3284034B4671617E9F1E" originalValue="μm">μm</normalizedToken>
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, J3 102-110
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<normalizedToken id="7FED2C7119019666007C6F4C533845A3" originalValue="μm">μm</normalizedToken>
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, J3-J4 118-151
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<normalizedToken id="01F23CD1791EB21267FB768E476F736F" originalValue="μm">μm</normalizedToken>
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, and J4 100-105
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<normalizedToken id="CCAFF8E4FBDA595BE8C91038C82E2414" originalValue="μm">μm</normalizedToken>
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. Dorsolateral
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<pageBreakToken id="AC27A777DC770C7444D313083FB69A15" pageId="3" pageNumber="74" start="start">soft</pageBreakToken>
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integument with four pairs of marginal setae (r6, R5‒R7). Posterolateral poroid structures gdZ1 and gdS4 slit-like, markedly adjacent each other, and placed close to setae Z2 or rarely between setae Z2 and S4.
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</paragraph>
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<caption id="FA48693A6B547ECF3565AC0B78042AD2" pageId="3" pageNumber="74">
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<paragraph id="5987BF63788BDD2F7EE1673DC73BE498" pageId="3" pageNumber="74">
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Figures 1-2.
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<taxonomicName id="B7CF260BEE854E6533683FF2F42C92B8" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps (Pachylaelaps) pyrenaicus" order="Mesostigmata" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="pyrenaicus" subGenus="Pachylaelaps">Pachylaelaps (Pachylaelaps) pyrenaicus</taxonomicName>
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, female, with setal notation of some idiosomal setae and glandular poroids. 1 Dorsal idiosoma 2 Ventral idiosoma.
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</paragraph>
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</caption>
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<paragraph id="597BD6B3E9A667A55A7928B1DAD91500" lastPageId="4" lastPageNumber="75" pageId="3" pageNumber="74">
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Ventral idiosoma (Figure 2). Sternal shield 272-280
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<normalizedToken id="8A8C460F5BEC4741C928ECD816F70025" originalValue="μm">μm</normalizedToken>
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long, proportionally 0.82-0.94 shorter than genitiventral shield, with concave anterior margin and two small projections close to bases of sternal setae st1. Genitiventral shield slightly shorter than wide or subequal in length and width (length 295-335
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<normalizedToken id="5A3554A4D99503DDFEDFBC518E00DEEB" originalValue="μm">μm</normalizedToken>
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, width 308-337
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<normalizedToken id="94ACA28265E2AD133522D9FAF891D563" originalValue="μm">μm</normalizedToken>
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, length/width 0.94-0.99). Anal shield subtriangular, 114-137
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<normalizedToken id="8705DD4C974D21897AC66546EA2E5A73" originalValue="μm">μm</normalizedToken>
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long and 170-199
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<normalizedToken id="083F8D6C3381C679B6F892772E56C1CC" originalValue="μm">μm</normalizedToken>
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wide (length/width 0.62-0.73); anus with circum-anal setae situated close to pos
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<pageBreakToken id="BA1DB69724FDEDE8358F278F40AF9189" pageId="4" pageNumber="75" start="start">terior</pageBreakToken>
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margin of shield. Peritremes well developed, relatively long, with anterior tip reaching dorsal surface close to setae z1. Peritrematal shields with weak longitudinal sculptural lines, other ventral shields distinctly and evenly reticulated on surface. Metapodal platelets minuscule, free and well separate from peritrematal shields. Ventral soft integument with seven pairs of ventral setae (JV2-JV5, ZV2, ZV4, ZV5). Ventral setae similar to those on dorsal idiosoma.
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</paragraph>
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<paragraph id="D540EE826BA9A810414B668B4547107A" pageId="4" pageNumber="75">Sperm induction structures (Figures 3, 13‒16). Tubes of sperm induction system relatively well developed, well sclerotized, broadened basally, and narrowed distally; worm-like distal section irregularly formed, folded, curved or with small bumps; basal section widely abutting to anterior margin of coxa IV.</paragraph>
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<caption id="8DBADAFA46DF61B8F4974F1EBAFF559A" pageId="4" pageNumber="75">
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<paragraph id="0006605566DD5E1CFACEAB2C718B57FA" pageId="4" pageNumber="75">
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Figures 3-7.
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<taxonomicName id="1A17B29234CD07A7B73366508B91D015" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps (Pachylaelaps) pyrenaicus" order="Mesostigmata" pageId="4" pageNumber="75" phylum="Arthropoda" rank="species" species="pyrenaicus" subGenus="Pachylaelaps">Pachylaelaps (Pachylaelaps) pyrenaicus</taxonomicName>
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, female. 3 Sperm induction structures 4 Ventral gnathosoma 5 Epistome 6 Chelicera, lateral view 7 Tarsus II, lateral view.
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</paragraph>
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</caption>
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<paragraph id="25E2C4D929648CA52CF21CA8E19E8EA6" pageId="5" pageNumber="76">
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<pageBreakToken id="0462F5E74D2C3B4A2BBF7F572911AAE9" pageId="5" pageNumber="76" start="start">Gnathosomal</pageBreakToken>
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structures (Figures 4‒6, 19). Corniculi elongated and horn-like; laciniae densely pilose, slightly longer than corniculi; deutosternum slightly widened medially, with six rows of denticles; subcapitular setae smooth and needle-shaped (Figure 4). Epistome with wide subtriangular base, elongate and narrow central neck and thin apical part crenelated on anterior margin; lateral margins of basal part with delicate denticulation; apical section not expanded or only very slightly expanded anteriorly, terminally truncate and with a row of four to seven prongs (Figure 5). Cheliceral digits relatively elongate and slender (Figures 6, 19), 100-110
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<normalizedToken id="14D74EA10C7A2DB211B80E09CCBC2C20" originalValue="μm">μm</normalizedToken>
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long; fixed digit of chelicera with terminal hook, small and obtuse subapical denticle, and one larger and flattened distal tooth associated with pilus dentilis; movable digit armed with relatively thin terminal hook and one subdistal tooth.
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</paragraph>
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<paragraph id="22057ECA1F41287451686488B9A26889" pageId="5" pageNumber="76">
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Legs. Leg setation normal for genus (
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<bibRefCitation id="9037F3165E59E70C91620832A506C21F" author="Masan, P" journalOrPublisher="NOI Press, Bratislava" pageId="24" pageNumber="95" title="A Review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European species (Acari: Mesostigmata: Eviphidoidea)." year="2007">
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<normalizedToken id="A9E9E9C01EAFA11A28716892773664AF" originalValue="Mašán">Masan</normalizedToken>
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2007
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</bibRefCitation>
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). Femur II with a small spine-like process on ventral distal surface, process associated with a seta. Tarsus II with two spur-like distal setae pl1 and pl2 (Figure 7).
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</paragraph>
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<paragraph id="BB3B29F9F4B7A3A63F7A1981C2A08CB6" pageId="5" pageNumber="76">
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Male. Idiosoma (Figure 8). Dorsal shield 810
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<normalizedToken id="645B0A9B96483AAC2A1ABDC492A97490" originalValue="μm">μm</normalizedToken>
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long and 492
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<normalizedToken id="7E8C5279F4001A551F1F2E55C3D5B1D7" originalValue="μm">μm</normalizedToken>
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wide, suboval (length/width 1.65). Sternal, genitiventral, peritrematal, metapodal, and anal plates are fused together to form an entire holoventral shield bearing nine pairs of setae (excluding three circum-anal setae); the shield irregularly reticulate on surface. Dorsolateral and ventral soft integument with eight pairs of setae (see diagnosis). Dorsal and ventral chaetotaxy and other characters as in female.
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</paragraph>
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<caption id="FBA9D25FF9F8A08B9D700CEC0D835EF0" pageId="5" pageNumber="76">
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<paragraph id="C2708F34F39918334F9E56EAD5C6C70D" pageId="5" pageNumber="76">
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Figures 8-12.
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<taxonomicName id="085AECCB8D48E71B819E004A1E0D50D1" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps (Pachylaelaps) pyrenaicus" order="Mesostigmata" pageId="5" pageNumber="76" phylum="Arthropoda" rank="species" species="pyrenaicus" subGenus="Pachylaelaps">Pachylaelaps (Pachylaelaps) pyrenaicus</taxonomicName>
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, male. 8 Ventral idiosoma 9 Chelicera, ventrolateral view 10 Palptibial projections 11 Projections on medial segments of leg II, ventral view 12 Tarsus II, lateral view.
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</paragraph>
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</caption>
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<caption id="1ED1B1256C21380B6B996DBBD41397C4" pageId="5" pageNumber="76">
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<paragraph id="E2DC40E007BE73DF324BA1DD0EFAE684" pageId="5" pageNumber="76">
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Figures 13-19.
|
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<taxonomicName id="9C05AB5F4B10C30E9950E40DDCC3835C" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps (Pachylaelaps) pyrenaicus" order="Mesostigmata" pageId="5" pageNumber="76" phylum="Arthropoda" rank="species" species="pyrenaicus" subGenus="Pachylaelaps">Pachylaelaps (Pachylaelaps) pyrenaicus</taxonomicName>
|
||
. 13-16 Sperm induction structures, female, variant forms 17-18 Palptibial projections, male, variant forms 19 Chelicera, female, lateral view. Not scaled.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="23DA3963AB5F6E01B929E9B8450F0B8B" pageId="6" pageNumber="77">
|
||
<pageBreakToken id="62FAF4A0AD53332D5AD0114C5E146234" pageId="6" pageNumber="77" start="start">Gnathosomal</pageBreakToken>
|
||
structures (Figures 9, 10, 17, 18). Cheliceral spermatodactyl elongated, ensiform, 162
|
||
<normalizedToken id="7D058418970142E79AC49403CB2A5995" originalValue="μm">μm</normalizedToken>
|
||
long (about 1.5 times as long as movable digit of chelicera), slightly widened in basal section, progressively tapering toward tip and slightly undulate medially; sperm ductus well defined (Figure 9). Palptibiae normal, not thickened (when compared with other palp segments), each bearing a pair of well-developed petal-like projections on proximal ventral surface (see Figures 17 and 18); outer petal markedly larger than inner one, and longer than cross-sectional radius of palptibia.
|
||
</paragraph>
|
||
<paragraph id="5083CF3C35E1BC12F0AB2CE41F279C2A" pageId="6" pageNumber="77">Legs. Medial segments of legs II spurred on their distal ventral surface: femur with one robust spur, genu and tibia each with a small knob-like spur, as in Figure 11. Femoral spur broadened basally, produced into obtuse and rounded apex; associated axillar seta pv1 inserted in a small tubercle (Figure 11). Terminal part of tarsus II with only one spur-like distal seta pl1; seta pl2 needle-shaped (Figure 12).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EB8236CF7E6D7E615F46385F4788D9E4" pageId="6" pageNumber="77" type="etymology">
|
||
<paragraph id="35502DD0CA3FD3AFF538D5263ED200C3" pageId="6" pageNumber="77">Etymology.</paragraph>
|
||
<paragraph id="CDD9F216DB48CA31180B1724C28D6751" pageId="6" pageNumber="77">The epithet of this species is derived from the Latin name "Pyrenaei Montes" and alludes to the type locality situated in the Pyrenees Mountains.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="635C8C806EB3CCD23122BB5D4FF66176" lastPageId="7" lastPageNumber="78" pageId="6" pageNumber="77" type="taxonomic notes">
|
||
<paragraph id="DEB6DAC53A36C3B2E3A67931E31C6D7E" pageId="6" pageNumber="77">Taxonomic notes.</paragraph>
|
||
<paragraph id="7532E46023288E05D4725F37E4350312" pageId="6" pageNumber="77">The new species may be distinguished from all other congeners especially by the following combination of characters: (1) in female, tubiform spermathecal structures irregular, with worm-like distal sections having some bends, folds and small bumps, and expanded base widely abutting the anterior margin of coxa IV; (2) female chelicera with flat to truncate subdistal tooth on fixed digit; (3) epistome with narrow central projection bearing a row of four to six denticles; (4) male palptibia with two well developed petal-like projections; (5) terminal part of male tarsus II with only one spur-like distal seta, pl1 (6) cheliceral spermatodactyl simple, ensiform, slightly undulate medially, without irregular convexities or projections on its margin.</paragraph>
|
||
<paragraph id="08D88BA4A85EF65A8D0C6CCA5C38450C" pageId="6" pageNumber="77">
|
||
<bibRefCitation id="8043EBF8B7C882424297D92EC9C11735" author="Masan, P" journalOrPublisher="NOI Press, Bratislava" pageId="24" pageNumber="95" title="A Review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European species (Acari: Mesostigmata: Eviphidoidea)." year="2007">
|
||
<normalizedToken id="5AB892189F09DF0CB546BF596BC0306F" originalValue="Mašán">Masan</normalizedToken>
|
||
(2007)
|
||
</bibRefCitation>
|
||
divided the European members of the subgenus
|
||
<taxonomicName id="FEFAF62F68D5BE5F89D2B9622F3758E0" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="genus">Pachylaelaps</taxonomicName>
|
||
into five clusters of species: (1) the
|
||
<taxonomicName id="CEC31BDD84AE141A395FBF1F6C1C3BC1" genus="Eviphidoidea" lsidName="Eviphidoidea bellicosus" pageId="6" pageNumber="77" rank="species" species="bellicosus">bellicosus</taxonomicName>
|
||
group (
|
||
<taxonomicName id="D4BA905ECC0AB06CA51D3F5782E636FE" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps bellicosus" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="bellicosus">Pachylaelaps bellicosus</taxonomicName>
|
||
and
|
||
<taxonomicName id="874917DD8C8CC5DA061C507D82615D6D" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps multidentatus" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="multidentatus">Pachylaelaps multidentatus</taxonomicName>
|
||
), with separate position of slit-like poroid structures gdZ1 and gdS4 on dorsal shield, multidentate cheliceral digits, spermathecal tubiform structures simple, transparent (weakly sclerotized) and relatively longer, and males apparently absent; (2) the
|
||
<taxonomicName id="06E9C89BE4E3656A85FDCD046A0A38F7" genus="Eviphidoidea" lsidName="Eviphidoidea denticulatus" pageId="6" pageNumber="77" rank="species" species="denticulatus">denticulatus</taxonomicName>
|
||
group (
|
||
<taxonomicName id="3E6570F36648D09107CCBE669E3BAD3A" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps denticulatus" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="denticulatus">Pachylaelaps denticulatus</taxonomicName>
|
||
only), possessing separate position of slit-like poroid structures gdZ1 and gdS4, three projections on male palptibia, one spur-like distal seta on tarsus II in male, and bidentate cheliceral digits; (3) the
|
||
<taxonomicName id="8512DD4F356476432AC0C31AAA895800" genus="Eviphidoidea" lsidName="Eviphidoidea ensifer" pageId="6" pageNumber="77" rank="species" species="ensifer">ensifer</taxonomicName>
|
||
group (
|
||
<taxonomicName id="0DDC50AB39983EFCB8459FAB49D64F0F" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps armimagnus" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="armimagnus">Pachylaelaps armimagnus</taxonomicName>
|
||
,
|
||
<taxonomicName id="96AE552A12CA772E5B838E34A1127B55" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps carpathimagnus" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="carpathimagnus">Pachylaelaps carpathimagnus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A75F489A620BBF06B2CCBBB09EC40CBE" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps ensifer" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="ensifer">Pachylaelaps ensifer</taxonomicName>
|
||
,
|
||
<taxonomicName id="B3C330924C26BD6DEB58FCE9CDDC0D3C" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps troglophilus" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="troglophilus">Pachylaelaps troglophilus</taxonomicName>
|
||
and
|
||
<taxonomicName id="0D9136AFEB1F8F244A5D01DB5E41613C" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps sacculimagnus" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="sacculimagnus">Pachylaelaps sacculimagnus</taxonomicName>
|
||
), characterized by the adjacent position of slit-like poroid structures gdZ1 and gdS4, robust size of idiosoma, spermathecal tubiform structures (if detectable) elongated and weakly sclerotized, and presence of 2‒4 palptibial projections in male and two spur-like distal setae on tarsus II in both adults; (4) the
|
||
<taxonomicName id="13703FA18808B90D735D346EE44606FF" genus="Eviphidoidea" lsidName="Eviphidoidea imitans" pageId="6" pageNumber="77" rank="species" species="imitans">imitans</taxonomicName>
|
||
group (
|
||
<taxonomicName id="FB10C2CD2432F194AB9EB1916838932D" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps imitans" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="imitans">Pachylaelaps imitans</taxonomicName>
|
||
,
|
||
<taxonomicName id="A04037EA7562ABD0962C8F49A1CF18EA" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps insularis" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="insularis">Pachylaelaps insularis</taxonomicName>
|
||
,
|
||
<taxonomicName id="D65B37A54511705A0879BA17980333F6" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps resinae" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="resinae">Pachylaelaps resinae</taxonomicName>
|
||
and
|
||
<taxonomicName id="0EF4FDF68DEF98379E3CC8D1B5FF2B03" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps terreus" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="terreus">Pachylaelaps terreus</taxonomicName>
|
||
), with adjacent slit-like poroid structures gdZ1 and gdS4 having their openings in a common infundibulum, spermathecal tubiform structures short, conical to cylindrical and strongly sclerotized, two palptibial projections in male, and small lobe-like convexity on ventral margin of cheliceral spermatodactyl; (5) the
|
||
<taxonomicName id="E2CFC4D46C14F7CDBFD2E6ACB7938516" genus="Eviphidoidea" lsidName="Eviphidoidea pectinifer" pageId="6" pageNumber="77" rank="species" species="pectinifer">pectinifer</taxonomicName>
|
||
group (
|
||
<taxonomicName id="5F7D3C4F8C0A1C48EE9B7BA2A8C27F32" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps littoralis" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="littoralis">Pachylaelaps littoralis</taxonomicName>
|
||
and
|
||
<taxonomicName id="9584137950978E0DA7529B125C61D94F" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps pectinifer" order="Mesostigmata" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="pectinifer">Pachylaelaps pectinifer</taxonomicName>
|
||
), characterized by the adjacent position of slit-like poroid structures gdZ1 and gdS4, Y-shaped spermathecal tubiform structures, absence of palptibial projections in male, and presence of two spur-like distal setae on tarsus II in both adults.
|
||
</paragraph>
|
||
<paragraph id="2B9E91C107544935D095F806A223D4AA" pageId="7" pageNumber="78">
|
||
<pageBreakToken id="E418E8301181354E338DCAEDB48E4415" pageId="7" pageNumber="78" start="start">In</pageBreakToken>
|
||
this classification scheme,
|
||
<taxonomicName id="637EC44549A04C047384AD2A2D8B7239" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps (Pachylaelaps) pyrenaicus" order="Mesostigmata" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="pyrenaicus" subGenus="Pachylaelaps">Pachylaelaps (Pachylaelaps) pyrenaicus</taxonomicName>
|
||
should be considered as a species with a separate position among the all above mentioned species groups because it possesses a unique combination of main diagnostic characters. Some morphological characters of
|
||
<taxonomicName id="EFBDC1926241E3DED67F302933B72095" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps (Pachylaelaps) pyrenaicus" order="Mesostigmata" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="pyrenaicus" subGenus="Pachylaelaps">Pachylaelaps (Pachylaelaps) pyrenaicus</taxonomicName>
|
||
are not consistent with those found typically in the individual species groups. The adjacent position of slit-like poroid structures gdZ1 and gdS4 on dorsal shield and unidentate cheliceral digits in the new species are in contradiction with the definition of the
|
||
<taxonomicName id="A64E3111AA02C3E218B9662A696ACC33" genus="Eviphidoidea" lsidName="Eviphidoidea bellicosus" pageId="7" pageNumber="78" rank="species" species="bellicosus">bellicosus</taxonomicName>
|
||
and
|
||
<taxonomicName id="C773BB6F6ED882AA76FE0372D175EBBC" genus="Eviphidoidea" lsidName="Eviphidoidea denticulatus" pageId="7" pageNumber="78" rank="species" species="denticulatus">denticulatus</taxonomicName>
|
||
groups. The male palptibia has two petal-like projections in
|
||
<taxonomicName id="1CE132F5002571377F4A66F4EFB713AB" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps (Pachylaelaps) pyrenaicus" order="Mesostigmata" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="pyrenaicus" subGenus="Pachylaelaps">Pachylaelaps (Pachylaelaps) pyrenaicus</taxonomicName>
|
||
, where the
|
||
<taxonomicName id="C31BD72360E505595B85A984137009ED" genus="Eviphidoidea" lsidName="Eviphidoidea pectinifer" pageId="7" pageNumber="78" rank="species" species="pectinifer">pectinifer</taxonomicName>
|
||
group species does not have these structures developed. In the robust species of the
|
||
<taxonomicName id="DE2590DDF4EDA77E028F222E17337012" genus="Eviphidoidea" lsidName="Eviphidoidea ensifer" pageId="7" pageNumber="78" rank="species" species="ensifer">ensifer</taxonomicName>
|
||
group, tarsus II has two spur-like distal setae in adults, but this character is found in the smaller new species only in females. In addition, in
|
||
<taxonomicName id="90D6156B37B66A52413AC44EA768BA86" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pachylaelaps (Pachylaelaps) pyrenaicus" order="Mesostigmata" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="pyrenaicus" subGenus="Pachylaelaps">Pachylaelaps (Pachylaelaps) pyrenaicus</taxonomicName>
|
||
, the tubular structures of sperm induction system have a distinctive form which is not known in the other species of the genus, but is especially different from the members of the
|
||
<taxonomicName id="F45D05CDF182D7027CF56D7D9EB2EDD9" genus="Eviphidoidea" lsidName="Eviphidoidea imitans" pageId="7" pageNumber="78" rank="species" species="imitans">imitans</taxonomicName>
|
||
group that are characterized by short, conical to cylindrical, and strongly sclerotized spermathecal tubes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |