188 lines
20 KiB
XML
188 lines
20 KiB
XML
<document id="C0A627CB640BA78D0D97858E8C1E2431" ID-DOI="10.3897/zookeys.1223.139346" ID-ZooBank="451749E9-009E-43E9-A6F1-892035BDF1B0" ID-publisher-id="139346" URI-arpha="6EA2BAFE-82AB-5570-95B8-EE1F5FA60D46" URI-zoobank="http://zoobank.org/451749E9-009E-43E9-A6F1-892035BDF1B0" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1736349736103" checkinUser="pensoft" docAuthor="Wesener, Thomas, Akkari, Nesrine & Golovatch, Sergei I." docDate="2025" docId="F2E9C6692D455CED8CBEECC193233FE4" docLanguage="en" docName="ZooKeys 1223: 185-220" docOrigin="ZooKeys 1223" docSource="https://zookeys.pensoft.net/article/139346/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Dalodesmus Cook 1896" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="220" masterDocId="6EA2BAFE82AB557095B8EE1F5FA60D46" masterDocTitle="Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida)" masterLastPageNumber="220" masterPageNumber="185" pageNumber="185" updateTime="1736355520980" updateUser="admin">
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<mods:titleInfo id="EA8DF083DB953545F0099E569F03B380">
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<mods:title id="415A132BEF15667B8BF1419A5100899B">Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida)</mods:title>
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<mods:name id="786CB8F7FEEA09B5968AEEFF6F1E12B0" type="personal">
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<mods:roleTerm id="BE5D1FD801D075A1F833D62F59A049E2">Author</mods:roleTerm>
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<mods:namePart id="A7EEB081D46961B605A2FF61A49AC58B">Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier id="85CC535FF8C583F9C50B5A46F4CBE61D" type="ORCID">0000-0002-2028-3541</mods:nameIdentifier>
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<mods:affiliation id="507E0D86C20A9C7453F148BF531B245F">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, D- 53113 Bonn, Germany</mods:affiliation>
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<mods:name id="90D102A24D2C93D9D98C323B9EF5C44C" type="personal">
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<mods:roleTerm id="EB919EA9D921AB5ADC6AE5580B66CE49">Author</mods:roleTerm>
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<mods:namePart id="2745D1EFBA687F6528C3BCB039706EC9">Akkari, Nesrine</mods:namePart>
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<mods:nameIdentifier id="949F402FDBF160341FA0A6F80E2356FF" type="ORCID">0000-0001-5019-4833</mods:nameIdentifier>
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<mods:affiliation id="6C9A9355047FD8737C2DAC8FC5343590">Third Zoological Department, Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:roleTerm id="639808B9AC777BAC2C903180912867F2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="87FE0C099EC3EA0AD7869FFF6142C32B">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="6491217BF7CC3F463DCD6249CB9DD7A4">Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow 119071, Russia</mods:affiliation>
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<mods:typeOfResource id="1547D8A08377361F63A00C3335730E4D">text</mods:typeOfResource>
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<mods:title id="2D987BB02583EA322BB31FE6475F1961">ZooKeys</mods:title>
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<mods:part id="933F4DA3101B234883E430CE7DF480A7">
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<mods:date id="5E0692F74663D66E3B5201EADFD389DF">2025</mods:date>
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<mods:number id="36FC9AB7058E41B1A8DB389F319C2AEE">2025-01-08</mods:number>
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<mods:detail id="A3BAA85F875C387991EDD0BC0C1B838D" type="volume">
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<mods:number id="5C6BFF4BE33F134C882A0A4F037B2024">1223</mods:number>
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<mods:start id="A8EA6552C6DC773E31C345B91A046AE1">185</mods:start>
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<mods:classification id="AB3D4171F0CF1C25238DE8596F46A831">journal article</mods:classification>
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<mods:identifier id="AC9ACE4E550A30642622F98434E25735" type="DOI">10.3897/zookeys.1223.139346</mods:identifier>
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<mods:identifier id="DDEA3BDD600E05FCB7CA1795F4625CCE" type="ZooBank">451749E9-009E-43E9-A6F1-892035BDF1B0</mods:identifier>
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<treatment id="F2E9C6692D455CED8CBEECC193233FE4" ID-DOI="http://doi.org/10.5281/zenodo.14616983" ID-Zenodo-Dep="14616983" ID-arpha="F2E9C669-2D45-5CED-8CBE-ECC193233FE4" LSID="urn:lsid:plazi:treatment:F2E9C6692D455CED8CBEECC193233FE4" httpUri="http://treatment.plazi.org/id/F2E9C6692D455CED8CBEECC193233FE4" scope_class="Diplopoda" scope_family="Dalodesmidae" scope_order="Polydesmida">
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<subSubSection id="601B2BB36554ACD825CEC370CBB3D9FC" type="nomenclature">
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<paragraph id="176384DED8861D4646677FBBC5502E06">
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||
<heading id="34445BB92BB32FC4FF6DB2DE57F5374B" reason="label">Genus</heading>
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||
<taxonomicName id="09C981531FAF7D09FACEB2A4B7607751" ID-CoL="7P7Z8" authority="Cook, 1896" authorityName="Cook" authorityYear="1896" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
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<emphasis id="672D8B9D000B58159753DEBE3C2C8BA7" italics="true">Dalodesmus</emphasis>
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<bibRefCitation id="B388CB0C4D19A24BCC81742C1B301249" author="Cook OF" journalOrPublisher="Brandtia" pagination="19-28" refId="B6" refString="Cook OF (1896) Cryptodesmus and its allies. Brandtia 5: 19 - 28." title="Cryptodesmus and its allies." volume="5" year="1896">Cook, 1896</bibRefCitation>
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</taxonomicName>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="E2801F4BD7622402C32746DD6B137BC0" type="reference_group">
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<paragraph id="2AAAB149FFC496D2DC6138DB6C21624B">
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||
<treatmentCitationGroup id="42F41E45E2AF9869C19519A44689E5A2">
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<treatmentCitation id="DD5C97A0F254DECB1A6C159CDBCA6425">
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<taxonomicName id="0B5BD5D0B856BC25F37BAA1DCE2A9C60" ID-CoL="7P7Z8" authority="Cook, 1896" authorityName="Cook" authorityYear="1896" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
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<emphasis id="4E2DE36F025E4DBC62C4953B87DB39DE" italics="true">Dalodesmus</emphasis>
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||
<bibRefCitation id="30B864255F4E41BD60DCE82C0D919689" author="Cook OF" journalOrPublisher="Brandtia" pagination="19-28" refId="B6" refString="Cook OF (1896) Cryptodesmus and its allies. Brandtia 5: 19 - 28." title="Cryptodesmus and its allies." volume="5" year="1896">Cook, 1896</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
; type species
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<taxonomicName id="0B6FD5C1829C306D2A17C6810FB5A9D2" ID-CoL="6CBHJ" authority="Cook, 1896" authorityName="Cook" authorityYear="1896" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="tectus">
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<emphasis id="3DA7ACEBD55523B7EA9D5661D822D357" italics="true">D. tectus</emphasis>
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<treatmentCitation id="D9C3580CD19A45996FBF4803A70985EC">
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||
<bibRefCitation id="104B0909D0F1762DC4849853DA444079" author="Cook OF" journalOrPublisher="Brandtia" pagination="19-28" refId="B6" refString="Cook OF (1896) Cryptodesmus and its allies. Brandtia 5: 19 - 28." title="Cryptodesmus and its allies." volume="5" year="1896">Cook, 1896</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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, by monotypy.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="F6A8DC14E386CD3ADF50302C2C2E2C35">
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||
<treatmentCitationGroup id="D20FDA2348F3A8A5F5A10D0F516201FE">
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||
<treatmentCitation id="83CE1AB833AD1DC4317857D3DC18DFC7">
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||
<taxonomicName id="35A114707A5C63877A3D7D62AAF2D59E" ID-CoL="BFHCG" authority="Attems, 1898" authorityName="Attems" authorityYear="1898" class="Diplopoda" family="Dalodesmidae" genus="Tubercularium" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
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<emphasis id="B00E9AA19AC799295149D54937DFC086" italics="true">Tubercularium</emphasis>
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<bibRefCitation id="4E3ECE11B11FD791513339197AC4F697" author="Attems C" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe" pagination="221-482" refId="B1" refString="Attems C (1898) System der Polydesmiden. I. Theil. Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe 67: 221 - 482." title="System der Polydesmiden. I. Theil." volume="67" year="1898">Attems, 1898</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
; type species:
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<taxonomicName id="B6FFA11F8358BE97494B3D8EC047579E" ID-CoL="59J9Y" authority="Attems, 1898" authorityName="Attems" authorityYear="1898" class="Diplopoda" family="Dalodesmidae" genus="Tubercularium" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="odontopezum">
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||
<emphasis id="65E01BCE6DFB66E3D84D9E2BF5449069" italics="true">T. odontopezum</emphasis>
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||
<treatmentCitation id="8A1E302DDEEA4844A4151B9DF126C53E">
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||
<bibRefCitation id="B7918F0773784F6CEDEA4C28F20EA758" author="Attems C" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe" pagination="221-482" refId="B1" refString="Attems C (1898) System der Polydesmiden. I. Theil. Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, Mathematisch-Naturwissenschaftliche Classe 67: 221 - 482." title="System der Polydesmiden. I. Theil." volume="67" year="1898">Attems, 1898</bibRefCitation>
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||
</treatmentCitation>
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||
</taxonomicName>
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||
, by monotypy, synonymised by
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||
<treatmentCitation id="2CEF67481F5160EB68C85EE99E3D2B9E">
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||
<bibRefCitation id="236F877F3A958532CC0096486C09C571" author="Jeekel CAW" firstAuthor="Jeekel" journalOrPublisher="Entomologische Berichten" pagination="236-239" refId="B17" refString="Jeekel CAW (1965) The identity of Dalodesmus tectus Cook, 1896, and the status of the family names Dalodesmidae Cook, 1896, Vanhoeffeniidae Attems, 1914 and Sphaeriotrichopodidae Attems, 1914 (Diplopoda, Polydesmida). Entomologische Berichten 25: 236–239." title="The identity of Dalodesmus tectus Cook, 1896, and the status of the family names Dalodesmidae Cook, 1896, Vanhoeffeniidae Attems, 1914 and Sphaeriotrichopodidae Attems, 1914 (Diplopoda, Polydesmida)." volume="25" year="1965">Jeekel (1965)</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="1BB0D6CE3A7B56AEBAB3FDECF396F048">
|
||
<treatmentCitationGroup id="CE36630575F03E60CAA2C44D9B25B2F2">
|
||
<treatmentCitation id="1CA46B9307785E6B9561A2B9CEE6E0BA">
|
||
<taxonomicName id="BA9F96F6DFEC07BC2513CEF601742C65" ID-CoL="7PWX5" authority="de Saussure & Zehntner, 1901" authorityName="de Saussure & Zehntner" authorityYear="1901" class="Diplopoda" family="Cryptodesmidae" genus="Pterodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
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||
<emphasis id="7CA834D7B84F55C825C1E5A27D2D9518" italics="true">Pterodesmus</emphasis>
|
||
<bibRefCitation id="3CE18C8C41AE74840E6D4E1C78B53317" author="de Saussure H & Zehntner L" issue="4" journalOrPublisher="Abhandlungen der Senckenbergischen naturforschenden Gesellschaft" pagination="429-460" refId="B8" refString="de Saussure H, Zehntner L (1901) Myriopoden aus Madagaskar und Zanzibar, gesammelt von Dr. A. Voeltzkow. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft 26 (4): 429 - 460." title="Myriopoden aus Madagaskar und Zanzibar, gesammelt von Dr. A. Voeltzkow." volume="26" year="1901">de Saussure & Zehntner, 1901</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
; invalidly proposed without a type species, also preoccupied by
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||
<taxonomicName id="873DA0A98DBD681E672A78E7D64EED84" ID-CoL="7PWX5" authority="Cook, 1896" authorityName="Cook" authorityYear="1896" class="Diplopoda" family="Cryptodesmidae" genus="Pterodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
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||
<emphasis id="AAC8D0B558E6ED37166D287DC7585149" italics="true">Pterodesmus</emphasis>
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||
<treatmentCitation id="C26479D3F53F80BD7C944F21EA7977FC">
|
||
<bibRefCitation id="0C3116652B37CF657AAA1AF9F3A534A7" author="Cook OF" journalOrPublisher="Brandtia" pagination="19-28" refId="B6" refString="Cook OF (1896) Cryptodesmus and its allies. Brandtia 5: 19 - 28." title="Cryptodesmus and its allies." volume="5" year="1896">Cook, 1896</bibRefCitation>
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||
</treatmentCitation>
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||
</taxonomicName>
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||
; synonymised by
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<treatmentCitation id="1975C59FD1F09C284645F406DB389EAF">
|
||
<bibRefCitation id="79229BEF6C5F198C24D6BA1A3CC6D7BA" author="Jeekel CAW" firstAuthor="Jeekel" journalOrPublisher="Entomologische Berichten" pagination="236-239" refId="B17" refString="Jeekel CAW (1965) The identity of Dalodesmus tectus Cook, 1896, and the status of the family names Dalodesmidae Cook, 1896, Vanhoeffeniidae Attems, 1914 and Sphaeriotrichopodidae Attems, 1914 (Diplopoda, Polydesmida). Entomologische Berichten 25: 236–239." title="The identity of Dalodesmus tectus Cook, 1896, and the status of the family names Dalodesmidae Cook, 1896, Vanhoeffeniidae Attems, 1914 and Sphaeriotrichopodidae Attems, 1914 (Diplopoda, Polydesmida)." volume="25" year="1965">Jeekel (1965)</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0EHLAG" type="notes">
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<paragraph id="375CE1F3C95209F43C5357C1063D1322">
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<heading id="F486E1089571C1E9B30F0526B7A63C87" reason="title">Note.</heading>
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||
</paragraph>
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<paragraph id="C8562AA80C8361B40C7062C085254FCB">Species included: 6 (including two new described below).</paragraph>
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</subSubSection>
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||
<subSubSection id="SECID0EMLAG" type="diagnosis">
|
||
<paragraph id="60AA94EEF93804D80617F7322CC8B714">
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||
<heading id="1ACBF0B7DD2602C3F923FFC3A784606C" reason="title">Diagnosis.</heading>
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||
</paragraph>
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||
<paragraph id="17EE765293F474EB2B7CBEB8B841101E">
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||
Body medium-sized,
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||
<quantity id="2F0DF22DB294A8C359F063B7A979C2CD" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.8" metricValueMin="1.7" unit="mm" value="22.5" valueMax="28.0" valueMin="17.0">17–28 mm</quantity>
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||
long and
|
||
<quantity id="EFB83A212449DF97C305813CE243FEE6" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.3" metricValueMin="3.7" unit="mm" value="4.5" valueMax="5.3" valueMin="3.7">3.7–5.3 mm</quantity>
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||
wide. Midbody metaterga with 4–6 transverse, often rather irregular rows of piligerous tuberculations or obliterated areations between always strongly developed and laterally similarly piligerous crenulate / tuberculate paraterga, these being sub-horizontal to upturned. Gonopods less elaborate and more simple than in the other genera, both coxites and densely setose prefemorites equally short and fused medially; femorites (fe) mostly slender, subcylindrical, at most only insignificantly flattened sagittally, contiguous medially and fused in basal 1 / 3, with a sternal rudiment visible ventrally at the very base, bare to sometimes sparsely setose until acropodite, the latter (= acropodite) basically tripartite, a mostly distinct, occasionally branching and only rarely missing solenomere branch (sl) lying between and typically flanked by two or three branches of a rather simple to elaborate solenophore: one medial (mb), this only rarely subdivided into an apical (ab) and a subapical branch (sb), and the other lateral (lb).
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||
</paragraph>
|
||
<paragraph id="9B69B9BD516B4559F977B5B4CBB2DB45">
|
||
<taxonomicName id="C64A848A4F36683B3C9C27C1D2458687" authorityName="Cook" authorityYear="1896" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
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||
<emphasis id="433E40BB5BAC73767FA0AAE1F398B4C7" italics="true">Dalodesmus</emphasis>
|
||
</taxonomicName>
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||
differs from
|
||
<taxonomicName id="6941053B3E88B8314F168499A3A8308C" authorityName="de Saussure & Zehntner" authorityYear="1902" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
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||
<emphasis id="9E792F2DE5DFE75D328CF112AFA75430" italics="true">Phymatodesmus</emphasis>
|
||
</taxonomicName>
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||
in the much larger size (>
|
||
<quantity id="C08F11044CC413E0D24066BAEB32D474" metricMagnitude="-2" metricUnit="m" metricValue="2.0" unit="mm" value="20.0">20 mm</quantity>
|
||
, vs ~
|
||
<quantity id="AF5364FBD13E39E9E6C379A73A65CCED" metricMagnitude="-2" metricUnit="m" metricValue="1.0" unit="mm" value="10.0">10 mm</quantity>
|
||
), the strongly developed, apically pointed paratergites (
|
||
<emphasis id="AFA7DE30C7733AC3DB44A001D3E4FFB8" italics="true">vs</emphasis>
|
||
short and rectangular in
|
||
<taxonomicName id="612394F7EB942356C3BEEB632880EA9F" authorityName="de Saussure & Zehntner" authorityYear="1902" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="95AF2C4332401C9080E92993FFE0B1ED" italics="true">Phymatodesmus</emphasis>
|
||
</taxonomicName>
|
||
), the presence of large, oval to polygonal, often irregular, piligerous tuberculations or areations (vs circular cones / tuberculations in
|
||
<taxonomicName id="53ED9ED7E8E9EA27401A7A51D2A9BC22" authorityName="de Saussure & Zehntner" authorityYear="1902" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="97A0BEC28271849686008994DE6A9214" italics="true">Phymatodesmus</emphasis>
|
||
</taxonomicName>
|
||
), and the 2 + 2 setae on the paraprocts not being borne on distinct knobs (vs borne on distinct knobs in
|
||
<taxonomicName id="64AD9BA2DFE173A8BD8B2EC2D0F4881C" authorityName="de Saussure & Zehntner" authorityYear="1902" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2F682A016FD24DDDB8A9EF794AB69DCA" italics="true">Phymatodesmus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="91E076A7ED92D8BCB4AEF6CE37BC37A3">
|
||
In gonopodal structure, the genus
|
||
<taxonomicName id="718F8E2E325B22B4C210EB2331ADD745" authorityName="Cook" authorityYear="1896" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CE56CF8024F37B954ED2494633F1FC12" italics="true">Dalodesmus</emphasis>
|
||
</taxonomicName>
|
||
differs from both
|
||
<taxonomicName id="28C5657106C320745FDF195DE48569C7" authorityName="de Saussure & Zehntner" authorityYear="1902" class="Diplopoda" family="Dalodesmidae" genus="Phymatodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CC3ECE77DECAF8D0CE300D09C1390127" italics="true">Phymatodesmus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="24764878454949810EE9484E61142E1D" authorityName="Brolemann" authorityYear="1916" class="Diplopoda" family="Dalodesmidae" genus="Eutubercularium" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3CB17467A2A7B7EDE1AB4FAB47EFACC3" italics="true">Eutubercularium</emphasis>
|
||
</taxonomicName>
|
||
in the femorites being long and slender,> 2 × as long as acropodites, subcylindrical, diverging in distal 2 / 3 to 1 / 2, bare to only poorly setose ventrally and / or laterally (vs femorites stout, only ~ 2 × as long as acropodites, clearly flattened dorsoventrally, contiguous all along or nearly so, and densely setose on both ventral and lateral sides).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |