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<document ID-CLB-Dataset="26430" ID-DOI="http://dx.doi.org/10.3897/phytokeys.124.34679" ID-GBIF-Dataset="0ed8fe19-ae7d-4551-b04c-40506ffc4274" ID-PMC="PMC6598937" ID-Pensoft-Pub="1314-2003-124-87" ID-Pensoft-UUID="CD370A37FFD8203C596C8A6F9E10FFC0" ID-PubMed="31274979" ID-Zenodo-Dep="3261874" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2003-124-87" ModsDocOrigin="PhytoKeys 124" ModsDocTitle="Insights on the systematics and morphology of Humiriaceae (Malpighiales): androecial and extrafloral nectary variation, two new combinations, and a new Sacoglottis from Guyana" checkinTime="1561202160728" checkinUser="pensoft" docAuthor="Wurdack, Kenneth J. &amp; Zartman, Charles E." docDate="2019" docId="F410C88615ABA3439E006A673192B91F" docLanguage="en" docName="PhytoKeys 124: 87-121" docOrigin="PhytoKeys 124" docSource="http://dx.doi.org/10.3897/phytokeys.124.34679" docTitle="Vantanea spiritu-sancti K. Wurdack &amp; C. E. Zartman 2019, comb. nov." docType="treatment" docVersion="8" id="CD370A37FFD8203C596C8A6F9E10FFC0" lastPageNumber="97" masterDocId="CD370A37FFD8203C596C8A6F9E10FFC0" masterDocTitle="Insights on the systematics and morphology of Humiriaceae (Malpighiales): androecial and extrafloral nectary variation, two new combinations, and a new Sacoglottis from Guyana" masterLastPageNumber="121" masterPageNumber="87" pageNumber="97" updateTime="1732825798615" updateUser="ExternalLinkService">
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<mods:title id="8F1C30C266171B1F7256B8490673BA8F">Insights on the systematics and morphology of Humiriaceae (Malpighiales): androecial and extrafloral nectary variation, two new combinations, and a new Sacoglottis from Guyana</mods:title>
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<mods:namePart id="55D964E740B074B4F065FCA6AD79E933">Wurdack, Kenneth J.</mods:namePart>
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<mods:namePart id="5139613F1CEB69B4B6846FFC38937597">Zartman, Charles E.</mods:namePart>
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<mods:date id="1D7426A0B960339079AFA3B9A6E69B6B">2019</mods:date>
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<mods:identifier id="0D95DB71BD4362E1C02832F66FA36764" type="DOI">http://dx.doi.org/10.3897/phytokeys.124.34679</mods:identifier>
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<treatment id="F410C88615ABA3439E006A673192B91F" ID-GBIF-Taxon="158363992" LSID="urn:lsid:plazi:treatment:F410C88615ABA3439E006A673192B91F" httpUri="http://treatment.plazi.org/id/F410C88615ABA3439E006A673192B91F" lastPageNumber="97" pageId="10" pageNumber="97" scope_family="Humiriaceae" scope_order="Malpighiales">
<subSubSection id="35E18BDBECDE3B8F095C6F85A801224A" pageId="10" pageNumber="97" type="nomenclature">
<paragraph id="905E71AE14499BD6660DFB6C7976F637" pageId="10" pageNumber="97">
<taxonomicName id="7958A43500AE4FB4A6E0F06753BF68BF" ID-CoL="7FGJR" LSID="5DD004A4-DCF8-5203-A52C-117A6609EF30" authority="(Cuatrec.) K. Wurdack &amp; C. E. Zartman" authorityName="K. Wurdack &amp; C. E. Zartman" authorityYear="2019" baseAuthorityName="K. Wurdack &amp; C. E. Zartman" baseAuthorityYear="2019" class="Magnoliopsida" family="Humiriaceae" genus="Vantanea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vantanea spiritu-sancti" order="Malpighiales" pageId="10" pageNumber="97" phylum="Tracheophyta" rank="species" species="spiritu-sancti" status="comb. nov.">
<pageBreakToken id="4F6564B9BB46CAB1F400FFB7A317355C" pageId="10" pageNumber="97" start="start">Vantanea</pageBreakToken>
spiritu-sancti (Cuatrec.) K.Wurdack &amp; C.E.Zartman
</taxonomicName>
<taxonomicNameLabel id="97628F7382158FE207FC7F352C886134" pageId="10" pageNumber="97">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="71601BCBC09C8F37273684FAE1C152C1" pageId="10" pageNumber="97" type="reference_group">
<paragraph id="33DE0C605BF9D490C5D47E949035AEC9" pageId="10" pageNumber="97">
<taxonomicName id="0A8EDC260F1F94C4F301BA34281ECB93" class="Magnoliopsida" family="Humiriaceae" genus="Humiriastrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Humiriastrum spiritu-sancti" order="Malpighiales" pageId="10" pageNumber="97" phylum="Tracheophyta" rank="species" species="spiritu-sancti">Humiriastrum spiritu-sancti</taxonomicName>
Cuatrec., Ciencia, Mexico 23(4): 137. 1964. Type: BRAZIL.
<normalizedToken id="42B7F1137EF4F2E109039D8C1A866DB1" originalValue="Espírito">Espirito</normalizedToken>
Santo, Mun. de Santa Tereza: Lombardia, 25 Jan 1954,
<emphasis id="720095EFA017F24DC79EACFE5E100DD8" italics="true" pageId="10" pageNumber="97">G. Dalcolmo s.n.</emphasis>
(holotype: RB-86212; isotype: US-2827596).
</paragraph>
</subSubSection>
<subSubSection id="89970D9D2BAFA7ACDA1DEA3F7CFF0C7E" pageId="10" pageNumber="97" type="notes">
<paragraph id="B895C73F91EFE364DFA37EAF95CAE535" pageId="10" pageNumber="97">Notes.</paragraph>
<paragraph id="BA002D74FA03096773438F7A3CB965B7" pageId="10" pageNumber="97">
<taxonomicName id="135C77B6165CA7918FF922BB3342C8BB" class="Magnoliopsida" family="Humiriaceae" genus="Humiriastrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Humiriastrum spiritu-sancti" order="Malpighiales" pageId="10" pageNumber="97" phylum="Tracheophyta" rank="species" species="spiritu-sancti">
<emphasis id="912A771399B5C39347FA789EF570F1D8" italics="true" pageId="10" pageNumber="97">Humiriastrum spiritu-sancti</emphasis>
</taxonomicName>
was poorly known until recent ample collections allowed a fuller characterization.
<bibRefCitation id="9E0B90897229BD77A262089CBF522CB2" DOI="https://doi.org/10.1590/2175-7860200859109" author="Giordano, LCS" journalOrPublisher="Rodriguesia" pageId="21" pageNumber="108" pagination="151 - 154" refId="B13" refString="Giordano, LCS, Bove, CP, 2008. Taxonomic considerations and amended description of Humiriastrum spiritu-sancti, Humiriaceae. Rodriguesia 59 (1): 151 - 154, DOI: https://doi.org/10.1590/2175-7860200859109" title="Taxonomic considerations and amended description of Humiriastrum spiritu-sancti, Humiriaceae." url="https://doi.org/10.1590/2175-7860200859109" volume="59" year="2008">Giordano and Bove (2008)</bibRefCitation>
described its thecae as unilocular, ovary cells as uniovulate, and fruits as having five apical foramina. The transfer here to
<taxonomicName id="3B82FDCF33F14720192C8C0B929C10FC" authorityName="Aublet" authorityYear="1775" class="Magnoliopsida" family="Humiriaceae" genus="Vantanea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vantanea" order="Malpighiales" pageId="10" pageNumber="97" phylum="Tracheophyta" rank="genus">
<emphasis id="F7474FD3BD66FE70E0B36F0334B0D67D" italics="true" pageId="10" pageNumber="97">Vantanea</emphasis>
</taxonomicName>
is supported by anther structure that we interpret as two lateral bisporangiate thecae with internal septa (Fig.
<figureCitation id="79F9C33001ADDD62E1C5939545E1F0A0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Stamen structure and anatomy of Humiriaceae. A Vantanea compacta anther, lateral B Vantanea compacta anther, dorsal C Vantanea depleta anther, lateral D Vantanea spiritu-sancti anther, lateral E Schistostemon oblongifolius clearing of androecium with 3 stamen types, ventral F Vantanea spiritu-sancti longitudinal section of ovary with 2 superposed ovules per locule; lower left ovule partial G Hylocarpa heterocarpa anthers (left to right): disporangiate dorsal, ventral; sterile, dorsal. Sources: A, B Hatschbach 21265 C Hammel &amp; Trainer 12954 D, F Silva et al. 1436 E Maas et al. 6804 G Ducke [JBRJ- 30137] (all US)." figureDoi="10.3897/phytokeys.124.34679.figure3" httpUri="https://binary.pensoft.net/fig/311889" pageId="10" pageNumber="97">3D</figureCitation>
; see Results), two superposed ovules per locule (Fig.
<figureCitation id="5DB965DF88C0DC5DE675A4717F3F6EFF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Stamen structure and anatomy of Humiriaceae. A Vantanea compacta anther, lateral B Vantanea compacta anther, dorsal C Vantanea depleta anther, lateral D Vantanea spiritu-sancti anther, lateral E Schistostemon oblongifolius clearing of androecium with 3 stamen types, ventral F Vantanea spiritu-sancti longitudinal section of ovary with 2 superposed ovules per locule; lower left ovule partial G Hylocarpa heterocarpa anthers (left to right): disporangiate dorsal, ventral; sterile, dorsal. Sources: A, B Hatschbach 21265 C Hammel &amp; Trainer 12954 D, F Silva et al. 1436 E Maas et al. 6804 G Ducke [JBRJ- 30137] (all US)." figureDoi="10.3897/phytokeys.124.34679.figure3" httpUri="https://binary.pensoft.net/fig/311889" pageId="10" pageNumber="97">3F</figureCitation>
), apparent lack of foramina, as well as previously reported pollen details (Bove and Melham 2000), and preliminary molecular phylogenetic placement (Zartman et al., unpublished data). Upon examination of endocarps from
<emphasis id="1678092E537E3B7E7E8BC9A670B23BD3" italics="true" pageId="10" pageNumber="97">Amorim et al. 1391</emphasis>
(US-3258470, as sectioned for
<bibRefCitation id="78C101185152FABC0AC2E827AB77129E" DOI="https://doi.org/10.1086/651229" author="Herrera, F" journalOrPublisher="International Journal of Plant Sciences" pageId="21" pageNumber="108" pagination="392 - 408" refId="B15" refString="Herrera, F, Manchester, SR, Jaramillo, C, MacFadden, B, da, Silva-Caminha SA, 2010. Phytogeographic history and phylogeny of the Humiriaceae. International Journal of Plant Sciences 171 (4): 392 - 408, DOI: https://doi.org/10.1086/651229" title="Phytogeographic history and phylogeny of the Humiriaceae." url="https://doi.org/10.1086/651229" volume="171" year="2010">Herrera et al. 2010</bibRefCitation>
), as well as the illustration from
<bibRefCitation id="3AA61BE7BD20A7BAFBCA6D91F16283FD" DOI="https://doi.org/10.1590/2175-7860200859109" author="Giordano, LCS" journalOrPublisher="Rodriguesia" pageId="21" pageNumber="108" pagination="151 - 154" refId="B13" refString="Giordano, LCS, Bove, CP, 2008. Taxonomic considerations and amended description of Humiriastrum spiritu-sancti, Humiriaceae. Rodriguesia 59 (1): 151 - 154, DOI: https://doi.org/10.1590/2175-7860200859109" title="Taxonomic considerations and amended description of Humiriastrum spiritu-sancti, Humiriaceae." url="https://doi.org/10.1590/2175-7860200859109" volume="59" year="2008">Giordano and Bove (2008)</bibRefCitation>
, we could not find well-developed apical foramina that otherwise characterize
<taxonomicName id="EAEB371BB94C6E0A10C36E90B510CD67" authorityName="Cuatrecasas" authorityYear="1961" baseAuthorityName="Urb." class="Magnoliopsida" family="Humiriaceae" genus="Humiriastrum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Humiriastrum" order="Malpighiales" pageId="10" pageNumber="97" phylum="Tracheophyta" rank="genus">
<emphasis id="A8067154DB62A34BE8A14744E4DA3158" italics="true" pageId="10" pageNumber="97">Humiriastrum</emphasis>
</taxonomicName>
. Furthermore, the androecial bauplan of
<taxonomicName id="B69DC48D6FDB8C409AAFC2659A80FB9A" class="Dicotyledoneae" family="Humiriaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="V. spiritu-sancti" order="Malpighiales" pageId="10" pageNumber="97" phylum="Angiospermae" rank="species" species="spiritu-sancti">
<emphasis id="69B78969A192677F479E2AAC2B39C8DA" italics="true" pageId="10" pageNumber="97">V. spiritu-sancti</emphasis>
</taxonomicName>
with a low number of alternating-length stamens is similar to that of
<taxonomicName id="39FDF40808EE2E17FFDFD1EA15238978" class="Dicotyledoneae" family="Humiriaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="V. depleta" order="Malpighiales" pageId="10" pageNumber="97" phylum="Angiospermae" rank="species" species="depleta">
<emphasis id="2D821134CB84150B49DA2B1A1F1EA2C0" italics="true" pageId="10" pageNumber="97">V. depleta</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="0C7AA8E93B2F06C4928A403504CB0724" ID-Zenodo-Dep="3262018" doi="10.3897/phytokeys.124.34679.figure7" httpUri="https://binary.pensoft.net/fig/311893" pageId="10" pageNumber="97" start="Figure 7" startId="F7">
<paragraph id="8A46031F9E38FBED301F5885FF57B057" pageId="10" pageNumber="97">
<emphasis id="050092E225CBCF994C5843E3F7F748B4" bold="true" pageId="10" pageNumber="97">Figure 7.</emphasis>
Illustration of
<taxonomicName id="C252687ACE4E55B09495269DB0962A06" authorityName="K. Wurdack &amp; C. E. Zartman" authorityYear="2019" class="Magnoliopsida" family="Humiriaceae" genus="Sacoglottis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sacoglottis perryi" order="Malpighiales" pageId="10" pageNumber="97" phylum="Tracheophyta" rank="species" species="perryi">
<emphasis id="4033F6BB3908BB445DE81CDCE3A8EC98" italics="true" pageId="10" pageNumber="97">Sacoglottis perryi</emphasis>
</taxonomicName>
.
<emphasis id="FE94051EF5FEA45DE46C2B9C5011B95F" bold="true" pageId="10" pageNumber="97">A</emphasis>
Habit
<emphasis id="482B8F548C0AC58115BE7EB5AFA8729B" bold="true" pageId="10" pageNumber="97">B</emphasis>
bud
<emphasis id="AB5AC95FFD24862BDC2DED3523CFA37D" bold="true" pageId="10" pageNumber="97">C</emphasis>
bud with petals removed
<emphasis id="EB46DDC83C5C6967071F20B64F34E751" bold="true" pageId="10" pageNumber="97">D</emphasis>
open flower, axial
<emphasis id="1BC09A4BCE123A8EA3CDD828E47D5207" bold="true" pageId="10" pageNumber="97">E</emphasis>
open flower, lateral
<emphasis id="9E1CB564FC6166D53FA8F971418E2AEF" bold="true" pageId="10" pageNumber="97">F</emphasis>
gynoecium
<emphasis id="0F8BC2A9488637CE2609AB9D7EA20C3F" bold="true" pageId="10" pageNumber="97">G</emphasis>
stamen cluster of 2 types, ventral
<emphasis id="7EF840D0741746510D33781FE7488B6C" bold="true" pageId="10" pageNumber="97">H</emphasis>
floral diagram
<emphasis id="3EB2ED7E45214072492106DFDF554594" bold="true" pageId="10" pageNumber="97">I</emphasis>
post-anthetic flower
<emphasis id="2C61342EB876086E44AD06098A538A98" bold="true" pageId="10" pageNumber="97">J</emphasis>
young fruit
<emphasis id="58041ED6ADC745BBB40C33DE2817E89F" bold="true" pageId="10" pageNumber="97">K, L</emphasis>
fruit
<emphasis id="88FDA8233952BCA3ADA04DF9FD758914" bold="true" pageId="10" pageNumber="97">M</emphasis>
fruit with 2 central seeds and endocarp lacunae, transverse section. Source:
<emphasis id="3E76B05CAF422416989FDECBCC34A762" bold="true" pageId="10" pageNumber="97">
<normalizedToken id="5007B0A3A419DE49203928634BF83224" originalValue="AM">A-M</normalizedToken>
</emphasis>
from specimens and life photos of
<emphasis id="6E8C1F4A2D02BC98AB257F3F51D06905" italics="true" pageId="10" pageNumber="97">Redden 7264</emphasis>
(US).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>