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<mods:title id="C6E17AEDB2DFBBC9EA1F9D8DC21B9BE3">Systematic revision of Johnmannia Irwin and Lyneborg (Diptera: Therevidae): Atypical metallic stiletto flies from Australian mesic habitats</mods:title>
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<taxonomicName id="B9FB4D12FFBE84271526D1410EF9FA3D" ID-CoL="56Q2" ID-ENA="1562684" authority="Irwin &amp; Lyneborg, 1989" authorityName="Irwin &amp; Lyneborg" authorityYear="1989" box="[264,729,1398,1424]" class="Insecta" family="Therevidae" genus="Johnmannia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
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<emphasis id="4C8FEA83FFBE84271526D1410D82FA3D" bold="true" box="[264,418,1398,1424]" italics="true" pageId="5" pageNumber="6">Johnmannia</emphasis>
<bibRefCitation id="1A6A4B60FFBE84271584D1410EF9FA3D" author="Irwin" box="[426,729,1398,1424]" lastPageId="5" lastPageNumber="23" pageId="5" pageNumber="22" refString="Irwin, M. E. &amp; Lyneborg, L. (1989) Family Therevidae. In: Evenhuis, N. L. (Ed) Catalog of the Diptera of the Australasian and Oceanian Regions, Bishop Museum Press and E. J. Brill, Honolulu, pp. 1155." type="book chapter" year="1989">Irwin &amp; Lyneborg, 1989</bibRefCitation>
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(
<figureCitation id="E6C02A14FFBE8427153ED1AE0D58FA1E" box="[272,376,1433,1459]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="9.[264,367,1411,1435]" captionStartId-1="4.[264,367,1827,1851]" captionStartId-2="12.[264,367,1702,1726]" captionStartId-3="13.[264,367,1764,1788]" captionStartId-4="17.[264,367,1753,1777]" captionStartId-5="19.[264,367,1730,1754]" captionTargetBox-0="[296,1276,703,1381]" captionTargetBox-1="[314,1263,300,1787]" captionTargetBox-2="[275,1323,287,1688]" captionTargetBox-3="[264,1323,411,1723]" captionTargetBox-4="[302,1284,290,1733]" captionTargetBox-5="[264,1323,456,1713]" captionTargetId-0="figure@9.[264,1305,689,1400]" captionTargetId-1="figure@4.[264,1300,258,1820]" captionTargetId-2="figure@12.[264,1323,287,1691]" captionTargetId-3="figure@13.[264,1323,411,1738]" captionTargetId-4="figure@17.[302,1285,290,1750]" captionTargetId-5="figure@19.[264,1323,456,1713]" captionTargetPageId-0="9" captionTargetPageId-1="4" captionTargetPageId-2="12" captionTargetPageId-3="13" captionTargetPageId-4="17" captionTargetPageId-5="19" captionText-0="FIGURE 1. Habitus of Johnmannia powerae sp. nov. Ψ MEI 0 25305, lateral." captionText-1="FIGURE 2. One of 6 most parsimonious trees with unambiguous character and state changes mapped on branches, and Bremer supports given above nodes." captionText-2="FIGURE 3. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 dorsal, A E; Ψ ANIC 29 0 0 3454 frontal, F G. (A) whole body, (B) occiput and frontal pruinescence, (C) thoracic pruinescence, (D) wing pattern and venation, (E) abdomen, (F) frontal pruinescence, (G) antennae." captionText-3="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." captionText-4="FIGURE 6. Johnmannia tasmanica Ψ UQIC 40944 lateral A C, body H J; Ψ ANIC 29 0 0 3419 genitalia, D G. (A) body, (B) head and thorax, (C) antenna, (D) furca, (E) accessory glands, SD, SPSD, common SPSD, (F) S 8, (G) T 8 with apodeme, (H) head frontal, (I) wing pattern and venation, with terminology of venation and cells, (J) mid femoral pruinescence." captionText-5="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri-0="https://zenodo.org/record/170864/files/figure.png" httpUri-1="https://zenodo.org/record/170863/files/figure.png" httpUri-2="https://zenodo.org/record/170865/files/figure.png" httpUri-3="https://zenodo.org/record/170866/files/figure.png" httpUri-4="https://zenodo.org/record/170867/files/figure.png" httpUri-5="https://zenodo.org/record/170868/files/figure.png" pageId="5" pageNumber="6">Figs 17</figureCitation>
)
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<taxonomicName id="B9FB4D12FFBE84271526D1DD0E74F9AD" authority="Paramonov, 1950: 525" authorityName="Paramonov" authorityPageNumber="525" authorityYear="1950" box="[264,596,1513,1537]" class="Marchantiopsida" family="Aytoniaceae" genus="Mannia" kingdom="Plantae" order="Marchantiales" pageId="5" pageNumber="6" phylum="Bryophyta" rank="genus">
<emphasis id="4C8FEA83FFBE84271526D1DD0D7BF9AC" box="[264,347,1514,1537]" italics="true" pageId="5" pageNumber="6">Mannia</emphasis>
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: 525
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;
<treatmentCitation id="FF5A1080FFBE84271671D1DE0F2AF9AD" author="Liepa" box="[607,778,1513,1536]" page="12" pageId="5" pageNumber="6" year="1969">
<bibRefCitation id="1A6A4B60FFBE84271671D1DE0EC2F9AD" author="Liepa" box="[607,738,1513,1536]" pageId="5" pageNumber="23" refString="Liepa, Z. R. (1969) Lists of the Scientific Works and Described Species of the Late Dr S. J. Paramonov, with locations of types. The Journal of the Entomological Society of Australia (NSW), 5, 3 - 22." type="journal article" year="1969">Liepa, 1969</bibRefCitation>
: 12
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. [Preoccupied
<treatmentCitation id="FF5A1080FFBE84271789D1DE08A6F9AD" author="Davidson" box="[935,1158,1513,1536]" page="156" pageId="5" pageNumber="6" year="1874">
<bibRefCitation id="1A6A4B60FFBE84271789D1DE0874F9AD" author="Davidson" box="[935,1108,1513,1536]" pageId="5" pageNumber="22" refString="Davidson, T. (1874) On the Tertiary Brachiopoda of Belgium. Geological Magazine, 1, 150 - 159." type="journal article" year="1874">Davidson, 1874</bibRefCitation>
: 156
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(Fossil Brachiopoda);
<treatmentCitation id="FF5A1080FFBE842715BED23E0E63F98D" author="Prout" box="[400,579,1545,1568]" page="382" pageId="5" pageNumber="6" year="1915">
<bibRefCitation id="1A6A4B60FFBE842715BED23E0E30F98D" author="Prout" box="[400,528,1545,1568]" pageId="5" pageNumber="24" refString="Prout, L. B. (1915) Die spannerartigen Nachtfalter. In: Seitz, A. (Ed) Die Gross-Schmetterlinge der Erde. Eine systematische Bearbeitung der bis jetzt bekannten Gross-Schmetterlinge. I. Abteilung: Die Gross-Schmetterlinge des palaarktischen Faunengebietes., vol. 4, Verlag, Stuttgart, pp. I - V, 1 - 479, pls 1 - 25." type="journal article" year="1915">Prout, 1915</bibRefCitation>
: 382
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(Lepidoptera)].
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</paragraph>
<paragraph id="7E443691FFBE84271526D21D0932F9ED" blockId="5.[264,1323,1513,1601]" box="[264,1298,1577,1601]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="5EEB11BFFFBE84271526D21D0932F9ED" box="[264,1298,1577,1601]" pageId="5" pageNumber="6">
<taxonomicName id="B9FB4D12FFBE84271526D21D0EEDF9ED" ID-CoL="56Q2" authority="Irwin &amp; Lyneborg, 1989: 357" authorityName="Irwin &amp; Lyneborg" authorityPageNumber="357" authorityYear="1989" box="[264,717,1577,1601]" class="Insecta" family="Therevidae" genus="Johnmannia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="4C8FEA83FFBE84271526D21D0DACF9EC" box="[264,396,1578,1601]" italics="true" pageId="5" pageNumber="6">Johnmannia</emphasis>
<treatmentCitation id="FF5A1080FFBE842715BDD21E0EEDF9ED" author="Irwin" box="[403,717,1577,1600]" page="357" pageId="5" pageNumber="6" year="1989">
<bibRefCitation id="1A6A4B60FFBE842715BDD21E0EBAF9ED" author="Irwin" box="[403,666,1577,1600]" lastPageId="5" lastPageNumber="23" pageId="5" pageNumber="22" refString="Irwin, M. E. &amp; Lyneborg, L. (1989) Family Therevidae. In: Evenhuis, N. L. (Ed) Catalog of the Diptera of the Australasian and Oceanian Regions, Bishop Museum Press and E. J. Brill, Honolulu, pp. 1155." type="book chapter" year="1989">Irwin &amp; Lyneborg, 1989</bibRefCitation>
: 357
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</taxonomicName>
[new name for
<taxonomicName id="B9FB4D12FFBE8427175BD21D0FE8F9EC" box="[885,968,1578,1601]" class="Marchantiopsida" family="Aytoniaceae" genus="Mannia" kingdom="Plantae" order="Marchantiales" pageId="5" pageNumber="6" phylum="Bryophyta" rank="genus">
<emphasis id="4C8FEA83FFBE8427175BD21D0FE8F9EC" box="[885,968,1578,1601]" italics="true" pageId="5" pageNumber="6">Mannia</emphasis>
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];
<treatmentCitation id="FF5A1080FFBE842717F1D21E092EF9ED" author="Winterton" box="[991,1294,1577,1601]" page="276" pageId="5" pageNumber="6" year="1999">
<bibRefCitation id="1A6A4B60FFBE842717F1D21E08F9F9ED" author="Winterton" box="[991,1241,1577,1601]" pageId="5" pageNumber="24" refString="Winterton, S. L., Merritt, D. J., O'Toole, A., Yeates, D. K. &amp; Irwin, M. E. (1999 c) Morphology and histology of the spermathecal sac, a novel structure in the female reproductive system of Therevidae (Diptera: Asiloidea). International Journal of Insect Morphology and Embryology, 28, 273 - 279." type="journal article" year="1999" yearSuffix="c">
Winterton
<emphasis id="4C8FEA83FFBE8427107FD21D08A0F9EC" box="[1105,1152,1578,1601]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
., 1999c
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: 276
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.
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</paragraph>
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<emphasis id="4C8FEA83FFBE84271526D2460D82F927" box="[264,418,1649,1674]" italics="true" pageId="5" pageNumber="6">
<typeStatus id="A1408833FFBE84271526D2460D1FF927" box="[264,319,1649,1674]" pageId="5" pageNumber="6">Type</typeStatus>
species.
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<taxonomicName id="B9FB4D12FFBE8427159FD2460F42F926" authority="Paramonov, 1950" authorityName="Paramonov" authorityYear="1950" box="[433,866,1649,1675]" class="Marchantiopsida" family="Aytoniaceae" genus="Mannia" kingdom="Plantae" order="Marchantiales" pageId="5" pageNumber="6" phylum="Bryophyta" rank="species" species="tasmanica">
<emphasis id="4C8FEA83FFBE8427159FD2460EABF927" box="[433,651,1649,1674]" italics="true" pageId="5" pageNumber="6">Mannia tasmanica</emphasis>
<bibRefCitation id="1A6A4B60FFBE842716BDD2460F42F926" author="Paramonov" box="[659,866,1649,1675]" pageId="5" pageNumber="24" refString="Paramonov, S. J. (1950) Notes on Australian Diptera (I - V). III A note on two Australian therevids. The Annals and Magazine of Natural History, 3, 525 - 529." type="journal article" year="1950">Paramonov, 1950</bibRefCitation>
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, by original designation
</paragraph>
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<subSubSection id="36E1651AFFBE84251526D2F608B7FC5B" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="7E443691FFBE84241526D2F60E9AFCC6" blockId="5.[264,1321,1729,1875]" lastBlockId="6.[264,1324,284,1898]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="4C8FEA83FFBE84271526D2F60DA7F977" box="[264,391,1729,1754]" italics="true" pageId="5" pageNumber="6">Diagnosis.</emphasis>
Colouration black (
<figureCitation id="E6C02A14FFBE842716AED2F60EEAF976" box="[640,714,1729,1755]" captionStart="FIGURE 6" captionStartId="17.[264,367,1753,1777]" captionTargetBox="[302,1284,290,1733]" captionTargetId="figure@17.[302,1285,290,1750]" captionTargetPageId="17" captionText="FIGURE 6. Johnmannia tasmanica Ψ UQIC 40944 lateral A C, body H J; Ψ ANIC 29 0 0 3419 genitalia, D G. (A) body, (B) head and thorax, (C) antenna, (D) furca, (E) accessory glands, SD, SPSD, common SPSD, (F) S 8, (G) T 8 with apodeme, (H) head frontal, (I) wing pattern and venation, with terminology of venation and cells, (J) mid femoral pruinescence." httpUri="https://zenodo.org/record/170867/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation>
A) with metallic blue or green reflectivity; scape elongate, with uniform long dark setae (
<figureCitation id="E6C02A14FFBE842716F1D2DE0F03F8AE" box="[735,803,1769,1795]" captionStart="FIGURE 3" captionStartId="12.[264,367,1702,1726]" captionTargetBox="[275,1323,287,1688]" captionTargetId="figure@12.[264,1323,287,1691]" captionTargetPageId="12" captionText="FIGURE 3. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 dorsal, A E; Ψ ANIC 29 0 0 3454 frontal, F G. (A) whole body, (B) occiput and frontal pruinescence, (C) thoracic pruinescence, (D) wing pattern and venation, (E) abdomen, (F) frontal pruinescence, (G) antennae." httpUri="https://zenodo.org/record/170865/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
FG); face with pruinescence laterally from antennae along compound eye, narrowed medially (
<figureCitation id="E6C02A14FFBE84271741D3260F96F886" box="[879,950,1809,1835]" captionStart="FIGURE 3" captionStartId="12.[264,367,1702,1726]" captionTargetBox="[275,1323,287,1688]" captionTargetId="figure@12.[264,1323,287,1691]" captionTargetPageId="12" captionText="FIGURE 3. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 dorsal, A E; Ψ ANIC 29 0 0 3454 frontal, F G. (A) whole body, (B) occiput and frontal pruinescence, (C) thoracic pruinescence, (D) wing pattern and venation, (E) abdomen, (F) frontal pruinescence, (G) antennae." httpUri="https://zenodo.org/record/170865/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
F), covering anteriorly directed flange of gena at base of compound eye (
<figureCitation id="E6C02A14FFBE842716DCD30E0F1BF8FE" box="[754,827,1849,1875]" captionStart="FIGURE 4" captionStartId="13.[264,367,1764,1788]" captionTargetBox="[264,1323,411,1723]" captionTargetId="figure@13.[264,1323,411,1738]" captionTargetPageId="13" captionText="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." httpUri="https://zenodo.org/record/170866/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
B); occiput with gold pruinescence, indistinct row of dark postocular setae almost length of ocellar triangle from compound eye; 4 6 short black post­spiracular setae (Fig. 5B) (metepisternal (
<bibRefCitation id="1A6A4B60FFBD842417F9D573084BFEF3" author="Yeates" box="[983,1131,324,350]" pageId="6" pageNumber="24" refString="Yeates, D. (1994) Cladistics and classification of the Bombyliidae (Diptera: Asiloidea). Bulletin of the American Museum of Natural History, 219, 1 - 191." type="journal article" year="1994">Yeates 1994</bibRefCitation>
) or hypopleural pile
<bibRefCitation id="1A6A4B60FFBD84241513D55B0E2DFE2B" author="Lyneborg" box="[317,525,364,390]" pageId="6" pageNumber="23" refString="Lyneborg, L. (2001) The Australian stiletto-flies of the Anabarhynchus genus-group Therevidae (Diptera: Therevidae). Entomonograph, Apollo Books, Steenstrup, Denmark, 256 pp." type="book" year="2001">(Lyneborg 2001))</bibRefCitation>
; wing cell m3 open (Fig. 5C); seta present antero­ventrally subapically on hind femur (
<figureCitation id="E6C02A14FFBD842415D0D5AE0E63FE1E" box="[510,579,409,435]" captionStart="FIGURE 1" captionStartId="9.[264,367,1411,1435]" captionTargetBox="[296,1276,703,1381]" captionTargetId="figure@9.[264,1305,689,1400]" captionTargetPageId="9" captionText="FIGURE 1. Habitus of Johnmannia powerae sp. nov. Ψ MEI 0 25305, lateral." httpUri="https://zenodo.org/record/170864/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
); without velutum patches on fore and hind femora and ventral surface of gonocoxites; wing with black infuscation distally and medially (Fig. 5C) or entire blade (
<figureCitation id="E6C02A14FFBD8424158AD5DE0DCBFDAE" box="[420,491,489,515]" captionStart="FIGURE 7" captionStartId="19.[264,367,1730,1754]" captionTargetBox="[264,1323,456,1713]" captionTargetId="figure@19.[264,1323,456,1713]" captionTargetPageId="19" captionText="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri="https://zenodo.org/record/170868/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
F), without orange infuscation; two scutellar setae (
<figureCitation id="E6C02A14FFBD84241060D5DE08B5FDAE" box="[1102,1173,489,515]" captionStart="FIGURE 3" captionStartId="12.[264,367,1702,1726]" captionTargetBox="[275,1323,287,1688]" captionTargetId="figure@12.[264,1323,287,1691]" captionTargetPageId="12" captionText="FIGURE 3. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 dorsal, A E; Ψ ANIC 29 0 0 3454 frontal, F G. (A) whole body, (B) occiput and frontal pruinescence, (C) thoracic pruinescence, (D) wing pattern and venation, (E) abdomen, (F) frontal pruinescence, (G) antennae." httpUri="https://zenodo.org/record/170865/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
C) (rarely 4, with outer setae weak); abdomen strongly recurved apically (
<figureCitation id="E6C02A14FFBD842417FFD6260834FD86" box="[977,1044,529,555]" captionStart="FIGURE 1" captionStartId="9.[264,367,1411,1435]" captionTargetBox="[296,1276,703,1381]" captionTargetId="figure@9.[264,1305,689,1400]" captionTargetPageId="9" captionText="FIGURE 1. Habitus of Johnmannia powerae sp. nov. Ψ MEI 0 25305, lateral." httpUri="https://zenodo.org/record/170864/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
), appears globular dorsally (
<figureCitation id="E6C02A14FFBD8424157CD60E0DB9FDFE" box="[338,409,569,595]" captionStart="FIGURE 3" captionStartId="12.[264,367,1702,1726]" captionTargetBox="[275,1323,287,1688]" captionTargetId="figure@12.[264,1323,287,1691]" captionTargetPageId="12" captionText="FIGURE 3. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 dorsal, A E; Ψ ANIC 29 0 0 3454 frontal, F G. (A) whole body, (B) occiput and frontal pruinescence, (C) thoracic pruinescence, (D) wing pattern and venation, (E) abdomen, (F) frontal pruinescence, (G) antennae." httpUri="https://zenodo.org/record/170865/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
E); spermathecal sac arrangement trilobate with each spermathecal sac with narrow tube to outer lobe (
<figureCitation id="E6C02A14FFBD84241669D6560EAFFDD6" box="[583,655,609,635]" captionStart="FIGURE 4" captionStartId="13.[264,367,1764,1788]" captionTargetBox="[264,1323,411,1723]" captionTargetId="figure@13.[264,1323,411,1738]" captionTargetPageId="13" captionText="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." httpUri="https://zenodo.org/record/170866/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
C); spermathecal duct broadening basally before joining individual spermathecal sac duct (
<figureCitation id="E6C02A14FFBD8424168DD6BE0ECDFD0E" box="[675,749,649,675]" captionStart="FIGURE 4" captionStartId="13.[264,367,1764,1788]" captionTargetBox="[264,1323,411,1723]" captionTargetId="figure@13.[264,1323,411,1738]" captionTargetPageId="13" captionText="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." httpUri="https://zenodo.org/record/170866/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
E); subepandrial sclerite well developed sclerotised plate (
<figureCitation id="E6C02A14FFBD842415B9D6860DC0FD66" box="[407,480,689,715]" captionStart="FIGURE 7" captionStartId="19.[264,367,1730,1754]" captionTargetBox="[264,1323,456,1713]" captionTargetId="figure@19.[264,1323,456,1713]" captionTargetPageId="19" captionText="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri="https://zenodo.org/record/170868/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
KM); outer gonocoxal process well developed, pointed (Fig. 5GI); inner gonocoxal process long, narrow, apically acuminate (Fig. 5H); gonostylus longer than inner gonocoxal process, dorsal recurved triangulate hook 1/3 length of gonostylus from apex (
<figureCitation id="E6C02A14FFBD842415A1D71E0DF4FCEE" box="[399,468,809,835]" captionStart="FIGURE 7" captionStartId="19.[264,367,1730,1754]" captionTargetBox="[264,1323,456,1713]" captionTargetId="figure@19.[264,1323,456,1713]" captionTargetPageId="19" captionText="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri="https://zenodo.org/record/170868/files/figure.png" pageId="6" pageNumber="7">
Fig.
<date id="0A451051FFBD842415EBD71E0DF4FCEE" box="[453,468,809,835]" pageId="6" pageNumber="7">7</date>
</figureCitation>
IJ); hypandrium triangular, joined laterally to gonocoxites (Fig.
<date id="0A451051FFBD842410E4D71E08C5FCEE" box="[1226,1253,809,835]" pageId="6" pageNumber="7">5I</date>
); distiphallus directed ventrally (Fig. 5L).
</paragraph>
<paragraph id="7E443691FFBD84241516D74E0908FA8E" blockId="6.[264,1324,284,1898]" pageId="6" pageNumber="7">
<emphasis id="4C8FEA83FFBD84241516D74E0E91FC3F" box="[312,689,889,914]" italics="true" pageId="6" pageNumber="7">Generic Redescription. Female</emphasis>
.
<emphasis id="4C8FEA83FFBD842416EBD74E0F25FC3F" box="[709,773,889,914]" italics="true" pageId="6" pageNumber="7">Head</emphasis>
shape laterally oval (
<figureCitation id="E6C02A14FFBD8424103AD74E087FFC3E" box="[1044,1119,889,915]" captionStart="FIGURE 6" captionStartId="17.[264,367,1753,1777]" captionTargetBox="[302,1284,290,1733]" captionTargetId="figure@17.[302,1285,290,1750]" captionTargetPageId="17" captionText="FIGURE 6. Johnmannia tasmanica Ψ UQIC 40944 lateral A C, body H J; Ψ ANIC 29 0 0 3419 genitalia, D G. (A) body, (B) head and thorax, (C) antenna, (D) furca, (E) accessory glands, SD, SPSD, common SPSD, (F) S 8, (G) T 8 with apodeme, (H) head frontal, (I) wing pattern and venation, with terminology of venation and cells, (J) mid femoral pruinescence." httpUri="https://zenodo.org/record/170867/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
B). Frons glossy black; upper frons width greater than width of ocellar triangle (
<figureCitation id="E6C02A14FFBD842417D7D7960860FC16" box="[1017,1088,929,955]" captionStart="FIGURE 3" captionStartId="12.[264,367,1702,1726]" captionTargetBox="[275,1323,287,1688]" captionTargetId="figure@12.[264,1323,287,1691]" captionTargetPageId="12" captionText="FIGURE 3. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 dorsal, A E; Ψ ANIC 29 0 0 3454 frontal, F G. (A) whole body, (B) occiput and frontal pruinescence, (C) thoracic pruinescence, (D) wing pattern and venation, (E) abdomen, (F) frontal pruinescence, (G) antennae." httpUri="https://zenodo.org/record/170865/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
C), short dark setae on ocellar triangle, deep broad medial concavity; lower frons rounded dark glossy callus or antennal tubercule (
<figureCitation id="E6C02A14FFBD842415DCD7C60E1BFBA6" box="[498,571,1009,1035]" captionStart="FIGURE 4" captionStartId="13.[264,367,1764,1788]" captionTargetBox="[264,1323,411,1723]" captionTargetId="figure@13.[264,1323,411,1738]" captionTargetPageId="13" captionText="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." httpUri="https://zenodo.org/record/170866/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
B); short, dark, setae laterally above antennae; face with dense, silver pruinescence. Occiput rounded (
<figureCitation id="E6C02A14FFBD842416F9D02E0F00FB9E" box="[727,800,1049,1075]" captionStart="FIGURE 3" captionStartId="12.[264,367,1702,1726]" captionTargetBox="[275,1323,287,1688]" captionTargetId="figure@12.[264,1323,287,1691]" captionTargetPageId="12" captionText="FIGURE 3. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 dorsal, A E; Ψ ANIC 29 0 0 3454 frontal, F G. (A) whole body, (B) occiput and frontal pruinescence, (C) thoracic pruinescence, (D) wing pattern and venation, (E) abdomen, (F) frontal pruinescence, (G) antennae." httpUri="https://zenodo.org/record/170865/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
B), indistinct rows of dark postocular setae distant from compound eye; dense, gold pruinescence at least ventrally; very long fine dark setae ventrally. Gena covered with fine long, dark setae, overlain with dense, gold pruinescence; distinct protrusion of gena at base of compound eye (
<figureCitation id="E6C02A14FFBD84241039D0A6087EFB06" box="[1047,1118,1169,1195]" captionStart="FIGURE 4" captionStartId="13.[264,367,1764,1788]" captionTargetBox="[264,1323,411,1723]" captionTargetId="figure@13.[264,1323,411,1738]" captionTargetPageId="13" captionText="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." httpUri="https://zenodo.org/record/170866/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
B). Palp two­segmented, apical segment spatulate, fine long dark setae basally, less dense medially (
<figureCitation id="E6C02A14FFBD842410D7D08E0D39FB56" captionStart="FIGURE 4" captionStartId="13.[264,367,1764,1788]" captionTargetBox="[264,1323,411,1723]" captionTargetId="figure@13.[264,1323,411,1738]" captionTargetPageId="13" captionText="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." httpUri="https://zenodo.org/record/170866/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
B); labellum dark, long dark setae medially, short dark setae apically. Antenna slightly longer than head (
<figureCitation id="E6C02A14FFBD842415F7D13E0E3CFA8E" box="[473,540,1289,1315]" captionStart="FIGURE 3" captionStartId="12.[264,367,1702,1726]" captionTargetBox="[275,1323,287,1688]" captionTargetId="figure@12.[264,1323,287,1691]" captionTargetPageId="12" captionText="FIGURE 3. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 dorsal, A E; Ψ ANIC 29 0 0 3454 frontal, F G. (A) whole body, (B) occiput and frontal pruinescence, (C) thoracic pruinescence, (D) wing pattern and venation, (E) abdomen, (F) frontal pruinescence, (G) antennae." httpUri="https://zenodo.org/record/170865/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
FG); scape as long or longer than postpedicel, wider than pedicel;
</paragraph>
<paragraph id="7E443691FFBD84251526D10E0857FDA4" blockId="6.[264,1324,284,1898]" lastBlockId="7.[264,1324,284,1899]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
scape and pedicel covered with uniformly long dark setae on all surfaces; postpedicel (1st antennal flagellomere) long (postpedicel/pedicel = 4.46.3); laterally flattened, broadest sub­basally, tapering apically, indentation on lateromedial surface; basal stylomere (2nd antennal flagellomere) short broad cylinder, antennal apical stylomere (3rd antennal flagellomere) conical with pale brown, short pointed style.
<emphasis id="4C8FEA83FFBD842417FFD1E80806FA55" box="[977,1062,1503,1528]" italics="true" pageId="6" pageNumber="7">Thorax</emphasis>
: Scutum glossy dark, reflective metallic dark blue to green, numerous fine, short, dark setae; gold pruinescence anteriorly. Scutellum black, reflective, metallic dark blue to green. Pleuron glossy, dark; overlain with areas of reddish­gold pruinescence, denser ventrally (
<figureCitation id="E6C02A14FFBD84241003D2600855F9DC" box="[1069,1141,1623,1649]" captionStart="FIGURE 4" captionStartId="13.[264,367,1764,1788]" captionTargetBox="[264,1323,411,1723]" captionTargetId="figure@13.[264,1323,411,1738]" captionTargetPageId="13" captionText="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." httpUri="https://zenodo.org/record/170866/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
A); pale, short, dense setae around anterior spiracle. Scutal chaetotaxy: np 45; sa 23; pa 1; dc 12; sc 1 (2 rarely: outer finer, shorter, not equal).
<emphasis id="4C8FEA83FFBD842416CAD2900F3BF96D" box="[740,795,1703,1728]" italics="true" pageId="6" pageNumber="7">Legs</emphasis>
dark reddish­brown to black, with short dark setae; 12 large black anteroventral subapical macrosetae on hind femur; tarsomeres sometimes with orange bands, ventral surface of 4th and 5th tarsomeres lacking setae, partial loss on 3rd.
<collectingCountry id="06EC7601FFBD84241571D3130DBAF893" box="[351,410,1828,1854]" name="India" pageId="6" pageNumber="7">Hind</collectingCountry>
legs long.
<emphasis id="4C8FEA83FFBD8424160DD3120E7DF893" box="[547,605,1829,1854]" italics="true" pageId="6" pageNumber="7">Wing</emphasis>
covered with microtrichia, variably infuscated, with darker opaque triangular basal area. Triangulate darker opaque area between apex of Sc and R1, extending to join of CuA2 and A1, over m­cu crossvein and CuA2; across base of r2+3, d, m3 and cua1; apex of bm and br, indistinct posteriorly (
<figureCitation id="E6C02A14FFBC8425175FD57E0F97FECE" box="[881,951,329,355]" captionStart="FIGURE 3" captionStartId="12.[264,367,1702,1726]" captionTargetBox="[275,1323,287,1688]" captionTargetId="figure@12.[264,1323,287,1691]" captionTargetPageId="12" captionText="FIGURE 3. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 dorsal, A E; Ψ ANIC 29 0 0 3454 frontal, F G. (A) whole body, (B) occiput and frontal pruinescence, (C) thoracic pruinescence, (D) wing pattern and venation, (E) abdomen, (F) frontal pruinescence, (G) antennae." httpUri="https://zenodo.org/record/170865/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
D). Apical half of wing darker, paler posteriorly (
<figureCitation id="E6C02A14FFBC842515C6D5430E15FE23" box="[488,565,372,398]" captionStart="FIGURE 3" captionStartId="12.[264,367,1702,1726]" captionTargetBox="[275,1323,287,1688]" captionTargetId="figure@12.[264,1323,287,1691]" captionTargetPageId="12" captionText="FIGURE 3. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 dorsal, A E; Ψ ANIC 29 0 0 3454 frontal, F G. (A) whole body, (B) occiput and frontal pruinescence, (C) thoracic pruinescence, (D) wing pattern and venation, (E) abdomen, (F) frontal pruinescence, (G) antennae." httpUri="https://zenodo.org/record/170865/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
D). Halter stem and knob dark.
<emphasis id="4C8FEA83FFBC842517F8D5430865FE20" box="[982,1093,372,397]" italics="true" pageId="7" pageNumber="8">Abdomen</emphasis>
: Conical, apically recurved thus appears globular in dorsal view (
<figureCitation id="E6C02A14FFBC84251719D5AB0F5EFE1B" box="[823,894,412,438]" captionStart="FIGURE 3" captionStartId="12.[264,367,1702,1726]" captionTargetBox="[275,1323,287,1688]" captionTargetId="figure@12.[264,1323,287,1691]" captionTargetPageId="12" captionText="FIGURE 3. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 dorsal, A E; Ψ ANIC 29 0 0 3454 frontal, F G. (A) whole body, (B) occiput and frontal pruinescence, (C) thoracic pruinescence, (D) wing pattern and venation, (E) abdomen, (F) frontal pruinescence, (G) antennae." httpUri="https://zenodo.org/record/170865/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
E). Dark, glossy, reflective, metallic dark blue to green; setae sparse, fine short, T12 with longer setae laterally; reddish­gold pruinescence on T1 basally, over entire S14, densest laterally at ST sutures.
</paragraph>
<paragraph id="7E443691FFBC84251516D62E08B7FC5B" blockId="7.[264,1324,284,1899]" pageId="7" pageNumber="8">
<emphasis id="4C8FEA83FFBC84251516D62E0D86FD9F" box="[312,422,537,562]" italics="true" pageId="7" pageNumber="8">Genitalia</emphasis>
: T8 (
<figureCitation id="E6C02A14FFBC842515C7D62E0E13FD9E" box="[489,563,537,563]" captionStart="FIGURE 4" captionStartId="13.[264,367,1764,1788]" captionTargetBox="[264,1323,411,1723]" captionTargetId="figure@13.[264,1323,411,1738]" captionTargetPageId="13" captionText="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." httpUri="https://zenodo.org/record/170866/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
G) subquadrate, anterior process long narrowed medially, 0.38 length of T8; furca (
<figureCitation id="E6C02A14FFBC842515D7D6730E61FDF3" box="[505,577,580,606]" captionStart="FIGURE 6" captionStartId="17.[264,367,1753,1777]" captionTargetBox="[302,1284,290,1733]" captionTargetId="figure@17.[302,1285,290,1750]" captionTargetPageId="17" captionText="FIGURE 6. Johnmannia tasmanica Ψ UQIC 40944 lateral A C, body H J; Ψ ANIC 29 0 0 3419 genitalia, D G. (A) body, (B) head and thorax, (C) antenna, (D) furca, (E) accessory glands, SD, SPSD, common SPSD, (F) S 8, (G) T 8 with apodeme, (H) head frontal, (I) wing pattern and venation, with terminology of venation and cells, (J) mid femoral pruinescence." httpUri="https://zenodo.org/record/170867/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
D) heavily sclerotised, long (1.7 x as long as wide), narrowed posteriorly, closed; T810 joined through sclerotised band; acanthophorites with 68 broad dark brown to black A1 setae and 1015 long fine, black A2 setae; S8 (
<figureCitation id="E6C02A14FFBC8425106ED6AE08A5FD1E" box="[1088,1157,665,691]" captionStart="FIGURE 6" captionStartId="17.[264,367,1753,1777]" captionTargetBox="[302,1284,290,1733]" captionTargetId="figure@17.[302,1285,290,1750]" captionTargetPageId="17" captionText="FIGURE 6. Johnmannia tasmanica Ψ UQIC 40944 lateral A C, body H J; Ψ ANIC 29 0 0 3419 genitalia, D G. (A) body, (B) head and thorax, (C) antenna, (D) furca, (E) accessory glands, SD, SPSD, common SPSD, (F) S 8, (G) T 8 with apodeme, (H) head frontal, (I) wing pattern and venation, with terminology of venation and cells, (J) mid femoral pruinescence." httpUri="https://zenodo.org/record/170867/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
F) with posterior margin indented medially, forming blunt rounded lobes; three spermathecae (
<figureCitation id="E6C02A14FFBC842510EED6F30926FD73" box="[1216,1286,708,734]" captionStart="FIGURE 4" captionStartId="13.[264,367,1764,1788]" captionTargetBox="[264,1323,411,1723]" captionTargetId="figure@13.[264,1323,411,1738]" captionTargetPageId="13" captionText="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." httpUri="https://zenodo.org/record/170866/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
C); spermathecal duct threadlike, twice length of furca to sclerotised cuticular valve, broadening basally (
<figureCitation id="E6C02A14FFBC842515B1D7230DC9FC83" box="[415,489,788,814]" captionStart="FIGURE 4" captionStartId="13.[264,367,1764,1788]" captionTargetBox="[264,1323,411,1723]" captionTargetId="figure@13.[264,1323,411,1738]" captionTargetPageId="13" captionText="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." httpUri="https://zenodo.org/record/170866/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
E) for the length of furca before joining individual spermathecal sac duct, less than length of furca from spermathecal sac duct join; spermathecae indistinct, unpigmented, unsclerotised; spermathecal sac arrangement trilobate (
<figureCitation id="E6C02A14FFBC84251010D75308A7FCD3" box="[1086,1159,868,894]" captionStart="FIGURE 4" captionStartId="13.[264,367,1764,1788]" captionTargetBox="[264,1323,411,1723]" captionTargetId="figure@13.[264,1323,411,1738]" captionTargetPageId="13" captionText="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." httpUri="https://zenodo.org/record/170866/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
C), with spermathecal sac joining common spermathecal sac duct less than length of furca from furca; long ovoid median spermathecal sac with narrow tube to longer oval lobe (
<figureCitation id="E6C02A14FFBC8425105ED7830896FC63" box="[1136,1206,948,974]" captionStart="FIGURE 4" captionStartId="13.[264,367,1764,1788]" captionTargetBox="[264,1323,411,1723]" captionTargetId="figure@13.[264,1323,411,1738]" captionTargetPageId="13" captionText="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." httpUri="https://zenodo.org/record/170866/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
F); pair of ovoid outer spermathecal sacs with narrow tube to large round lobe (
<figureCitation id="E6C02A14FFBC84251003D7EB0854FC5B" box="[1069,1140,988,1014]" captionStart="FIGURE 4" captionStartId="13.[264,367,1764,1788]" captionTargetBox="[264,1323,411,1723]" captionTargetId="figure@13.[264,1323,411,1738]" captionTargetPageId="13" captionText="FIGURE 4. Johnmannia powerae sp. nov. Ψ ANIC 29 0 0 3454 lateral, A B; Ψ ANIC 29 0 0 3453 genitalia, C G. (A) thoracic pruinescence, (B) head, frons and genal flange; (C) SP, SD, SPSD, common SPSD, (D) S 8, furca, S 10, (E) SD, SPSD, common SPSD, (F) lateral (left) and medial SPS, lobes, fine tubes, (G) T 8 with apodeme. Morphological abbreviations in Appendix 1." httpUri="https://zenodo.org/record/170866/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
C).
</paragraph>
</subSubSection>
<subSubSection id="36E1651AFFBC842A1516D0330F3BFEF3" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="7E443691FFBC84251516D033090BF8C6" blockId="7.[264,1324,284,1899]" pageId="7" pageNumber="8">
<emphasis id="4C8FEA83FFBC84251516D0330D54FBB0" box="[312,372,1028,1053]" italics="true" pageId="7" pageNumber="8">Male</emphasis>
. Similar to female except: larger and darker. Upper frons narrower dorsally (
<figureCitation id="E6C02A14FFBC842510D5D0330D38FBEB" captionStart="FIGURE 7" captionStartId="19.[264,367,1730,1754]" captionTargetBox="[264,1323,456,1713]" captionTargetId="figure@19.[264,1323,456,1713]" captionTargetPageId="19" captionText="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri="https://zenodo.org/record/170868/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
C). Apical half of wing darker (Fig. 5C).
<emphasis id="4C8FEA83FFBC842516D2D01B0F4BFBE8" box="[764,875,1068,1093]" italics="true" pageId="7" pageNumber="8">Genitalia</emphasis>
: Gonocoxites with dense tufts of posteriorly directed setae along ventral ridge on basomedial margin (Fig.
<date id="0A451051FFBC84251069D0630841FBC3" box="[1095,1121,1108,1134]" pageId="7" pageNumber="8">5I</date>
), without medial atrium (
<bibRefCitation id="1A6A4B60FFBC8425154AD04B0E55FB3B" author="Winterton" box="[356,629,1148,1174]" pageId="7" pageNumber="24" refString="Winterton, S. L., Irwin, M. E. &amp; Yeates, D. K. (1999 a) Phylogenetic revision of the Taenogera Krober genus-group (Diptera: Therevidae) with descriptions of two new genera. Australian Journal of Entomology, 38, 274 - 290." type="journal article" year="1999" yearSuffix="a">
Winterton
<emphasis id="4C8FEA83FFBC842515CBD04B0E00FB38" box="[485,544,1148,1173]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
1999a
</bibRefCitation>
). Epandrium (Fig. 5D) narrowed posteriorly, with posterolateral extensions, indented medially on posterior surface. Subepandrial sclerite (
<figureCitation id="E6C02A14FFBC842510EFD0930925FB13" box="[1217,1285,1188,1214]" captionStart="FIGURE 7" captionStartId="19.[264,367,1730,1754]" captionTargetBox="[264,1323,456,1713]" captionTargetId="figure@19.[264,1323,456,1713]" captionTargetPageId="19" captionText="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri="https://zenodo.org/record/170868/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
K M) with very well developed, sclerotised, broad plate strongly attached to basiphallus. T8 (
<figureCitation id="E6C02A14FFBC8425153FD0C10D7AFABD" box="[273,346,1270,1296]" captionStart="FIGURE 7" captionStartId="19.[264,367,1730,1754]" captionTargetBox="[264,1323,456,1713]" captionTargetId="figure@19.[264,1323,456,1713]" captionTargetPageId="19" captionText="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri="https://zenodo.org/record/170868/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
OP) emarginate, broadly narrowed medially, setae along posterolateral margin, with lateral spiracular pore (Fig. 5J); S8 (
<figureCitation id="E6C02A14FFBC842516D7D1290F60FA95" box="[761,832,1310,1336]" captionStart="FIGURE 7" captionStartId="19.[264,367,1730,1754]" captionTargetBox="[264,1323,456,1713]" captionTargetId="figure@19.[264,1323,456,1713]" captionTargetPageId="19" captionText="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri="https://zenodo.org/record/170868/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
N) trapezoid, narrowed basally, posterior margin longer than basal margin, black setae along posterior margin. Outer gonocoxal process (Fig. 5GI) well developed, pointed; ventral lobe (
<figureCitation id="E6C02A14FFBC84251786D1460FD1FA26" box="[936,1009,1393,1419]" captionStart="FIGURE 7" captionStartId="19.[264,367,1730,1754]" captionTargetBox="[264,1323,456,1713]" captionTargetId="figure@19.[264,1323,456,1713]" captionTargetPageId="19" captionText="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri="https://zenodo.org/record/170868/files/figure.png" pageId="7" pageNumber="8">
Fig.
<date id="0A451051FFBC842517CCD1460FD1FA26" box="[994,1009,1393,1419]" pageId="7" pageNumber="8">7</date>
</figureCitation>
I) broad, flattened, sclerotised; gonocoxites distinctly separate, not fused ventromedially (
<figureCitation id="E6C02A14FFBC84251019D1AE08A6FA1E" box="[1079,1158,1433,1459]" captionStart="FIGURE 7" captionStartId="19.[264,367,1730,1754]" captionTargetBox="[264,1323,456,1713]" captionTargetId="figure@19.[264,1323,456,1713]" captionTargetPageId="19" captionText="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri="https://zenodo.org/record/170868/files/figure.png" pageId="7" pageNumber="8">
Fig.
<date id="0A451051FFBC84251058D1AE08A6FA1E" box="[1142,1158,1433,1459]" pageId="7" pageNumber="8">7</date>
</figureCitation>
I); gonocoxal apodeme (Fig. 5GI) as long as ventral lobe, expanded apically to form broad gonocoxal apodeme plate. Inner gonocoxal process (
<figureCitation id="E6C02A14FFBC842516D5D1DE0F63F9AE" box="[763,835,1513,1539]" captionStart="FIGURE 7" captionStartId="19.[264,367,1730,1754]" captionTargetBox="[264,1323,456,1713]" captionTargetId="figure@19.[264,1323,456,1713]" captionTargetPageId="19" captionText="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri="https://zenodo.org/record/170868/files/figure.png" pageId="7" pageNumber="8">
Fig.
<date id="0A451051FFBC8425171AD1DE0F63F9AE" box="[820,835,1513,1539]" pageId="7" pageNumber="8">7</date>
</figureCitation>
I) long, narrow, apically acuminate, cupshaped on ventroapical surface, glabrous. Gonostylus (Fig. 5GI) longer than inner gonocoxal process, basally broad and twisted, some setae on basolateral surface, many setae on medial surface; dorsal recurved triangulate subapical hook, 1/3 length of gonostylus from apex; apical portion of gonostylus narrowing, recurved dorsomedially, cup­shaped on dorsal surface. Hypandrium (Fig. 5G) triangular, separate from gonocoxites but joined laterally. Distiphallus (
<figureCitation id="E6C02A14FFBC842515C6D2EE0E11F95E" box="[488,561,1753,1779]" captionStart="FIGURE 7" captionStartId="19.[264,367,1730,1754]" captionTargetBox="[264,1323,456,1713]" captionTargetId="figure@19.[264,1323,456,1713]" captionTargetPageId="19" captionText="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri="https://zenodo.org/record/170868/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
KM) smoothly recurved from base, directed ventrally, longer than ejaculatory apodeme, spinulose on dorsal surface; dorsal apodeme (
<figureCitation id="E6C02A14FFBC84251059D336089FF8B6" box="[1143,1215,1793,1819]" captionStart="FIGURE 7" captionStartId="19.[264,367,1730,1754]" captionTargetBox="[264,1323,456,1713]" captionTargetId="figure@19.[264,1323,456,1713]" captionTargetPageId="19" captionText="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri="https://zenodo.org/record/170868/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
KM) of parameral sheath with anteriorly directed lateral arm; ventral apodeme of parameral sheath short; ejaculatory apodeme (Fig. 5L) broad anteriorly; lateral ejaculatory apodeme (
<figureCitation id="E6C02A14FFBC842510D5D366090BF8C6" box="[1275,1323,1873,1899]" captionStart="FIGURE 7" captionStartId="19.[264,367,1730,1754]" captionTargetBox="[264,1323,456,1713]" captionTargetId="figure@19.[264,1323,456,1713]" captionTargetPageId="19" captionText="FIGURE 7. Johnmannia kosciuszkoensis sp. nov. ɗ ANIC 29 0 0 4008 body A F, genitalia G P. (A) body, lateral, (B) abdomen; C E head, (C) frontal, (D) lateral, (E) occiput; (F) wing pattern and venation; G H epandrium, (G) dorsal, (H) lateral; I J gonocoxal complex, (I) ventral, (J) lateral; K M aedeagus, (K) dorsal; (L) lateral, (M) ventral; (N) S 8, ventral; O P, T 8, (O) dorsal, (P) lateral with spiracle." httpUri="https://zenodo.org/record/170868/files/figure.png" pageId="7" pageNumber="8">Fig.</figureCitation>
</paragraph>
<paragraph id="7E443691FFB3842A1526D52B0F3BFEF3" blockId="8.[264,1323,284,790]" pageId="8" pageNumber="9">7KM) narrow, ribbon­like, separation less than base of distiphallus; basal ejaculatory apodeme broad, spinulose on ventral surface.</paragraph>
</subSubSection>
<subSubSection id="36E1651AFFB3842A1516D55B0887FDDB" pageId="8" pageNumber="9" type="discussion">
<paragraph id="7E443691FFB3842A1516D55B086BFE7B" blockId="8.[264,1323,284,790]" pageId="8" pageNumber="9">
<emphasis id="4C8FEA83FFB3842A1516D55B0D94FE28" box="[312,436,364,389]" italics="true" pageId="8" pageNumber="9">Comments</emphasis>
. Very rare. In this genus, female specimens have more often been collected, described, and given
<typeStatus id="A1408833FFB3842A1626D5A30E4FFE03" box="[520,623,404,430]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
status. We will continue this trend by basing our descriptions upon the female, and then comparing the male to that description.
</paragraph>
<paragraph id="7E443691FFB3842A1516D5D30887FDDB" blockId="8.[264,1323,284,790]" pageId="8" pageNumber="9">
<bibRefCitation id="1A6A4B60FFB3842A1516D5D30E4BFE53" author="Irwin" box="[312,619,484,510]" lastPageId="8" lastPageNumber="23" pageId="8" pageNumber="22" refString="Irwin, M. E. &amp; Lyneborg, L. (1989) Family Therevidae. In: Evenhuis, N. L. (Ed) Catalog of the Diptera of the Australasian and Oceanian Regions, Bishop Museum Press and E. J. Brill, Honolulu, pp. 1155." type="book chapter" year="1989">Irwin &amp; Lyneborg (1989)</bibRefCitation>
erected the name
<taxonomicName id="B9FB4D12FFB3842A1762D5D30FFDFE50" box="[844,989,484,509]" class="Insecta" family="Therevidae" genus="Johnmannia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="4C8FEA83FFB3842A1762D5D30FFDFE50" box="[844,989,484,509]" italics="true" pageId="8" pageNumber="9">Johnmannia</emphasis>
</taxonomicName>
citing preoccupation of the name
<taxonomicName id="B9FB4D12FFB3842A157FD63B0D8CFD88" box="[337,428,524,549]" class="Marchantiopsida" family="Aytoniaceae" genus="Mannia" kingdom="Plantae" order="Marchantiales" pageId="8" pageNumber="9" phylum="Bryophyta" rank="genus">
<emphasis id="4C8FEA83FFB3842A157FD63B0D8CFD88" box="[337,428,524,549]" italics="true" pageId="8" pageNumber="9">Mannia</emphasis>
</taxonomicName>
by
<bibRefCitation id="1A6A4B60FFB3842A15F5D63B0E8DFD8B" author="Dewalque" box="[475,685,524,550]" pageId="8" pageNumber="22" refString="Dewalque, G. (1868) Prodrome d'une description geologique de la Belgique, Carmanne, Liege, 442 pp." type="book" year="1868">Dewalque (1868)</bibRefCitation>
. However, the
<bibRefCitation id="1A6A4B60FFB3842A174FD63B0800FD8B" author="Dewalque" box="[865,1056,524,550]" pageId="8" pageNumber="22" refString="Dewalque, G. (1868) Prodrome d'une description geologique de la Belgique, Carmanne, Liege, 442 pp." type="book" year="1868">Dewalque 1868</bibRefCitation>
: 432 name for a fossil had been declared a
<emphasis id="4C8FEA83FFB3842A15D2D6030E83FDE0" box="[508,675,564,589]" italics="true" pageId="8" pageNumber="9">nomen nudum</emphasis>
(
<bibRefCitation id="1A6A4B60FFB3842A169DD6030F68FDE3" author="Neave" box="[691,840,564,590]" pageId="8" pageNumber="24" refString="Neave, S. A. (Ed) (1940) Nomenclator Zoologicus. A List of the Names of Genera and Subgenera in Zoology from the Tenth Edition of Linnaeus 1758 to the end of 1935., The Zoological Society of London, London, 1065 pp." type="book" year="1940">Neave 1940</bibRefCitation>
). The name
<taxonomicName id="B9FB4D12FFB3842A17F2D6030817FDE0" box="[988,1079,564,589]" class="Marchantiopsida" family="Aytoniaceae" genus="Mannia" kingdom="Plantae" order="Marchantiales" pageId="8" pageNumber="9" phylum="Bryophyta" rank="genus">
<emphasis id="4C8FEA83FFB3842A17F2D6030817FDE0" box="[988,1079,564,589]" italics="true" pageId="8" pageNumber="9">Mannia</emphasis>
</taxonomicName>
was preoccupied by
<bibRefCitation id="1A6A4B60FFB3842A1526D66B0DEDFDDB" author="Davidson" box="[264,461,604,630]" pageId="8" pageNumber="22" refString="Davidson, T. (1874) On the Tertiary Brachiopoda of Belgium. Geological Magazine, 1, 150 - 159." type="journal article" year="1874">Davidson (1874)</bibRefCitation>
for a fossil brachiopod, and again by
<bibRefCitation id="1A6A4B60FFB3842A17A7D66B083EFDDB" author="Prout" box="[905,1054,604,630]" pageId="8" pageNumber="24" refString="Prout, L. B. (1915) Die spannerartigen Nachtfalter. In: Seitz, A. (Ed) Die Gross-Schmetterlinge der Erde. Eine systematische Bearbeitung der bis jetzt bekannten Gross-Schmetterlinge. I. Abteilung: Die Gross-Schmetterlinge des palaarktischen Faunengebietes., vol. 4, Verlag, Stuttgart, pp. I - V, 1 - 479, pls 1 - 25." type="journal article" year="1915">Prout (1915)</bibRefCitation>
for a moth.
</paragraph>
</subSubSection>
<subSubSection id="36E1651AFFB3842A1516D6B30FDBFCBB" pageId="8" pageNumber="9" type="distribution">
<paragraph id="7E443691FFB3842A1516D6B30E34FD6B" blockId="8.[264,1323,284,790]" pageId="8" pageNumber="9">
<emphasis id="4C8FEA83FFB3842A1516D6B30DE7FD30" box="[312,455,644,669]" italics="true" pageId="8" pageNumber="9">Distribution</emphasis>
. SE Queensland; Lansdowne (
<geoCoordinate id="1BCF5056FFB3842A1718D6B30FB2FD33" box="[822,914,644,670]" direction="south" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="-31.783333">31°47'S</geoCoordinate>
­
<geoCoordinate id="1BCF5056FFB3842A1780D6B3083DFD33" box="[942,1053,644,670]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="152.53334">152°32'E</geoCoordinate>
), Kosciuszko National Park, NSW; Tasmania.
</paragraph>
<paragraph id="7E443691FFB3842A1516D6E30FDBFCBB" blockId="8.[264,1323,284,790]" pageId="8" pageNumber="9">
<emphasis id="4C8FEA83FFB3842A1516D6E30DDEFD40" box="[312,510,724,749]" italics="true" pageId="8" pageNumber="9">Included species</emphasis>
:
<taxonomicName id="B9FB4D12FFB3842A163CD6E30FEEFD43" authority="Paramonov" authorityName="Paramonov" box="[530,974,724,750]" class="Insecta" family="Therevidae" genus="Johnmannia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tasmanica">
<emphasis id="4C8FEA83FFB3842A163CD6E30F06FD40" box="[530,806,724,749]" italics="true" pageId="8" pageNumber="9">Johnmannia tasmanica</emphasis>
(Paramonov)
</taxonomicName>
,
<taxonomicName id="B9FB4D12FFB3842A17F0D6E30D92FCBB" authority="Lambkin &amp; Recsei" authorityName="Lambkin &amp; Recsei" class="Insecta" family="Therevidae" genus="Johnmannia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="powerae">
<emphasis id="4C8FEA83FFB3842A17F0D6E308C0FD40" box="[990,1248,724,749]" italics="true" pageId="8" pageNumber="9">Johnmannia powerae</emphasis>
Lambkin &amp; Recsei
</taxonomicName>
,
<taxonomicName id="B9FB4D12FFB3842A1590D6CB0FDBFCBB" authority="Lambkin &amp; Recsei." authorityName="Lambkin &amp; Recsei." box="[446,1019,764,790]" class="Insecta" family="Therevidae" genus="Johnmannia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kosciuszkoensis">
<emphasis id="4C8FEA83FFB3842A1590D6CB0F2FFCB8" box="[446,783,764,789]" italics="true" pageId="8" pageNumber="9">Johnmannia kosciuszkoensis</emphasis>
Lambkin &amp; Recsei.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>