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<document ID-CLB-Dataset="7802" ID-DOI="http://dx.doi.org/10.3897/zookeys.1023.59811" ID-GBIF-Dataset="ff06869b-5933-4bfd-a781-51c864788a77" ID-PMC="PMC7969580" ID-Pensoft-Pub="1313-2970-1023-49" ID-Pensoft-UUID="50CDE76120CD5FE19696920D05B8C2DA" ID-PubMed="33776514" ID-ZooBank="8299700AD3904354BA5ADDD004165B32" ModsDocID="1313-2970-1023-49" checkinTime="1615410365580" checkinUser="pensoft" docAuthor="Wang, Xian-Yi, Li, Zi-Zhong &amp; Dai, Ren-huai" docDate="2021" docId="F42E4C2261C55BAF92091B0500F35FCD" docLanguage="en" docName="ZooKeys 1023: 49-60" docOrigin="ZooKeys 1023" docPubDate="2021-03-10" docSource="http://dx.doi.org/10.3897/zookeys.1023.59811" docTitle="Calodia quadrimacula Wang &amp; Li &amp; Dai 2021, sp. nov." docType="treatment" docUuid="8C89F5F1-C4DC-4FA5-92ED-6AC3C1D9268F" docUuidSource="ZooBank" docVersion="7" id="50CDE76120CD5FE19696920D05B8C2DA" lastPageNumber="49" masterDocId="50CDE76120CD5FE19696920D05B8C2DA" masterDocTitle="Two new species of the leafhopper genus Calodia Nielson (Hemiptera, Cicadellidae, Coelidiinae) from China, with a checklist and key to Chinese species" masterLastPageNumber="60" masterPageNumber="49" pageNumber="49" updateTime="1732821750904" updateUser="ExternalLinkService">
<mods:mods id="AD922174FD970902C96EC973729623C1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1E93179A050FD6C5FC6EC58EE29ECB54">
<mods:title id="C4F4BE1B71D2A19164FB9BAEED209198">Two new species of the leafhopper genus Calodia Nielson (Hemiptera, Cicadellidae, Coelidiinae) from China, with a checklist and key to Chinese species</mods:title>
</mods:titleInfo>
<mods:name id="DE82FA684A4981C20F1781A679BE3B91" type="personal">
<mods:role id="58BB5B976519FE2D9BFCD061F95C6C95">
<mods:roleTerm id="CB82F4B1F7C47F06138794B4E0AD489F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27637365959BB4B8545B8677F429D82D">Wang, Xian-Yi</mods:namePart>
<mods:affiliation id="CFE567B4B9D8A341B065D91A2A5E0DF6">Institute of Entomology, Guizhou University; Guizhou Provincial Key Laboratory for Agricultural Pest Management of the Mountainous Region, Guiyang 550025, China</mods:affiliation>
</mods:name>
<mods:name id="B692F861CDCDBBF7A383D408E3B55BDF" type="personal">
<mods:role id="2FA12F1EFA7910C4BA6AC49A391439FE">
<mods:roleTerm id="A58A9ED4FAFE3542B018A6742CFCDF8B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="04DC658A93B9D3418061D2849CFCB313">Li, Zi-Zhong</mods:namePart>
<mods:affiliation id="B61972A948075198AE741FF5EBB0A457">Institute of Entomology, Guizhou University; Guizhou Provincial Key Laboratory for Agricultural Pest Management of the Mountainous Region, Guiyang 550025, China</mods:affiliation>
</mods:name>
<mods:name id="3F8589BB66EC2D62465C25E67D9BB793" type="personal">
<mods:role id="3510B0DE8D27CEF530DFB7A04D81872E">
<mods:roleTerm id="F0E7997F4973B531493848A60F306606">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB3A1924A5FACB094497A0DE91C819D0">Dai, Ren-huai</mods:namePart>
<mods:nameIdentifier id="EC49F74CF82E9C53E3612C9754F1DA55" type="ORCID">https://orcid.org/0000-0001-7652-6808</mods:nameIdentifier>
<mods:affiliation id="FFDB2CB77EA0E84014953F548E9B5F71">Institute of Entomology, Guizhou University; Guizhou Provincial Key Laboratory for Agricultural Pest Management of the Mountainous Region, Guiyang 550025, China</mods:affiliation>
<mods:nameIdentifier id="058A2F940286C11BABDBD377312D4D1C" type="email">dmolbio@126.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="75B10F6AE8FA021A0B18AC47DC4139F5">text</mods:typeOfResource>
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<mods:title id="35FBD8FF946BA87EECDAA9805BE1124C">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="09019C0577BD0490999BE5316F8B9A02">
<mods:date id="43BBC48EBA2E20513D62D960722DE033">2021</mods:date>
<mods:detail id="41A466AF421C735F930A3878A5477FCC" type="pubDate">
<mods:number id="3BD4F7F8A3993027922F7E53801B40B2">2021-03-10</mods:number>
</mods:detail>
<mods:detail id="D56A437A9DA08F3A404DBDC47FA9B97A" type="volume">
<mods:number id="B10834DD0458F8AF9009B37121B66824">1023</mods:number>
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<mods:start id="1CC6787D6D84FF0C5215DDEA4EA8E707">49</mods:start>
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<mods:url id="22A7A0F61116A574AA1A7712082B9BF4">http://dx.doi.org/10.3897/zookeys.1023.59811</mods:url>
</mods:location>
<mods:classification id="714BD622BBA1E225D2D88517197AFF1C">journal article</mods:classification>
<mods:identifier id="E6C2681A4DF51150ED1277324B777B95" type="DOI">http://dx.doi.org/10.3897/zookeys.1023.59811</mods:identifier>
<mods:identifier id="C7EEB964361DA25F46135888179A2E1F" type="Pensoft-Pub">1313-2970-1023-49</mods:identifier>
<mods:identifier id="6F8A361C8A7306FF7C42AEA8C173F40B" type="ZooBank">8299700AD3904354BA5ADDD004165B32</mods:identifier>
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<treatment id="F42E4C2261C55BAF92091B0500F35FCD" ID-GBIF-Taxon="180302186" LSID="urn:lsid:zoobank.org:act:8C89F5F1-C4DC-4FA5-92ED-6AC3C1D9268F" httpUri="http://treatment.plazi.org/id/F42E4C2261C55BAF92091B0500F35FCD" lastPageNumber="49" pageId="0" pageNumber="49" scope_family="Cicadellidae" scope_order="Hemiptera" scope_subFamily="Coelidiinae">
<subSubSection id="DDCF4D949D4EDB2E2180B7E87686948F" pageId="0" pageNumber="49" type="nomenclature">
<paragraph id="DA4A93B1145EF7E8EA0F3234C60BD44C" pageId="0" pageNumber="49">
<taxonomicName id="CE46DA12EAA6E9805918CE8955B356BA" ID-CoL="8KGTM" LSID="http://zoobank.org/8C89F5F1-C4DC-4FA5-92ED-6AC3C1D9268F" authority="Wang &amp; Li &amp; Dai, 2021" authorityName="Wang &amp; Li &amp; Dai" authorityYear="2021" class="Insecta" family="Cicadellidae" genus="Calodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calodia quadrimacula" order="Hemiptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="quadrimacula" status="sp. nov.">Calodia quadrimacula</taxonomicName>
<taxonomicNameLabel id="3871BB54B434689507C912B7D6B34F8D" pageId="0" pageNumber="49">sp. nov.</taxonomicNameLabel>
<figureCitation id="72F819DBDD895FDB251A2509B6AD8704" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">Figs 14-23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8C9220FE3B3BBFB26C615B8CF476DBDA" pageId="0" pageNumber="49" type="type material">
<paragraph id="1E7C2311ECE584E4010DFB963D10BC08" pageId="0" pageNumber="49">
<typeStatus id="90F3010CCDBA7F77CEC0507459CABD6D">Type</typeStatus>
material.
</paragraph>
<paragraph id="0DAF9D835D16AE6DDD701D6EEA12FD1D" pageId="0" pageNumber="49">
<emphasis id="42D62DA39E810A2D149DC38B79A3A766" bold="true" pageId="0" pageNumber="49">
<emphasis id="EC93E4ED218FF0E72E5585BC3879CC92" italics="true" pageId="0" pageNumber="49">
<typeStatus id="FF6A437610D02A44C3EB59A2A9292195">Holotype</typeStatus>
</emphasis>
,
</emphasis>
<materialsCitation id="3CC1F770DDC62AFCD0EBD8B483E8B3B7" ID-GBIF-Occurrence="3063633302" collectingDate="2017-07-03" collectionCode="GUGC" collectorName="Caohai &amp; Paratype" county="Weining County" location="Bijie City" municipality="China" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
<specimenCount id="C8E3B057DF902A83B50FF3A8A5E1B811">♂</specimenCount>
,
<collectingMunicipality id="89F19C1F623075D65DF8409A585E5450">China</collectingMunicipality>
:
<collectingRegion id="CF3603F9E6FD4580CAC222F30025F9BD" country="China" name="Guizhou">Guizhou Province</collectingRegion>
,
<location id="FE117ED9DD1C697B941E0E6BA256F587" LSID="urn:lsid:plazi:treatment:F42E4C2261C55BAF92091B0500F35FCD:FE117ED9DD1C697B941E0E6BA256F587" county="Weining County" municipality="China" name="Bijie City" stateProvince="Guizhou">Bijie City</location>
,
<collectingCounty id="84A16F509EDF2AA3B918362666CD4274">Weining County</collectingCounty>
,
<location id="CF3A071A385849C505C13DA1C52B6CF6" LSID="urn:lsid:plazi:treatment:F42E4C2261C55BAF92091B0500F35FCD:CF3A071A385849C505C13DA1C52B6CF6" county="Weining County" municipality="China" name="Caohai Reserve" stateProvince="Guizhou">Caohai Reserve</location>
,
<date id="1A8E650699F1610AD9CFBAD71B9F91E0" value="2017-07-03">
<collectingDate id="FDBA2088D81789F61799D0C08F5F98E5" value="2017-07-03">3 July 2017</collectingDate>
</date>
, coll.
<collectorName id="CA3BDE132C499D5B02C93042C0E0A7D7">Caohai</collectorName>
expedition team (
<collectionCode id="0C7F59C94DD5ECA2581CCF1B107598DE">GUGC</collectionCode>
).
<emphasis id="15912E7ED616A7344B9450726BA79F0A" bold="true" italics="true" pageId="0" pageNumber="49">
<typeStatus id="C13188A6BAB2E4FEAA3D8CA034301EF0">
<collectorName id="0F320538E3BAC0567BB56E430D10D54E">Paratype</collectorName>
</typeStatus>
</emphasis>
</materialsCitation>
,
<specimenCount id="8548C95131B05DC1332DA2632F7A1CD2" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="D2CDFB6C84C1E475E33CFF4651FFE72C" type="female">6 ♀♀</specimenCount>
same information as
<typeStatus id="BB9D06023F6B289D545DE557AAD4CADB">holotype</typeStatus>
.
<materialsCitation id="9C8AE5070046D309D8626CD74BB33FA7" ID-GBIF-Occurrence="3063633304" collectingDate="2018-07-21" collectionCode="GUGC" collectorName="Xian-yi Wang" country="China" county="Xinping County" location="Yuxi City" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="84C4773C854DA276F6905B6D64290CBF" type="male">2 ♂♂</specimenCount>
,
<collectingCountry id="2EEDED3A60CD19CD1A4906CFF6163060" name="China">CHINA</collectingCountry>
:
<collectingRegion id="05397187612A577F6B675D02AC9A658B" country="China" name="Yunnan">Yunnan Province</collectingRegion>
,
<location id="3A957F7D31CBE7DB8696EC14FE6A6BD1" LSID="urn:lsid:plazi:treatment:F42E4C2261C55BAF92091B0500F35FCD:3A957F7D31CBE7DB8696EC14FE6A6BD1" country="China" county="Xinping County" name="Yuxi City" stateProvince="Yunnan">Yuxi City</location>
,
<collectingCounty id="C19260914ECC7226AD94C1D3C8244FCC">Xinping County</collectingCounty>
,
<date id="0B4FCE06DB5A21513E7B1E833295C004" value="2018-07-21">
<collectingDate id="37D36DD3369062A1DAA104BB9257D096" value="2018-07-21">21 July 2018</collectingDate>
</date>
, coll.
<collectorName id="6E4D3F3CD37B6E87ECB0BD098C5E5F98">Xian-yi Wang</collectorName>
(
<collectionCode id="E821BA98E4546491A6E333F23225B4D7">GUGC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9D4A92EB9DA4B09229347F29FA953366" pageId="0" pageNumber="49" type="diagnosis">
<paragraph id="D15AC718EAC86FA1D57F59064B63083E" pageId="0" pageNumber="49">Diagnosis.</paragraph>
<paragraph id="6CCAD4F4CDB009E5B337867C5EFD651E" pageId="0" pageNumber="49">
The new species is similar to
<taxonomicName id="93223A819C3EA3905E14E9B8B846CFBC" lsidName="C. harpagota" pageId="0" pageNumber="49" rank="species" species="harpagota">
<emphasis id="E3B4AD60B090501CEEE524AA1A714AB6" italics="true" pageId="0" pageNumber="49">C. harpagota</emphasis>
</taxonomicName>
Zhang, 1994, but differs in having the style apophysis with a subapical spur and the aedeagal shaft with angular projection on the ventral margin in lateral view and with two slender subapical processes.
</paragraph>
</subSubSection>
<subSubSection id="88F8F61DE2135818CE7666C3944B1AF3" pageId="0" pageNumber="49" type="description">
<paragraph id="29CA10C030568679AC64D4AF4DEB204F" pageId="0" pageNumber="49">Description.</paragraph>
<paragraph id="170E9626A3DF28CEE160E3E06E65E7B6" pageId="0" pageNumber="49">
Middle-sized species. Body length (including tegmina): male,
<quantity id="5FA3DE6055FE56E7F2D5B1B124BB68E5" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="7.8" metricValueMin="7.2" unit="mm" value="7.5" valueMax="7.8" valueMin="7.2">7.2-7.8 mm</quantity>
, female,
<quantity id="5D39AC9696577C42B8B55C2423F7E795" metricMagnitude="-3" metricUnit="m" metricValue="8.15" metricValueMax="8.4" metricValueMin="7.9" unit="mm" value="8.15" valueMax="8.4" valueMin="7.9">7.9-8.4 mm</quantity>
.
</paragraph>
<paragraph id="F0D17A756C9B05827698E7F7D7696F86" pageId="0" pageNumber="49">
<emphasis id="7D30DE98ADFA24BFC69E6A8C8251D45C" bold="true" italics="true" pageId="0" pageNumber="49">Coloration.</emphasis>
Ground color brown. Crown yellow with two pairs of brown spots medially, ocelli black (Fig.
<figureCitation id="2EB737C099FE290C0C535F563975407A" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">14</figureCitation>
). Face (Fig.
<figureCitation id="7957027E55D0FC6722BFCAAC49E6ADF3" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">16</figureCitation>
) with clypeus and clypellus black; area between laterofrontal sutures and eye yellow. Pronotum (Fig.
<figureCitation id="BF349224F30106026C222D48284224C2" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">14</figureCitation>
) dark brown, with yellow markings. Mesonotum (Fig.
<figureCitation id="B05911318CC4F84B00916C60A4BE907E" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">14</figureCitation>
) with basal triangles and one round spot on either side of median line anterior to scutoscutellar suture, black. Venation black, with numerous, small, brown spots. Legs (Fig.
<figureCitation id="0A8497187E0AF89C35170F072C1DF39C" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">15</figureCitation>
) dark brown to black.
</paragraph>
<caption id="CD69597AD0AEC1562A43A76B15AD1858" doi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49" start="Figures 14–23" startId="F2">
<paragraph id="C3D49C209C39F854F97896EE53382396" pageId="0" pageNumber="49">
<emphasis id="DCD7BE8806576EE40357EA758F1CBF90" bold="true" pageId="0" pageNumber="49">Figures 14-23.</emphasis>
<taxonomicName id="F065A22710F38C08672EEB93C5D5ADB3" authorityName="Wang &amp; Li &amp; Dai" authorityYear="2021" class="Insecta" family="Cicadellidae" genus="Calodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calodia quadrimacula" order="Hemiptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="quadrimacula">
<emphasis id="D7AAFBAD06D355518F232E8BC08AE1F5" italics="true" pageId="0" pageNumber="49">Calodia quadrimacula</emphasis>
</taxonomicName>
sp. nov.
<specimenCount id="50EF03940C36665AB4EA0BFE28CE8C3F" type="adult">
<emphasis id="47B636D01132420FCAE1590BD32D2BBF" bold="true" pageId="0" pageNumber="49">14</emphasis>
adult
</specimenCount>
, dorsal view
<specimenCount id="D43AD33503CC7D940D9DF99509C7A155" type="adult">
<emphasis id="96827D1C8501D37E24118F14A559317F" bold="true" pageId="0" pageNumber="49">15</emphasis>
adult
</specimenCount>
, lateral view
<emphasis id="C85F8495E878BB4DCF322E914D91A33B" bold="true" pageId="0" pageNumber="49">16</emphasis>
face
<specimenCount id="27606DABA56E6ADADEF28C608EC0E200" type="male">
<emphasis id="0B4D944260B40C8A444174DAFE6CDC76" bold="true" pageId="0" pageNumber="49">17</emphasis>
male
</specimenCount>
pygofer side, lateral view
<emphasis id="6CF9EC2E7ECF2B5E074690666B9DC4C6" bold="true" pageId="0" pageNumber="49">18</emphasis>
connective, dorsal view
<specimenCount id="5EAA8386EA14A14F24F21DCFB4515907" type="male">
<emphasis id="FDB2B30EFE48D1C905E171F269A5BD9A" bold="true" pageId="0" pageNumber="49">19</emphasis>
male
</specimenCount>
subgential plate, ventral view
<emphasis id="07229B171DD2B76B4A8DEE609DB2566E" bold="true" pageId="0" pageNumber="49">20</emphasis>
style, dorsal view (
<collectingRegion id="654120DEF9CB32B3DCCE950EEF334B3F" country="China" name="Yunnan">Yunnan</collectingRegion>
)
<emphasis id="68BB4DF962CAD47204A0F807902AE72A" bold="true" pageId="0" pageNumber="49">21</emphasis>
style, dorsal view (
<collectingRegion id="7DE77C2F2EE37C47C48317819D4961D4" country="China" name="Guizhou">Guizhou</collectingRegion>
)
<emphasis id="B802E75C9DDE610CF461215F9E7AADA6" bold="true" pageId="0" pageNumber="49">22</emphasis>
aedeagus and dorsal connective, dorsal view
<emphasis id="9737A4629594C3EC22CB2E267B05F08D" bold="true" pageId="0" pageNumber="49">23</emphasis>
aedeagus and dorsal connective, lateral view. Scale bars:
<quantity id="840CFBC7EBF2F8F55E65A882D6F73B23" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
(
<emphasis id="D10275F5D1CF9BC56CC4337E245EF717" bold="true" pageId="0" pageNumber="49">14-16</emphasis>
);
<quantity id="F44EAE3B90157CF2F8A15F626CEA8A07" metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
(
<emphasis id="2C1752EDA179F0D0D1DAAA917F295A53" bold="true" pageId="0" pageNumber="49">17-23</emphasis>
).
</paragraph>
</caption>
<paragraph id="960BFE6099E264C1A6ECE4C4084D5205" pageId="0" pageNumber="49">
<emphasis id="38C875947D99E8B5C5318385674E243E" bold="true" italics="true" pageId="0" pageNumber="49">Morphology.</emphasis>
Head narrower than pronotum, anterior margin broadly obtuse; crown as wide as diameter of one eye, produced anteriorly beyond eyes; ocelli on anterior margin of crown; coronal suture extending to level of ocelli (Fig.
<figureCitation id="EA9FAE660ED8E7D828305BF41CC04266" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">14</figureCitation>
); clypeus (Fig.
<figureCitation id="306C013C75718D9C9CE35D99616B8BCB" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">16</figureCitation>
) flat, narrow, laterally expanded under antennal sockets, apex constricted, base inflated longitudinally, apically with lateral margins expanded. Pronotum (Fig.
<figureCitation id="983BA607B2F63D94B082512F5002DE6E" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">14</figureCitation>
) surface bullate. Mesonotum (Figs
<figureCitation id="6778A4676E1A12CE0E894F215204B2C8" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">14</figureCitation>
,
<figureCitation id="37F1DAD5064D00B025B3EE8BE26391CE" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">15</figureCitation>
) nearly as long as pronotum.
</paragraph>
<paragraph id="CD0E36740CF780C9730E3444AAD9CB5E" pageId="0" pageNumber="49">
<emphasis id="883DD3CED536B07B37088BF411700631" bold="true" italics="true" pageId="0" pageNumber="49">Male genitalia.</emphasis>
Pygofer with caudal lobe (Fig.
<figureCitation id="73350C206A0A9FC9B89BBC78A549A8E7" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">17</figureCitation>
) broadly triangular in lateral view. Subgenital plate (Fig.
<figureCitation id="4FDAE49DA271FF24CB9B8EB37429E4DC" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">19</figureCitation>
) long with base slightly broad, sparsely setose apically. Style (Figs
<figureCitation id="76012C75A8D246B4A0BEB1C2A2DB5E6A" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">20</figureCitation>
,
<figureCitation id="175CD8B6DBED6AACA6894520C846F92B" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">21</figureCitation>
) well developed, base expanded, apex bilobed, with a subapical short spur. Connective (Fig.
<figureCitation id="155566ECA991D83C5E9AB98F430662F8" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">18</figureCitation>
) Y-shaped with stem short. Aedeagal shaft (Figs
<figureCitation id="5D3168E5723887C7E5ABC58BBA630254" captionStart="Figures 14–23" captionStartId="F2" captionText="Figures 14 - 23. Calodia quadrimacula sp. nov. 14 adult, dorsal view 15 adult, lateral view 16 face 17 male pygofer side, lateral view 18 connective, dorsal view 19 male subgential plate, ventral view 20 style, dorsal view (Yunnan) 21 style, dorsal view (Guizhou) 22 aedeagus and dorsal connective, dorsal view 23 aedeagus and dorsal connective, lateral view. Scale bars: 1 mm (14 - 16); 0.5 mm (17 - 23)." figureDoi="10.3897/zookeys.1023.59811.figures14-23" httpUri="https://binary.pensoft.net/fig/518693" pageId="0" pageNumber="49">22-23</figureCitation>
) sinuated, ventral margin in lateral view with angular projection slightly distad of half length, curved apically in lateral view, apex with group of spines, with two subapical retrose processes arising on same side, distal process about twice as long as proximal one, with outer margin, serrate proximal process glabrous; gonopore large, subapical, situated laterally more proximal than proximal process.
</paragraph>
</subSubSection>
<subSubSection id="A7F30ACBC235B8B9631F3A0F2F97C542" pageId="0" pageNumber="49" type="etymology">
<paragraph id="71AAA24CA87B4BD57CF4A90DA583F8F8" pageId="0" pageNumber="49">Etymology.</paragraph>
<paragraph id="002CA4E53133B21EF52AD2414735282D" pageId="0" pageNumber="49">
The new species name is derived from the words &quot;
<emphasis id="9D585A9DD0BD4FAF65E6F64BDFDF3B00" italics="true" pageId="0" pageNumber="49">quadri</emphasis>
&quot; and &quot;
<emphasis id="BD6A41AED9BB0313A9F98340692BB8A6" italics="true" pageId="0" pageNumber="49">macula</emphasis>
&quot;, referring to the scutellum with four black plaques.
</paragraph>
</subSubSection>
<subSubSection id="A5FB5EEFEF4C3B10DF8509136EFC1865" pageId="0" pageNumber="49" type="distribution">
<paragraph id="4EB8A60BC298A5E11AE107830A158490" pageId="0" pageNumber="49">Distribution.</paragraph>
<paragraph id="A0EC3BC2514E0E4DD00DF27E3D2F4F7F" pageId="0" pageNumber="49">
<materialsCitation id="39A4619D79CB3040534B93B94A108EC6" ID-GBIF-Occurrence="3063633301" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="China" location="Guizhou" specimenCount="1" stateProvince="Guizhou">
<collectingCountry id="6420C36571C6053702F13E86A793E3A0" name="China">China</collectingCountry>
(
<collectingRegion id="1D32491956396A5A4312F27A7C3A78F2" country="China" name="Guizhou">Guizhou</collectingRegion>
,
<collectingRegion id="9E115F5872BDBA8F45F597554913A8C9" country="China" name="Yunnan">Yunnan</collectingRegion>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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