228 lines
20 KiB
XML
228 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.722.21192" ID-GBIF-Dataset="a84fe3cb-e432-4f4a-8322-331b8b942069" ID-PMC="PMC5740414" ID-Pensoft-Pub="1313-2970-722-1" ID-PubMed="29308027" ID-ZBK="CAF34BA6EEF3486C905B96F03A0F20B4" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-722-1" ModsDocOrigin="ZooKeys 722" ModsDocTitle="Two new species of the millipede genus Glyphiulus Gervais, 1847 from Laos (Diplopoda, Spirostreptida, Cambalopsidae)" checkinTime="1513190152484" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I., Inkhavilay, Khamla, Sutcharit, Chirasak, Srisonchai, Ruttapon & Panha, Somsak" docDate="2017" docId="F4BC69E75F3698790BDA0EC75101C7EF" docLanguage="en" docName="ZooKeys 722: 1-18" docOrigin="ZooKeys 722" docSource="http://dx.doi.org/10.3897/zookeys.722.21192" docTitle="Glyphiulus semicostulifer Likhitrakarn, Golovatch & Panha, sp. n." docType="treatment" docUuid="6FCD27DF-7566-42F4-92D5-63F6EDDBA66B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="9" masterDocId="FFA1FF8FFFB8FFF4FFCDFFEBFFBEFF94" masterDocTitle="Two new species of the millipede genus Glyphiulus Gervais, 1847 from Laos (Diplopoda, Spirostreptida, Cambalopsidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="4" updateTime="1668165208222" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of the millipede genus Glyphiulus Gervais, 1847 from Laos (Diplopoda, Spirostreptida, Cambalopsidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Inkhavilay, Khamla</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Srisonchai, Ruttapon</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>722</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>18</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.722.21192</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.722.21192</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-722-1</mods:identifier>
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<mods:identifier type="ZBK">CAF34BA6EEF3486C905B96F03A0F20B4</mods:identifier>
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<mods:identifier type="ZooBank">CAF34BA6EEF3486C905B96F03A0F20B4</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="137687667" LSID="urn:lsid:zoobank.org:act:6FCD27DF-7566-42F4-92D5-63F6EDDBA66B" httpUri="http://treatment.plazi.org/id/F4BC69E75F3698790BDA0EC75101C7EF" lastPageId="8" lastPageNumber="9" pageId="3" pageNumber="4">
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<subSubSection pageId="3" pageNumber="4" type="nomenclature">
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<paragraph pageId="3" pageNumber="4">
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<taxonomicName LSID="http://zoobank.org/6FCD27DF-7566-42F4-92D5-63F6EDDBA66B" authority="Likhitrakarn, Golovatch & Panha" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus semicostulifer" order="Spirostreptida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="semicostulifer">Glyphiulus semicostulifer Likhitrakarn, Golovatch & Panha</taxonomicName>
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<taxonomicNameLabel pageId="3" pageNumber="4">sp. n.</taxonomicNameLabel>
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Figs 1C, D, 5, 6, 7
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="type material">
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<paragraph pageId="4" pageNumber="5">
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<pageBreakToken pageId="4" pageNumber="5" start="start">Type</pageBreakToken>
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material.
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</paragraph>
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<paragraph pageId="4" pageNumber="5">
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Holotype ♂ (CUMZ), Laos, Luang Namtha Province, Viengphoukha District, Kao Rao Cave, 737 m a.s.l.,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="20.725">20°43'30"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="101.151115">101°09'04"E</geoCoordinate>
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, 12.10.2014, leg. C. Sutcharit and R. Srisonchai.
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</paragraph>
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<paragraph pageId="4" pageNumber="5">Paratypes: 6 ♂, 16 ♀, 8 juveniles (CUMZ), 1 ♂, 2 ♀ (ZMUM), same locality, together with holotype.</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="etymology">
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<paragraph pageId="4" pageNumber="5">Etymology.</paragraph>
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<paragraph pageId="4" pageNumber="5">
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To emphasize the obvious similarities to
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<taxonomicName lsidName="G. costulifer" pageId="4" pageNumber="5" rank="species" species="costulifer">G. costulifer</taxonomicName>
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Golovatch, Geoffroy,
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<normalizedToken originalValue="Mauriès">Mauries</normalizedToken>
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& VandenSpiegel, 2007.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="5" type="diagnosis">
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<paragraph pageId="4" pageNumber="5">Diagnosis.</paragraph>
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<paragraph pageId="4" pageNumber="5">
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This new species is particularly similar to
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<taxonomicName lsidName="G. costulifer" pageId="4" pageNumber="5" rank="species" species="costulifer">G. costulifer</taxonomicName>
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, with which it shares the following diagnostic characters: the unique carinotaxy formulae, coupled with anterior gonopod structural details. It differs from
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<taxonomicName lsidName="G. costulifer" pageId="4" pageNumber="5" rank="species" species="costulifer">G. costulifer</taxonomicName>
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by the more sparsely alveolate background fine structure of the metazonae, coupled with the gnathochilarium being considerably less densely setose on the caudal face, the paramedian coxal prongs on ♂ legs 1 much stronger and their telopodites 4-segmented, the apicoparamedian sternal projections on the anterior gonopods slightly longer and more slender, and the flagella of the posterior gonopods much longer. See also Key below.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6" type="description">
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<paragraph pageId="5" pageNumber="6">
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<pageBreakToken pageId="5" pageNumber="6" start="start">Description</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="5" pageNumber="6">Length of holotype ca 19.4 mm; adult paratypes 17.2-25.3 (♂) or 17.5-25.6 mm long (♀), juveniles 1.34-1.79 mm long; midbody segments round in cross-section (Fig. 5L), their width (horizontal diameter) and height (vertical diameter) similar, width in holotype 1.4 mm; paratypes 1.1-1.5 (♂), 0.9-1.5 (♀) or 0.9-1.1 mm (juveniles).</paragraph>
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<caption pageId="5" pageNumber="6">
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<paragraph pageId="5" pageNumber="6">
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Figure 5.
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<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus semicostulifer" order="Spirostreptida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="semicostulifer">Glyphiulus semicostulifer</taxonomicName>
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sp. n.,
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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,
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<normalizedToken originalValue="H–Q">H-Q</normalizedToken>
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♀ paratypes
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<normalizedToken originalValue="D–G">D-G</normalizedToken>
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♂ paratype
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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anterior part of body, lateral and dorsal views, respectively C collum and body segments 2 and 3, dorsal view D cephalic capsule, dorsal view E gnathochilarium, ventral view F apical part of antenna, ventral view G bacilliform sensilla on antennomere 5, lateral view H, I midbody segments, lateral and dorsal views, respectively J midbody prozona enlarged, dorsal view K limbus, dorsal view L cross-section of midbody segment M midbody sternite and coxae, frontal view N porostele, lateral view
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<normalizedToken originalValue="O–Q">O-Q</normalizedToken>
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posterior part of body, lateral, dorsal and ventral views, respectively.
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</paragraph>
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</caption>
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<paragraph pageId="5" pageNumber="6">Coloration of live animals dark brown to red-brownish (Fig. 1C), with contrasting light yellow head, antennae, collum, segments 2 and 3, sometimes segment 4 as well; telson yellow-brown, venter and legs brownish yellow to brownish red, ocellaria blackish, lateral longitudinal stripes above porosteles brownish; a thin axial line traceable due to darker median crests, sometimes body uniformly yellowish to brownish yellow (Fig. 1D); coloration in alcohol, after three years of preservation similar, but telson light brownish, venter and legs brownish yellow to brownish, lateral longitudinal stripes brownish to brownish red.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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Body with 52p+4a+T (holotype); paratypes with 52
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<normalizedToken originalValue="–67p+2–">-67p+2-</normalizedToken>
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4a+T (♂), 49
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<normalizedToken originalValue="–60p+2–">-60p+2-</normalizedToken>
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4a+T (♀) or 41-50+5a+T (juveniles). Eye patches transversely ovoid, each composed of 7-11 blackish, rather flat ocelli in 4 or 5 irregular longitudinal rows (Fig. 5D). Antennae short and clavate (Figs 1C, D, 5
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<normalizedToken originalValue="A–E">A-E</normalizedToken>
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, 7A), extending behind segment 3 laterally, antennomeres 5 and 6 each with a small distoventral group or corolla of bacilliform sensilla (Fig. 5F, G). Gnathochilarium with a clearly separated promentum (Figs 5E, 6B, 7B).
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</paragraph>
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<paragraph pageId="5" pageNumber="6">Head width = segment 2 <collum = midbody segment (close to 8th to 10th)> segment 3 = 5> 4 <7 <8 = 10; body abruptly tapering towards telson on a few posteriormost segments (Fig. 5P). Postcollar constriction very evident (Fig. 5B).</paragraph>
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<paragraph pageId="5" pageNumber="6">
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Collum nearly smooth, carinotaxy formula 1+2c+3-4+5c+6a+pc+ma (Fig. 5
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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), with 6+6 longitudinal crests starting from anterior edge, but both median crests interrupted in about caudal 2/3-3/4, being replaced there by similar 1+1+1 crests.
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</paragraph>
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<paragraph pageId="5" pageNumber="6">
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Following metaterga similarly strongly crested (Figs 1C, D, 5
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, H, I, O, P), especially from segment 5 on, whence porosteles commence (Fig. 5A, B), smaller tubercles in their stead on legless segments in front of telson due to loss of ozopores (Fig. 5P). Porosteles large, conical, round, directed caudolaterad, wider than high (Fig. 5N), ozoporiferous crests distinctly divided into two about midway, their frontal halves being higher (Fig. 5A, B, H, I, O, P). Carinotaxy formulae 2+I/i+3/3+I/i+2 on segments 2-3, as well as on the last 1-2 leg-bearing and legless segments (Fig. 5A, B, O, P); midbody segments showing all dorsal crests subdivided transversely (carinotaxy formulae 2/2+I/i+3/3+I/i+2/2) and sharper, especially so lateral crests (Fig. 5H, I).
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</paragraph>
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<paragraph lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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Tegument delicately alveolate-areolate (Fig. 5A, B,
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<normalizedToken originalValue="H–J">H-J</normalizedToken>
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, M, O, P), dull throughout. Fine longitudinal striations in front of stricture between pro- and metazona, and remaining surface of prozona very delicately shagreened (Fig. 5J). Metatergal setae absent. Segments 2 and 3 with long pleural flaps. Limbus extremely finely and more or less regularly denticulate (Fig. 5K). Epiproct (Fig. 5O, P) simple, with an evident lateral tubercle placed level with ozoporiferous crests, also with a rounded ridge in caudal part and an evident axial rib dorsally. Paraprocts rather regularly convex, each
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<pageBreakToken pageId="6" pageNumber="7" start="start">with</pageBreakToken>
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a faint premarginal sulcus medially (Fig. 5Q). Hypoproct bean-shaped, with 1+1 strongly separated setae near caudal margin (Fig. 5Q).
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</paragraph>
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<paragraph pageId="6" pageNumber="7">Ventral flaps behind gonopod aperture on ♂ segment 7 rather well distinguishable as low swellings forming a bare transverse ridge.</paragraph>
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<paragraph pageId="6" pageNumber="7">Legs rather short, on midbody segments about 3/4 length of segment height (Figs 5A, L, O, 7F). Claw at base with a strong accessory spine about 1/3-1/4 length of main claw. Tarsi and tarsal setae very delicately fringed (Fig. 7F).</paragraph>
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<paragraph pageId="7" pageNumber="8">
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<pageBreakToken pageId="7" pageNumber="8" start="start">♂</pageBreakToken>
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legs 1 highly characteristic (Figs 6A, 7C) in showing nearly fully developed, 4-segmented telopodites and a pair of large, subdigitiform, medially contiguous, but apically diverging coxal processes with a groups of long and strong setae at base.
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</paragraph>
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<caption pageId="7" pageNumber="8">
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<paragraph pageId="7" pageNumber="8">
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Figure 6.
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<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus semicostulifer" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="semicostulifer">Glyphiulus semicostulifer</taxonomicName>
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sp. n., ♂ paratype. A legs 1, frontal view B legs 2, caudal view C penes, caudal view D, E legs 3, caudal and frontal views, respectively F, G anterior gonopods, frontal and caudal views, respectively H microsetae at base of telopodite on anterior gonopod I, J microsetae on posterior coxosternum of anterior gonopod L, M posterior gonopods, caudal and frontal views, respectively K distal part of flagellum of posterior gonopod N microsetae at base of posterior gonopod, frontal view O paramedian lobes on posterior gonopods.
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</paragraph>
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</caption>
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<caption pageId="7" pageNumber="8">
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<paragraph pageId="7" pageNumber="8">
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Figure 7.
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<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus semicostulifer" order="Spirostreptida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="semicostulifer">Glyphiulus semicostulifer</taxonomicName>
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sp. n., ♂ holotype. A antenna, lateral view B gnathochilarium, ventral view C legs 1, frontal view D legs 2, caudal view E leg 3, caudal view F midbody leg G anterior gonopods, caudal view H, I posterior gonopods, frontal and caudal views, respectively. Scale bars: 0.2 mm.
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</paragraph>
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</caption>
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<paragraph pageId="7" pageNumber="8">♂ legs 2 nearly normal, only claw and, anteriorly, coxa somewhat reduced, and femur abbreviated on frontal face; penes broad, rounded, each with 4-5 strong setae distolaterally (Figs 6B, C, 7D).</paragraph>
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<paragraph pageId="7" pageNumber="8">♂ legs 3 modified in having coxa especially slender and elongate (Figs 6D, E, 7E).</paragraph>
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<paragraph pageId="8" pageNumber="9">
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<pageBreakToken pageId="8" pageNumber="9" start="start">Anterior</pageBreakToken>
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gonopods (Figs 6
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<normalizedToken originalValue="F–J">F-J</normalizedToken>
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, 7G) with a typical shield-like coxosternum, this being rather densely microsetose on caudal face (Fig. 6G, I, J), with a high, digitiform, apicomesal process (d). Telopodite typical, slender, movable, 1-segmented, lateral in position, with 3-5 strong apical setae and a field of microsetae at base (Fig. 6H).
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</paragraph>
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<paragraph pageId="8" pageNumber="9">
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Posterior gonopods (Figs 6
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<normalizedToken originalValue="K–O">K-O</normalizedToken>
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, 7H, I) highly compact, both contiguous basally until about midheight; two densely and strongly setose lobes (l) paramedially (Fig. 6O); each half with two higher central pieces with a seminal groove between, frontal piece (a) elongate; long, distally microplumose flagella (f) (Fig. 6
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<normalizedToken originalValue="K–M">K-M</normalizedToken>
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); caudal piece (b) subquadrate, membranous, micropapillate frontolaterally, with an apical field of coniform microsetae laterally (Fig. 6L, N).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="remarks">
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<paragraph pageId="8" pageNumber="9">Remark.</paragraph>
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<paragraph pageId="8" pageNumber="9">The javanicus-group is currently comprised of 23 species, including this new species, a fourth in this group to be reported from Laos.</paragraph>
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<caption pageId="8" pageNumber="9">
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<paragraph pageId="8" pageNumber="9">
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Figure 8. Distribution of
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<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Glyphiulus</taxonomicName>
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species in Laos (6 species): Open diamond:
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<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus semicostulifer" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="semicostulifer">Glyphiulus semicostulifer</taxonomicName>
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sp. n. Open circle:
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<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus bedosae" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bedosae">Glyphiulus bedosae</taxonomicName>
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Golovatch, Geoffroy,
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<normalizedToken originalValue="Mauriès">Mauries</normalizedToken>
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& VandenSpiegel, 2007. Filled triangle:
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<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus costulifer" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="costulifer">Glyphiulus costulifer</taxonomicName>
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Golovatch, Geoffroy,
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<normalizedToken originalValue="Mauriès">Mauries</normalizedToken>
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& VandenSpiegel, 2007. Open square:
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<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus subbedosae" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="subbedosae">Glyphiulus subbedosae</taxonomicName>
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sp. n. Inverted open triangle:
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<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus subcostulifer" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="subcostulifer">Glyphiulus subcostulifer</taxonomicName>
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Golovatch, Geoffroy,
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<normalizedToken originalValue="Mauriès">Mauries</normalizedToken>
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& VandenSpiegel, 2007. Filled circle:
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<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus percostulifer" order="Spirostreptida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="percostulifer">Glyphiulus percostulifer</taxonomicName>
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Golovatch, Geoffroy,
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<normalizedToken originalValue="Mauriès">Mauries</normalizedToken>
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& VandenSpiegel, 2007.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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||
</document> |