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<document ID-CLB-Dataset="21016" ID-DOI="http://dx.doi.org/10.3897/zookeys.964.55703" ID-GBIF-Dataset="8af068af-c393-4348-816f-b46b796f7ab0" ID-PMC="PMC7471130" ID-Pensoft-Pub="1313-2970-964-143" ID-Pensoft-UUID="3357582565E7539FB17394EEA89FE63A" ID-PubMed="32939150" ID-ZooBank="CB14C5685B55487E94A3AE980C2E839B" ModsDocID="1313-2970-964-143" checkinTime="1598623661843" checkinUser="pensoft" docAuthor="Lu, Xiao-Qiang &amp; Du, Xi-Cui" docDate="2020" docId="F2D86EA8E65355FA98E73EE7A737CF04" docLanguage="en" docName="ZooKeys 964: 143-159" docOrigin="ZooKeys 964" docSource="http://dx.doi.org/10.3897/zookeys.964.55703" docTitle="Nagiella quadrimaculalis" docType="treatment" docVersion="6" id="3357582565E7539FB17394EEA89FE63A" lastPageNumber="143" masterDocId="3357582565E7539FB17394EEA89FE63A" masterDocTitle="Revision of Nagiella Munroe (Lepidoptera, Crambidae), with the description of a new species from China" masterLastPageNumber="159" masterPageNumber="143" pageNumber="143" updateTime="1732754367688" updateUser="ExternalLinkService">
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<mods:title id="86028D6F49B6B1240C4B60CC3D41B3DA">Revision of Nagiella Munroe (Lepidoptera, Crambidae), with the description of a new species from China</mods:title>
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<mods:namePart id="48CA6B7D12D767E06115C9A31A558819">Lu, Xiao-Qiang</mods:namePart>
<mods:affiliation id="F4FF289C581798A42B7241EB99D13D34">College of Plant Protection, Southwest University, Chongqing, China</mods:affiliation>
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<mods:namePart id="08971CDFB52760774B7B73BFEB7D387E">Du, Xi-Cui</mods:namePart>
<mods:affiliation id="51C56D0FA85D46B0B2779BDBBBA266F7">College of Plant Protection, Southwest University, Chongqing, China</mods:affiliation>
<mods:nameIdentifier id="E6CB1FB9B6FBBB489DE26909BA86B44A" type="orcid">https://orcid.org/0000-0002-7796-7303</mods:nameIdentifier>
<mods:nameIdentifier id="1832618059698FCC40611BCDB523E763" type="email">duxicui@hotmail.com</mods:nameIdentifier>
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<mods:date id="F9877E7731828C423A2F5429AD762D49">2020</mods:date>
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<treatment id="F2D86EA8E65355FA98E73EE7A737CF04" ID-GBIF-Taxon="167407667" LSID="urn:lsid:plazi:treatment:F2D86EA8E65355FA98E73EE7A737CF04" httpUri="http://treatment.plazi.org/id/F2D86EA8E65355FA98E73EE7A737CF04" lastPageNumber="143" pageId="0" pageNumber="143" scope_family="Crambidae" scope_order="Lepidoptera">
<subSubSection id="F5B09A8263D6866CAED412F6F1574C2F" pageId="0" pageNumber="143" type="nomenclature">
<paragraph id="349D46916A3B3C4D56982854D16E96F8" pageId="0" pageNumber="143">
<taxonomicName id="0D37B4AE113B9DAB93347A960F6CF3B8" ID-CoL="45KHY" LSID="F2D86EA8-E653-55FA-98E7-3EE7A737CF04" authority="(Kollar &amp; Redtenbacher, 1844)" baseAuthorityName="Kollar &amp; Redtenbacher" baseAuthorityYear="1844" class="Insecta" family="Crambidae" genus="Nagiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nagiella quadrimaculalis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="quadrimaculalis">Nagiella quadrimaculalis (Kollar &amp; Redtenbacher, 1844)</taxonomicName>
<figureCitation id="1D29912F4179677B559628604D629B1D" captionStart="Figures 4–8" captionStartId="F3" captionText="Figures 4 - 8. Habitus of Nagiella species 4 N. hortulatoides male 5 N. inferior male 6 N. quadrimaculalis male 7 Nagia incomitata Swinhoe 1894 female, type, BMNH Pyr., London. 7 A head 8 Nagiella bispina sp. nov. male, holotype." figureDoi="10.3897/zookeys.964.55703.figures4-8" httpUri="https://binary.pensoft.net/fig/446787" pageId="0" pageNumber="143">Figures 6</figureCitation>
<figureCitation id="ED6E616B12375C7599050AA64DE1C02C" captionStart="Figures 9–15" captionStartId="F4" captionText="Figures 9 - 15. Genitalia of Nagiella species 9 N. hortulatoides: male, genitalia slide no. LXQ 18311 10, 13 N. inferior: 10 male, genitalia slide no. LXQ 18291 13 female, genitalia slide no. XLJ 14239 11, 14 N. quadrimaculalis: 11 male, genitalia slide no. LXQ 18310 14 female, genitalia slide no. LXQ 18306 12, 15 N. bispina sp. nov.: 12 male, holotype, genitalia slide no. XLJ 14011 15 female, paratype, slide no. XLJ 14009 15 A signa." figureDoi="10.3897/zookeys.964.55703.figures9-15" httpUri="https://binary.pensoft.net/fig/446788" pageId="0" pageNumber="143">, 11</figureCitation>
<figureCitation id="BFDAC81BFA6ECE9A65F1149773FDF777" captionStart="Figures 9–15" captionStartId="F4" captionText="Figures 9 - 15. Genitalia of Nagiella species 9 N. hortulatoides: male, genitalia slide no. LXQ 18311 10, 13 N. inferior: 10 male, genitalia slide no. LXQ 18291 13 female, genitalia slide no. XLJ 14239 11, 14 N. quadrimaculalis: 11 male, genitalia slide no. LXQ 18310 14 female, genitalia slide no. LXQ 18306 12, 15 N. bispina sp. nov.: 12 male, holotype, genitalia slide no. XLJ 14011 15 female, paratype, slide no. XLJ 14009 15 A signa." figureDoi="10.3897/zookeys.964.55703.figures9-15" httpUri="https://binary.pensoft.net/fig/446788" pageId="0" pageNumber="143">, 14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="87423E965B348E40C4E035CB796D6AE5" pageId="0" pageNumber="143" type="reference_group">
<paragraph id="18BDB629303EFC09ACF113790485E255" pageId="0" pageNumber="143">
<taxonomicName id="7D3DAE807573223F46A0D768A6C4A27E" authorityName="Kollar &amp; Redtenbacher" authorityYear="1844" class="Insecta" family="Crambidae" genus="Scopula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scopula quadrimaculalis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="quadrimaculalis">Scopula quadrimaculalis</taxonomicName>
Kollar &amp; Redtenbacher, 1844: 492.
</paragraph>
<paragraph id="DD054B73BF6A7A6ADEDEFEC311816615" pageId="0" pageNumber="143">
<taxonomicName id="E7C7B0809A10BEFE52A997A7BDC78DB6" authorityName="Walker" authorityYear="1866" class="Insecta" family="Crambidae" genus="Nagia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nagia desmialis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="desmialis">Nagia desmialis</taxonomicName>
Walker, 1866: 1320.
</paragraph>
<paragraph id="4EB46A2D649C08E2A37D239C0A9255F4" pageId="0" pageNumber="143">
<taxonomicName id="256DB66D082C17E2511F0BF65C681712" class="Insecta" family="Crambidae" genus="Omiodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omiodes quadrimaculalis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="quadrimaculalis">Omiodes quadrimaculalis</taxonomicName>
: Meyrick, 1890: 441.
</paragraph>
<paragraph id="5599237B49253027F521271221EE7A8E" pageId="0" pageNumber="143">
<taxonomicName id="5F3D7439A4227366B6D31DEE3CF6764B" authorityName="Motschulsky" authorityYear="1861" class="Insecta" family="Crambidae" genus="Botys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Botys quadrimaculalis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="quadrimaculalis">Botys quadrimaculalis</taxonomicName>
: Snellen, 1890: 589.
</paragraph>
<paragraph id="5A5A582072A14D8D6833911F0575604B" pageId="0" pageNumber="143">
<taxonomicName id="A2F973A5976346B18B8B44AA6E7D7FA0" authorityName="Hubner" authorityYear="1825" class="Insecta" family="Crambidae" genus="Sylepta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sylepta" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">Sylepta</taxonomicName>
[sic]
<taxonomicName id="666184937977C272D3E86057F148106F" class="Insecta" family="Crambidae" genus="Sylepta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="quadrimaculalis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="quadrimaculalis">quadrimaculalis</taxonomicName>
: Hampson, 1896: 336.
</paragraph>
<paragraph id="C017FA170A430007D524AE4399BB6AE7" pageId="0" pageNumber="143">
<taxonomicName id="ABA19A0A3FEB0265FE7306404ADDCAA5" authorityName="Hubner" authorityYear="1825" class="Insecta" family="Crambidae" genus="Sylepta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sylepta" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">Sylepta</taxonomicName>
[sic]
<taxonomicName id="35DA2527429FBF27C4EE75D858072F8F" authorityName="Hampson" authorityYear="1912" class="Insecta" family="Crambidae" genus="Sylepta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="desmialis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="desmialis">desmialis</taxonomicName>
: Swinhoe, 1906: 293.
</paragraph>
<paragraph id="763169C49590E2B81DA41C8EB6F05650" pageId="0" pageNumber="143">
<taxonomicName id="1014C20DBE0045EE80CA14D79E52D04B" baseAuthorityName="Kollar &amp; Redtenbacher" baseAuthorityYear="1844" class="Insecta" family="Crambidae" genus="Nagiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nagiella quadrimaculalis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="quadrimaculalis">Nagiella quadrimaculalis</taxonomicName>
: Munroe, 1976: 876.
</paragraph>
<paragraph id="7BE06DE7443E77D14CE0813DF6B04380" pageId="0" pageNumber="143">
<taxonomicName id="8590AC07C568EC3014671799AE0EDC3A" baseAuthorityName="Kollar &amp; Redtenbacher" baseAuthorityYear="1844" class="Insecta" family="Crambidae" genus="Pleuroptya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pleuroptya quadrimaculalis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="quadrimaculalis">Pleuroptya quadrimaculalis</taxonomicName>
: Inoue, 1982: 343.
</paragraph>
</subSubSection>
<subSubSection id="59BBA79E83A8452386C85BE88E879ECD" pageId="0" pageNumber="143" type="materials_examined">
<paragraph id="212C4D978B82F09517DC30A85B0A5D39" pageId="0" pageNumber="143">Material examined.</paragraph>
<paragraph id="23ECF0FA754FAD15CBDD52396F93E3E8" pageId="0" pageNumber="143">
China, Gansu: 4 ♂♂, 3 ♀♀, Kangxian County, Baiyun Mountain, 1200 m, 3.VII.2018, leg. Xiao-Qiang Lu &amp; Xi-Cui Du; Shanxi: 4 ♂♂, Lishanxiachuan Nature Reserve, 1560 m, 26.VII.2012, leg. Gui-Qing He; Shaanxi: 11 ♂♂, Ningshan County, Yangjuba Town, 1400 m, 4.VIII.2014, leg. Hai-Li Yu &amp; Jiu-Yang Luo; 7 ♂♂, 8 ♀♀, Yang County, 3500 m, 15.VIII.2017, leg. Jian-Yue Qiu &amp; Hao Xu; 7 ♂♂, 1 ♀, Taibai County, Huangbaiyuan, 1291 m, 16.VII.2018, leg. Qing-Ming Liu; Henan: 33 ♂♂, Neixiangbaotianman Nature Reserve, Luotiofeng, 1300 m, 8.VI.2017, leg. Jian-Yue Qiu &amp; Hao Xu; Hubei: 28 ♂♂, 3 ♀, Dabie Mountain, Taohua Village, 590 m, 20.VII.2010, leg. Li-Jun Xu; 4 ♂♂, Luotian County, Qingguantai, 580 m, 1.VII.2014, leg. Jiu-Yang Luo; 11 ♂♂, Changyang County, Hejiaping, 800 m, 18.VI.2018, leg. Xiao-Qiang Lu &amp; Xi-Cui Du; 4 ♂♂, Wufenghou River, 1100 m, 26, VII.2018, leg. Jian-Yue Qiu &amp; Hao Xu; Hunan: 7 ♂♂, Shimen County, Huping Mountain, Dadongping, 1400 m, 8.VI.2017, leg. Jian-Yue Qiu &amp; Hao Xu; 5 ♂♂, Sangzhi County, tianping Mountain, 1400 m, 15.VII.2018, leg. Jian-Yue Qiu &amp; Hao Xu; 6 ♂♂, 2 ♀♀, Yizhang County, Mangshan Nature Reserve, 1000 m, 2.VIII.2018, leg. Jian-Yue Qiu &amp; Hao Xu; Zhejiang: 10 ♂♂, Tianmu Mountain Nature Reserve, 400 m, 26-29.VII.2011, leg. Xiao-Bing Fu &amp; Xi-Cui Du; 11 ♂♂, 3 ♀♀, Qingliang Mountain, Shunxiwu, 300 m, 18-21.V.2012, leg. Xiao-Bing Fu; Jiangxi: 1 ♂, 1 ♀, Jinggangshan City, Xiaoxidong, 625 m, 30.V.2011, leg. Jin-Wei Li; Chongqing: 18 ♂♂, 10 ♀♀, Jingfo Mountain Nature Reserve, 1700 m, 12.VII.2010, leg. Xi-Cui Du &amp; Sheng-wen Shi; 10 ♂♂, Simian Mountain Nature Reserve, 1120 m, 19.VII. 2010, leg. Xi-Cui Du &amp; Li-fang Song; 3 ♂♂, Simian Mountain Nature Reserve, 1200 m, 15-19. VII. 2012, leg. Gui-Qing He &amp; Li-Jun Xu; 13 ♂♂, 1 ♀, Chengkou County, Xingtian Village, 1300 m, 1.VII.2013, leg. Gui-Qing He &amp; Li-Jun Xu; Sichuan: 4 ♂♂, 1 ♀, Luding County, Hailuogou, 3478 m, 4.VII.2012, leg. Jin-Wei Li; 19 ♂♂, Nanjiang County, Guangwu Mountain, 700 m, 3.VII.2013, leg. Gui-Qing He &amp; Li-Jun Xu; 21 ♂♂, 14 ♀♀, Xuyong County, huagaoxi Nature Reserve, 621 m, 26-30.VIII.2013, leg. Dan Xu &amp; Xue-Li Wei; 19 ♂♂, 3 ♀♀,
<normalizedToken id="4F444786C20DAF0026D5DF1295C36CBD" originalValue="An’zi">An'zi</normalizedToken>
River Nature Reserve, 1690 m, 4.VIII.2015, leg. Xi-Cui Du; Guizhou: 11 ♂♂, 1 ♀, Kuankuoshui Nature Reserve, 800 m, 10-17.VIII.2010, leg. Xi-Cui Du; 5 ♂♂, Libo County,
<normalizedToken id="C79B707EB5A851CA44660DA9C8832B77" originalValue="An’xiang">An'xiang</normalizedToken>
, 1345 m, 22.VII.2015, leg. Ji-Ping Wan; Yunnan: 1 ♂, 1 ♀, Ninglang County, Xichuan, 2400 m, 20.VII.2013, leg. Gui-Qing He; 19 ♂♂, 2 ♂♂, Malipo County, Daxichang, 1465 m, 7.VI.2015, leg. Man-Fei Tao; Dawei Mountain Nature Reserve, 2700 m, 27.V.2018, leg. Xiao-Qiang Lu &amp; Xi-Cui Du; 10 ♂♂, Huanglian Mountain Nature Reserve, 900 m, 23.VI.2018, leg. Xiao-Qiang Lu &amp; Xi-Cui Du; 15 ♂♂, 1 ♀, Xihuangbanna Prefecture, Menglun Town, 620 m, 17.V.2018, leg. Xiao-Qiang Lu &amp; Xi-Cui Du; Guangdong: 27 ♂♂, 15 ♀♀, Nanlingbabao Mountain Nature Reserve, 1070 m, 22.VIII.2010, leg. Xi-Cui Du; 2 ♂♂, 1 ♀, Shixing County, Baling Nature Reserve, 496 m, 29.V.2017, leg. Yong-Hong Duan (SYSU); Guangxi: 22 ♂♂, Hechi City, Jiuwan Mountain, 1600 m, 26.VII.2015, leg. Ji-Ping Wan; 4 ♂♂, Guilin City, Maoer Mountain Nature Reserve, 1100 m, 23.VII.2015, leg. Kai-Li Liu &amp; Jing-Xia Zhao; 9 ♂♂, Rongshui County, Peixiu Village, 1900 m, 24.VIII.2015, leg. Ji-Ping Wan; 12 ♂♂, Cenwanglao Mountain, Dalongping, 1290 m, 4.VIII.2014, leg. Xue-Li Wei &amp; Chao Ran; Fujian: 8 ♂♂, Wuyi Mountain Nature Reserve, Tongmu Village, 758 m, 20.VIII.2016, leg. Kai Chen &amp; Yong-Hong Duan (SYSU). Genitalia slide no.: XLJ13123 ♂, XLJ13158 ♂, XLJ13215 ♂, XLJ14029 ♂, XLJ14056 ♂, XLJ14075 ♂, XLJ14076 ♂, XLJ14133 ♂, XLJ14229 ♂, LXQ19304 ♂, LXQ19305 ♂, LXQ18308 ♂, LXQ18310 ♂, XLJ13124 ♀, XLJ13159 ♀, XLJ13216 ♀, XLJ114012 ♀, XLJ14030 ♀, LXQ18306 ♀.
</paragraph>
</subSubSection>
<subSubSection id="2FBC19900BDDD8F3E217462B22DAB313" pageId="0" pageNumber="143" type="diagnosis">
<paragraph id="EA3210E02ACF5D097A47E4328EC0F055" pageId="0" pageNumber="143">Diagnosis.</paragraph>
<paragraph id="A751BD0AFDFF39C9E981BFDD22A351D8" pageId="0" pageNumber="143">
Adult (Fig.
<figureCitation id="A2AFADF415527686C5A3094E82770565" captionStart="Figures 4–8" captionStartId="F3" captionText="Figures 4 - 8. Habitus of Nagiella species 4 N. hortulatoides male 5 N. inferior male 6 N. quadrimaculalis male 7 Nagia incomitata Swinhoe 1894 female, type, BMNH Pyr., London. 7 A head 8 Nagiella bispina sp. nov. male, holotype." figureDoi="10.3897/zookeys.964.55703.figures4-8" httpUri="https://binary.pensoft.net/fig/446787" pageId="0" pageNumber="143">6</figureCitation>
): Wings brown. Forewing length 12.0-20.0mm (wingspan 26.0-43.0 mm); a small white spot between the orbicular spot and discoidal spot; a large white sub-reniform spot between the discoidal spot and postmedial line, up to Rs2+Rs3 and down to CuA1; antemedial and postmedial line unclear. Hindwing with a large white irregular quadrilateral spot between the discoidal spot and postmedial line, dentated between M2 and M3. Male genitalia (Fig.
<figureCitation id="CBE96B82E08D741DE2660336AD343D20" captionStart="Figures 9–15" captionStartId="F4" captionText="Figures 9 - 15. Genitalia of Nagiella species 9 N. hortulatoides: male, genitalia slide no. LXQ 18311 10, 13 N. inferior: 10 male, genitalia slide no. LXQ 18291 13 female, genitalia slide no. XLJ 14239 11, 14 N. quadrimaculalis: 11 male, genitalia slide no. LXQ 18310 14 female, genitalia slide no. LXQ 18306 12, 15 N. bispina sp. nov.: 12 male, holotype, genitalia slide no. XLJ 14011 15 female, paratype, slide no. XLJ 14009 15 A signa." figureDoi="10.3897/zookeys.964.55703.figures9-15" httpUri="https://binary.pensoft.net/fig/446788" pageId="0" pageNumber="143">11</figureCitation>
): Uncus trapezoidal. Gnathos slender, finger-like. Clasper thickly finger-like. Female genitalia (Fig.
<figureCitation id="ED5F8C8F470BDF6C34FFF98549B2BB02" captionStart="Figures 9–15" captionStartId="F4" captionText="Figures 9 - 15. Genitalia of Nagiella species 9 N. hortulatoides: male, genitalia slide no. LXQ 18311 10, 13 N. inferior: 10 male, genitalia slide no. LXQ 18291 13 female, genitalia slide no. XLJ 14239 11, 14 N. quadrimaculalis: 11 male, genitalia slide no. LXQ 18310 14 female, genitalia slide no. LXQ 18306 12, 15 N. bispina sp. nov.: 12 male, holotype, genitalia slide no. XLJ 14011 15 female, paratype, slide no. XLJ 14009 15 A signa." figureDoi="10.3897/zookeys.964.55703.figures9-15" httpUri="https://binary.pensoft.net/fig/446788" pageId="0" pageNumber="143">14</figureCitation>
): Signum small, round.
</paragraph>
</subSubSection>
<subSubSection id="128B7FE4F75BE33855DED98D24FE0B4F" pageId="0" pageNumber="143" type="male genitalia">
<paragraph id="B95C3967BB17C4802208299261BC1D9F" pageId="0" pageNumber="143">Male genitalia</paragraph>
<paragraph id="8878273B9393C749D00D23029433933B" pageId="0" pageNumber="143">
(Fig.
<figureCitation id="FA42EB8AA427156DA6A21FC80AF868DA" captionStart="Figures 9–15" captionStartId="F4" captionText="Figures 9 - 15. Genitalia of Nagiella species 9 N. hortulatoides: male, genitalia slide no. LXQ 18311 10, 13 N. inferior: 10 male, genitalia slide no. LXQ 18291 13 female, genitalia slide no. XLJ 14239 11, 14 N. quadrimaculalis: 11 male, genitalia slide no. LXQ 18310 14 female, genitalia slide no. LXQ 18306 12, 15 N. bispina sp. nov.: 12 male, holotype, genitalia slide no. XLJ 14011 15 female, paratype, slide no. XLJ 14009 15 A signa." figureDoi="10.3897/zookeys.964.55703.figures9-15" httpUri="https://binary.pensoft.net/fig/446788" pageId="0" pageNumber="143">11</figureCitation>
). Uncus trapezoidal. Gnathos slender, finger-like. Valva elongate lingulate, with apex narrowed, posterior margin with a cluster of long setae near the middle; clasper thickly finger-like. Saccus conical, broad. Juxta peach-shaped. Phallus longitudinally wrinkled distally.
</paragraph>
</subSubSection>
<subSubSection id="AB045937F72863FD65506082D072990C" pageId="0" pageNumber="143" type="female genitalia">
<paragraph id="BEB21AADFAF23F3359DC6E8185A2F3D5" pageId="0" pageNumber="143">Female genitalia</paragraph>
<paragraph id="21093B862C7E380F0F4001E012E7AF4D" pageId="0" pageNumber="143">
(Fig.
<figureCitation id="390686CEC567AA137D56611C4FA12443" captionStart="Figures 9–15" captionStartId="F4" captionText="Figures 9 - 15. Genitalia of Nagiella species 9 N. hortulatoides: male, genitalia slide no. LXQ 18311 10, 13 N. inferior: 10 male, genitalia slide no. LXQ 18291 13 female, genitalia slide no. XLJ 14239 11, 14 N. quadrimaculalis: 11 male, genitalia slide no. LXQ 18310 14 female, genitalia slide no. LXQ 18306 12, 15 N. bispina sp. nov.: 12 male, holotype, genitalia slide no. XLJ 14011 15 female, paratype, slide no. XLJ 14009 15 A signa." figureDoi="10.3897/zookeys.964.55703.figures9-15" httpUri="https://binary.pensoft.net/fig/446788" pageId="0" pageNumber="143">14</figureCitation>
). Apophyses anteriores ca. twice the length of apophyses posteriores. Ductus bursae ca. twice the length of corpus bursae, distinctly narrowed near the base; ductus seminalis from approximately one third of the ductus bursae. Corpus bursae oval, with a small round signum.
</paragraph>
</subSubSection>
<subSubSection id="070B4D016B6772C6A2A9A4F62E00C135" pageId="0" pageNumber="143" type="distribution">
<paragraph id="8B1AAE29CFA3BFE67BF7288EAC537D28" pageId="0" pageNumber="143">Distribution.</paragraph>
<paragraph id="BEFBF5A8E4540C6F5F57B2EEC99EFB20" pageId="0" pageNumber="143">
China (Heilongjiang, Liaoning, Gansu, Shanxi, Shaanxi, Henan, Hebei, Hubei, Shandong, Hunan, Zhejiang, Jiangxi, Tibet, Sichuan, Chongqing, Guizhou, Yunnan, Guangdong, Guangxi, Hainan, Fujian, Taiwan), Korea, Japan, Indonesia, India (Sikkim), Nepal, Russia (far east), Malaysia (Walker 1866;
<bibRefCitation id="6645EE8ACFA77FEFDB1B9490FF611E4F" author="Inoue, H" editor="Inoue, H" journalOrPublisher="Kodansha, Tokyo," pageId="0" pageNumber="143" pagination="307 - 404" refId="B14" refString="Inoue, H, 1982. Pyralidae. In: Inoue, H, Ed., (Eds) Moths of Japan. Kodansha, Tokyo, 1: 307 - 404" title="Pyralidae." volume="1" volumeTitle="(Eds) Moths of Japan." year="1982">Inoue 1982</bibRefCitation>
;
<bibRefCitation id="749658800AD726C15AD3088A4EC5063B" DOI="https://doi.org/10.1080/00305316.2008.10417555" author="Du, XC" editor="Li, HH" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="143" pagination="237 - 305" refId="B4" refString="Du, XC, 2009. Spilomelinae. In: Li, HH, Ren, YD, Zhang, DD, Du, XC, Li, WC, You, P, Eds., Insect fauna of Henan (Lepidoptera: Pyraloidea). Science Press, Beijing: 237 - 305, DOI: https://doi.org/10.1080/00305316.2008.10417555" title="Spilomelinae." url="https://doi.org/10.1080/00305316.2008.10417555" volumeTitle="Insect fauna of Henan (Lepidoptera: Pyraloidea)." year="2009">Du 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D06DCEE12CE1E520FF17A658C399496E" pageId="0" pageNumber="143" type="host">
<paragraph id="C7768434B8609A4046F696906BDDCC6B" pageId="0" pageNumber="143">Host.</paragraph>
<paragraph id="9F462E3EE462A3CD642989E626C5C208" pageId="0" pageNumber="143">
<taxonomicName id="7AB1551542F103D82CC6C2EA938E1CF7" class="Magnoliopsida" family="Anacardiaceae" genus="Rhus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhus chinensis" order="Sapindales" pageId="0" pageNumber="143" phylum="Tracheophyta" rank="species" species="chinensis">
<emphasis id="4BBF5F94C045D587557127425A68395F" italics="true" pageId="0" pageNumber="143">Rhus chinensis</emphasis>
</taxonomicName>
Mill (
<taxonomicName id="AA4BCF11A6E22E038683057FBE8F1A3B" class="Magnoliopsida" family="Anacardiaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Sapindales" pageId="0" pageNumber="143" phylum="Tracheophyta" rank="family">Anacardiaceae</taxonomicName>
) (
<bibRefCitation id="4A950386279A102EF38B63595BAAF51E" author="Fan, LH" journalOrPublisher="Proft et Storch, Hafniae" pageId="0" pageNumber="143" refId="B6" refString="Fan, LH, Piao, MH, 2013. Larval descriptions for four species in the genus Pleuroptya (Lepidoptera: Crambidae: Spilomelinae) from China, with notes on biology, Entomotaxonomia 35(1): 45-52." title="Larval descriptions for four species in the genus Pleuroptya (Lepidoptera: Crambidae: Spilomelinae) from China, with notes on biology, Entomotaxonomia 35 (1): 45 - 52." year="2013">Fan and Piao 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D5A377B9DBDA56C0D9671DE3AB078332" pageId="0" pageNumber="143" type="remarks">
<paragraph id="182B741E9792105F3905C2FAFF508351" pageId="0" pageNumber="143">Remarks.</paragraph>
<paragraph id="54ABAFBFF5C843AE7C325AFA08B2C73A" pageId="0" pageNumber="143">
In addition to
<taxonomicName id="1D4A8BF35264F078F89846EF72F75C21" class="Magnoliopsida" family="Anacardiaceae" genus="Rhus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhus chinensis" order="Sapindales" pageId="0" pageNumber="143" phylum="Tracheophyta" rank="species" species="chinensis">
<emphasis id="3679B7B12A2473DA95629FEFDBC225E9" italics="true" pageId="0" pageNumber="143">Rhus chinensis</emphasis>
</taxonomicName>
Mill, another host,
<taxonomicName id="1CCF2845C8269DC834D3E118840F374B" authorityName="Makino" authorityYear="1903" baseAuthorityName="Thunb." class="Magnoliopsida" family="Apocynaceae" genus="Metaplexis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Metaplexis japonica" order="Gentianales" pageId="0" pageNumber="143" phylum="Tracheophyta" rank="species" species="japonica">
<emphasis id="AA6E7FC2744351E6EFC9C5EAAF57A207" italics="true" pageId="0" pageNumber="143">Metaplexis japonica</emphasis>
</taxonomicName>
Makino (
<taxonomicName id="63981AA585F32D858ECBF8315E1B2662" class="Magnoliopsida" family="Apocynaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Gentianales" pageId="0" pageNumber="143" phylum="Tracheophyta" rank="family">Apocynaceae</taxonomicName>
), was recorded by
<bibRefCitation id="405CC353B044F9221B3348A71BFD1FAC" author="Fan, LH" journalOrPublisher="Proft et Storch, Hafniae" pageId="0" pageNumber="143" refId="B6" refString="Fan, LH, Piao, MH, 2013. Larval descriptions for four species in the genus Pleuroptya (Lepidoptera: Crambidae: Spilomelinae) from China, with notes on biology, Entomotaxonomia 35(1): 45-52." title="Larval descriptions for four species in the genus Pleuroptya (Lepidoptera: Crambidae: Spilomelinae) from China, with notes on biology, Entomotaxonomia 35 (1): 45 - 52." year="2013">Fan and Piao (2013)</bibRefCitation>
in the same article according to
<bibRefCitation id="E7DCBF832152B78FDDCE513BD8F777C0" author="Yoshiyasu, Y" journalOrPublisher="Japanese Journal of Entomology" pageId="0" pageNumber="143" pagination="1 - 774" refId="B48" refString="Yoshiyasu, Y, 1991. New host record and feeding habit Glyphodes quadrimaculalis (Lepidoptera, Pyralidae). Japanese Journal of Entomology 59 (4): 1 - 774" title="New host record and feeding habit Glyphodes quadrimaculalis (Lepidoptera, Pyralidae)." volume="59" year="1991">Yoshiyasu (1991)</bibRefCitation>
. However, we found
<taxonomicName id="B8ED885C7C54A9B2F24058D0DB80A7C1" lsidName="M. japonica" pageId="0" pageNumber="143" rank="species" species="japonica">
<emphasis id="0BF152B4B965F73CFB87D6CD8BA09508" italics="true" pageId="0" pageNumber="143">M. japonica</emphasis>
</taxonomicName>
was recorded by
<bibRefCitation id="A744338E0CCCA749B756F8634C478C3E" author="Yoshiyasu, Y" journalOrPublisher="Japanese Journal of Entomology" pageId="0" pageNumber="143" pagination="1 - 774" refId="B48" refString="Yoshiyasu, Y, 1991. New host record and feeding habit Glyphodes quadrimaculalis (Lepidoptera, Pyralidae). Japanese Journal of Entomology 59 (4): 1 - 774" title="New host record and feeding habit Glyphodes quadrimaculalis (Lepidoptera, Pyralidae)." volume="59" year="1991">Yoshiyasu (1991)</bibRefCitation>
as the host of
<taxonomicName id="DB2EABB8E8E7FC0492C88BF40BF85C33" authorityName="Bremer &amp; Grey" authorityYear="1853" class="Insecta" family="Crambidae" genus="Glyphodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphodes quadrimaculalis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="quadrimaculalis">
<emphasis id="7D982B200C789537C58B1135497871CE" italics="true" pageId="0" pageNumber="143">Glyphodes quadrimaculalis</emphasis>
</taxonomicName>
(Bremer and Grey 1853) but not of
<taxonomicName id="5939A279AA7F93B493DF1D054AD47D2C" lsidName="N. quadrimaculalis" pageId="0" pageNumber="143" rank="species" species="quadrimaculalis">
<emphasis id="A9CB572A946A467EADD5B2D9533BF12C" italics="true" pageId="0" pageNumber="143">N. quadrimaculalis</emphasis>
</taxonomicName>
(Kollar and Redtenbacher).
<taxonomicName id="5791174E74FF1BEFE6353CE4EE965835" class="Magnoliopsida" family="Anacardiaceae" genus="Rhus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rhus chinensis" order="Sapindales" pageId="0" pageNumber="143" phylum="Tracheophyta" rank="species" species="chinensis">
<emphasis id="B58CEF546E911F254795943BB0B3DF8F" italics="true" pageId="0" pageNumber="143">Rhus chinensis</emphasis>
</taxonomicName>
Mill is the only host of
<taxonomicName id="09CAFA92E676F3B3437C6927F6DEFA5E" lsidName="N. quadrimaculalis" pageId="0" pageNumber="143" rank="species" species="quadrimaculalis">
<emphasis id="A326FDC9C7A960A2604AECC93B991C88" italics="true" pageId="0" pageNumber="143">N. quadrimaculalis</emphasis>
</taxonomicName>
(Kollar and Redtenbacher) known so far.
</paragraph>
<paragraph id="6E685F25D4DB5DF3D7E6D79F34A30FCC" pageId="0" pageNumber="143">
<bibRefCitation id="4B36001C4F83FB78011666869ABE6575" DOI="https://doi.org/10.1080/00222939408677791" author="Swinhoe, C" journalOrPublisher="Systematic Biology" pageId="0" pageNumber="143" refId="B38" refString="Swinhoe, C, 1894. New Pyrales from the Khasia Hills. Annals and Magazine of Natural History, including Zoology, Botany and Geology, London (ser. 6) 14(81): 197-210. https://doi.org/10.1080/00222939408677791" title="New Pyrales from the Khasia Hills. Annals and Magazine of Natural History, including Zoology, Botany and Geology, London (ser. 6) 14 (81): 197 - 210." url="https://doi.org/10.1080/00222939408677791" year="1894">Swinhoe (1894)</bibRefCitation>
stated that
<taxonomicName id="4A90672AEF3F2E77EEFCF81CB24CFA57" authorityName="C.Swinhoe" authorityYear="1894" class="Insecta" family="Erebidae" genus="Nagia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nagia incomitata" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="incomitata">
<emphasis id="93BD0C8CF422FEB84CCB7461F1E84843" italics="true" pageId="0" pageNumber="143">Nagia incomitata</emphasis>
</taxonomicName>
was between
<taxonomicName id="90F8259ECF3C5EBB2407AF2E8AB16E35" class="Insecta" family="Erebidae" genus="Nagia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nagia quadrimaculalis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="quadrimaculalis">
<emphasis id="1A3EA4FBDA5FC8CE0F6A10750423B50B" italics="true" pageId="0" pageNumber="143">Nagia quadrimaculalis</emphasis>
</taxonomicName>
and
<taxonomicName id="1ED5A77B5E174C4280A636CE4D330D32" lsidName="N. flavispila" pageId="0" pageNumber="143" rank="species" species="flavispila">
<emphasis id="A5EAF245E76082F1DA19295D117C0B13" italics="true" pageId="0" pageNumber="143">N. flavispila</emphasis>
</taxonomicName>
, but quite different to either. But
<taxonomicName id="52DD6D3E13B67C0950AD47A6FEC7AC88" lsidName="N. incomitata" pageId="0" pageNumber="143" rank="species" species="incomitata">
<emphasis id="B0FA832181FDD63A041F8DAF05797147" italics="true" pageId="0" pageNumber="143">N. incomitata</emphasis>
</taxonomicName>
was regarded as a synonym of
<taxonomicName id="6FB02E546164E181B159F7471EB4B5FE" lsidName="N. quadrimaculalis" pageId="0" pageNumber="143" rank="species" species="quadrimaculalis">
<emphasis id="D33426833B25DBA7B2CE45D21E615DE2" italics="true" pageId="0" pageNumber="143">N. quadrimaculalis</emphasis>
</taxonomicName>
because they were similar in habitus (
<bibRefCitation id="B2AEBA3E2176DD3ED4627234A7E9003B" author="Bae, YS" journalOrPublisher="Korea National Arboretum, Samsungad, Seoul" pageId="0" pageNumber="143" refId="B1" refString="Bae, YS, Byun, BK, Paek, MK, 2008. Pyralid moths of Korea (Lepidoptera: Pyraloidea). Korea National Arboretum, Samsungad, Seoul" title="Pyralid moths of Korea (Lepidoptera: Pyraloidea)." year="2008">Bae et al. 2008</bibRefCitation>
). We investigated the original description and type specimen of
<taxonomicName id="888C9B13D64BB67BF836490CD78CDFB8" lsidName="N. incomitata" pageId="0" pageNumber="143" rank="species" species="incomitata">
<emphasis id="22D220F325A7DAFA467163A2458740F0" italics="true" pageId="0" pageNumber="143">N. incomitata</emphasis>
</taxonomicName>
Swinhoe, 1894, and compared them with the description and photographs of
<taxonomicName id="CFD9AEA7ABD8BD6C1346612043AB2968" lsidName="N. quadrimaculalis" pageId="0" pageNumber="143" rank="species" species="quadrimaculalis">
<emphasis id="B4B636F8710FAB0D2868A38225047859" italics="true" pageId="0" pageNumber="143">N. quadrimaculalis</emphasis>
</taxonomicName>
(
<bibRefCitation id="61B7CFBE1C0AEF058B827C046918CDE0" author="Kollar, V" journalOrPublisher="Junk, s' Gravenhage" pageId="0" pageNumber="143" refId="B21" refString="Kollar, V, Redtenbacher, L, 1844. Aufzaehlung und Beschreibung der von Freiherrn Carl v. Huegel auf seiner Reise durch Kaschmir und das Himaleyagebirge gesammelten Insecten. Hallbergersche Verlagshandlung, Stuttgart, 395-564." title="Aufzaehlung und Beschreibung der von Freiherrn Carl v. Huegel auf seiner Reise durch Kaschmir und das Himaleyagebirge gesammelten Insecten. Hallbergersche Verlagshandlung, Stuttgart, 395 - 564." year="1844">Kollar and Redtenbacher 1844</bibRefCitation>
;
<bibRefCitation id="2AEAD7760FFF38868451616E58B820D6" DOI="https://doi.org/10.1080/00305316.2008.10417555" author="Du, XC" editor="Li, HH" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="143" pagination="237 - 305" refId="B4" refString="Du, XC, 2009. Spilomelinae. In: Li, HH, Ren, YD, Zhang, DD, Du, XC, Li, WC, You, P, Eds., Insect fauna of Henan (Lepidoptera: Pyraloidea). Science Press, Beijing: 237 - 305, DOI: https://doi.org/10.1080/00305316.2008.10417555" title="Spilomelinae." url="https://doi.org/10.1080/00305316.2008.10417555" volumeTitle="Insect fauna of Henan (Lepidoptera: Pyraloidea)." year="2009">Du 2009</bibRefCitation>
;
<bibRefCitation id="DFEA82E0EC7B6DB340D91131D1EA3DE3" author="Sasaki, A" editor="Nasu, Y" journalOrPublisher="Gakken Education Publishing, Tokyo" pageId="0" pageNumber="143" pagination="74 - 84" refId="B33" refString="Sasaki, A, Yamanaka, H, 2013. Spilomelini. In: Nasu, Y, Hirowatari, T, Kishida, Y, Eds., The Standard of Moths in Japan IV. Gakken Education Publishing, Tokyo: 74 - 84" title="Spilomelini." volumeTitle="The Standard of Moths in Japan IV." year="2013">Sasaki and Yamanaka 2013</bibRefCitation>
). The third segment of labial palpus of the former is slender and pointed distally (Fig.
<figureCitation id="D013DF44099B93874F0FA48513A6755B" captionStart="Figures 4–8" captionStartId="F3" captionText="Figures 4 - 8. Habitus of Nagiella species 4 N. hortulatoides male 5 N. inferior male 6 N. quadrimaculalis male 7 Nagia incomitata Swinhoe 1894 female, type, BMNH Pyr., London. 7 A head 8 Nagiella bispina sp. nov. male, holotype." figureDoi="10.3897/zookeys.964.55703.figures4-8" httpUri="https://binary.pensoft.net/fig/446787" pageId="0" pageNumber="143">7A</figureCitation>
), the forewing has no small white spot between the orbicular spot and discoidal spot, and the large white spot beyond the cell is down to the CuA2 (Fig.
<figureCitation id="5C225EF8ADB3924B9FF25B17F9CC79DF" captionStart="Figures 4–8" captionStartId="F3" captionText="Figures 4 - 8. Habitus of Nagiella species 4 N. hortulatoides male 5 N. inferior male 6 N. quadrimaculalis male 7 Nagia incomitata Swinhoe 1894 female, type, BMNH Pyr., London. 7 A head 8 Nagiella bispina sp. nov. male, holotype." figureDoi="10.3897/zookeys.964.55703.figures4-8" httpUri="https://binary.pensoft.net/fig/446787" pageId="0" pageNumber="143">7</figureCitation>
); while the third segment of labial palpus of the latter is stubby and blunt distally (Fig.
<figureCitation id="76D47481D9C58454CC6979BF451FFF00" captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Head and wing venation of Nagiella quadrimaculalis (Kollar &amp; Redtenbacher, 1844). Wing slide no. LXQ 20001, male." figureDoi="10.3897/zookeys.964.55703.figures2-3" httpUri="https://binary.pensoft.net/fig/446786" pageId="0" pageNumber="143">2</figureCitation>
), the forewing has a small white spot between the orbicular spot and discoidal spot, and the large white spot beyond the cell is down to the CuA1 (Fig.
<figureCitation id="89C5055D60A94001A8F28C5D1381FEAD" captionStart="Figures 4–8" captionStartId="F3" captionText="Figures 4 - 8. Habitus of Nagiella species 4 N. hortulatoides male 5 N. inferior male 6 N. quadrimaculalis male 7 Nagia incomitata Swinhoe 1894 female, type, BMNH Pyr., London. 7 A head 8 Nagiella bispina sp. nov. male, holotype." figureDoi="10.3897/zookeys.964.55703.figures4-8" httpUri="https://binary.pensoft.net/fig/446787" pageId="0" pageNumber="143">6</figureCitation>
). Therefore,
<taxonomicName id="F733888B13A7BECD2AFE16ED1E37C86C" lsidName="N. incomitata" pageId="0" pageNumber="143" rank="species" species="incomitata">
<emphasis id="222EDEDA2D25FFD301B940EB1FFE8CE0" italics="true" pageId="0" pageNumber="143">N. incomitata</emphasis>
</taxonomicName>
is not a synonym of
<taxonomicName id="7B18316AAAD84A6B591FF27D006CB369" lsidName="N. quadrimaculalis" pageId="0" pageNumber="143" rank="species" species="quadrimaculalis">
<emphasis id="A72C7FD8F6FD90E9B70FDC382D7AE413" italics="true" pageId="0" pageNumber="143">N. quadrimaculalis</emphasis>
</taxonomicName>
.
<taxonomicName id="21F2056E195A215F492766F812734987" authorityName="C.Swinhoe" authorityYear="1894" class="Insecta" family="Erebidae" genus="Nagia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nagia incomitata" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="incomitata">
<emphasis id="12EC83DCFAC4E78D1F5479D661128683" italics="true" pageId="0" pageNumber="143">Nagia incomitata</emphasis>
</taxonomicName>
was transferred to
<taxonomicName id="BDDDF230793A2DA8D0613247D4356254" authorityName="Butler" authorityYear="1887" class="Insecta" family="Crambidae" genus="Chalcidoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalcidoptera" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B8EBAD93EEC749B44BE054CD24080960" italics="true" pageId="0" pageNumber="143">Chalcidoptera</emphasis>
</taxonomicName>
Butler, 1887 by
<bibRefCitation id="DE37A064A566AF45882C6BD0E26ADEC1" DOI="https://doi.org/10.1080/03745480109443321" author="Swinhoe, C" journalOrPublisher="Clarendon Press, Oxford" pageId="0" pageNumber="143" refId="B40" refString="Swinhoe, C, 1901. New genera and species of Eastern and Australian moths. Annals and Magazine of Natural History, including Zoology, Botany and Geology, London (ser. 7) 8: 16-27. https://doi.org/10.1080/03745480109443321" title="New genera and species of Eastern and Australian moths. Annals and Magazine of Natural History, including Zoology, Botany and Geology, London (ser. 7) 8: 16 - 27." url="https://doi.org/10.1080/03745480109443321" year="1901">Swinhoe (1901)</bibRefCitation>
after stating previously that it did not belong into
<taxonomicName id="87E816E5544D3B73F7AF2A807A721F42" class="Insecta" family="Erebidae" genus="Nagia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nagia" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="C507375987904BB2720BE600E6A25D46" italics="true" pageId="0" pageNumber="143">Nagia</emphasis>
</taxonomicName>
(
<bibRefCitation id="037974068DDC852A7D99C00571134D80" author="Swinhoe, C" journalOrPublisher="Clarendon Press, Oxford" pageId="0" pageNumber="143" refId="B39" refString="Swinhoe, C, 1900. Noctuina, Geometrina and Pyralidina. In: Swinhoe C, Walsingham L, Durrant JH (Eds) Catalogue of eastern and Australian LepidopteraHeterocera in the Collection of the Oxford University Museum. Part II. Clarendon Press, Oxford" title="Noctuina, Geometrina and Pyralidina. In: Swinhoe C, Walsingham L, Durrant JH (Eds) Catalogue of eastern and Australian LepidopteraHeterocera in the Collection of the Oxford University Museum. Part II." year="1900">Swinhoe 1900</bibRefCitation>
).
<bibRefCitation id="FD3F9ECF8477565EC7028F6323D74D04" author="Hampson, GF" journalOrPublisher="The Fauna of British India, including Ceylon and Burma, London 4" pageId="0" pageNumber="143" refId="B9" refString="Hampson, GF, 1896. Moths. The Fauna of British India, including Ceylon and Burma, London 4" title="Moths." year="1896">Hampson (1896)</bibRefCitation>
, on the other hand, considered it a synonym of
<taxonomicName id="D791706BA56CBB04FC7158ABC2E94FAB" authorityName="Walker" authorityYear="1859" class="Insecta" family="Crambidae" genus="Nosophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nosophora chironalis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="chironalis">
<emphasis id="9821A934F1045A88B1B45665F6AFB9B7" italics="true" pageId="0" pageNumber="143">Nosophora chironalis</emphasis>
</taxonomicName>
(Walker, 1859), which he later revised (
<bibRefCitation id="868E4F16703B3CAE49EFCADCE02C1E10" author="Hampson, GF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="143" refId="B11" refString="Hampson, GF, 1903. The moths of India. Supplementary paper to the volumes in &quot;The fauna of British India.&quot; Series II. Part IX, X. The Journal of the Bombay Natural History Society 15: part IX: 19-37; part X: 206-226. [pl. C.]" title="The moths of India. Supplementary paper to the volumes in &quot; The fauna of British India. &quot; Series II. Part IX, X. The Journal of the Bombay Natural History Society 15: part IX: 19 - 37; part X: 206 - 226. [pl. C.]" year="1903">Hampson 1903</bibRefCitation>
) by reinstating it as
<taxonomicName id="7F4A69429412DF2217EBB5F58B5D0275" baseAuthorityName="C.Swinhoe" baseAuthorityYear="1894" class="Insecta" family="Crambidae" genus="Nosophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nosophora incomitata" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="incomitata">
<emphasis id="2E6B532639DCA67C9C3694C9806C2C01" italics="true" pageId="0" pageNumber="143">Nosophora incomitata</emphasis>
</taxonomicName>
, with the junior synonym
<taxonomicName id="54A7CC12A4002DBC2C184294DD8F5AB1" authorityName="Warren" authorityYear="1896" class="Insecta" family="Crambidae" genus="Nosophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nosophora triguttalis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="triguttalis">
<emphasis id="71F2889658311EACFF3793BCDD0A7280" italics="true" pageId="0" pageNumber="143">Nosophora triguttalis</emphasis>
</taxonomicName>
Warren, 1896. In the same publication on page 216,
<bibRefCitation id="0E6307609F05827CB1AB88D683B6A8D9" author="Hampson, GF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="143" refId="B11" refString="Hampson, GF, 1903. The moths of India. Supplementary paper to the volumes in &quot;The fauna of British India.&quot; Series II. Part IX, X. The Journal of the Bombay Natural History Society 15: part IX: 19-37; part X: 206-226. [pl. C.]" title="The moths of India. Supplementary paper to the volumes in &quot; The fauna of British India. &quot; Series II. Part IX, X. The Journal of the Bombay Natural History Society 15: part IX: 19 - 37; part X: 206 - 226. [pl. C.]" year="1903">Hampson (1903)</bibRefCitation>
synonymised the males of
<taxonomicName id="7833565FA246F4475F44445C177A5044" lsidName="N. incomitata" pageId="0" pageNumber="143" rank="species" species="incomitata">
<emphasis id="C0503F9BB74D4EB0462E426D04A45F7F" italics="true" pageId="0" pageNumber="143">N. incomitata</emphasis>
</taxonomicName>
with
<taxonomicName id="A2962366D211D2DA25010996655AD405" authorityName="Hubner" authorityYear="1825" class="Insecta" family="Crambidae" genus="Sylepta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sylepta" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="8922AEACA29ADE044DD2773269C72B47" italics="true" pageId="0" pageNumber="143">Sylepta</emphasis>
</taxonomicName>
[sic]
<taxonomicName id="EDDE5897DD86C57B710483403C7B15F7" class="Insecta" family="Crambidae" genus="Sylepta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="quadrimaculalis" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="quadrimaculalis">
<emphasis id="3553936CBB5561F950B24CDC1ABB825B" italics="true" pageId="0" pageNumber="143">quadrimaculalis</emphasis>
</taxonomicName>
. For the time being (i.e., until the type material has been investigated), we conclude as
<bibRefCitation id="DCEF4BA8894AB2E15C52A6D80DD2A230" author="Hampson, GF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="143" refId="B11" refString="Hampson, GF, 1903. The moths of India. Supplementary paper to the volumes in &quot;The fauna of British India.&quot; Series II. Part IX, X. The Journal of the Bombay Natural History Society 15: part IX: 19-37; part X: 206-226. [pl. C.]" title="The moths of India. Supplementary paper to the volumes in &quot; The fauna of British India. &quot; Series II. Part IX, X. The Journal of the Bombay Natural History Society 15: part IX: 19 - 37; part X: 206 - 226. [pl. C.]" year="1903">Hampson (1903)</bibRefCitation>
,
<bibRefCitation id="FD6AE824F809F2CF49C545ADAD45573E" DOI="https://doi.org/10.5962/bhl.title.124016" author="Klima, A" editor="Bryk, F" journalOrPublisher="Junk, s' Gravenhage" pageId="0" pageNumber="143" pagination="3 - 224" refId="B20" refString="Klima, A, 1939b. Pyralididae: Subfamily: Pyraustinae I. In: Bryk, F, Ed., Lepidopterorum Catalogous. Dr. W. Junk, s' Gravenhage 89: 3 - 224, DOI: https://doi.org/10.5962/bhl.title.124016" title="Pyralididae: Subfamily: Pyraustinae I." url="https://doi.org/10.5962/bhl.title.124016" volume="89" volumeTitle="Lepidopterorum Catalogous. Dr. W." year="1939 b">Klima (1939b)</bibRefCitation>
, and
<bibRefCitation id="17AB5E8D8DFFBA86FCBCB296A111E78F" author="Mandal, DK" journalOrPublisher="Records of the Zoological Survey of India" pageId="0" pageNumber="143" pagination="293 - 342" refId="B26" refString="Mandal, DK, Bhattacharya, DP, 1980. On the Pyraustinae (Lepidoptera: Pyralidae) from the Andaman, Nicobar and Great Nicobar islands, Indian ocean. Records of the Zoological Survey of India 77: 293 - 342" title="On the Pyraustinae (Lepidoptera: Pyralidae) from the Andaman, Nicobar and Great Nicobar islands, Indian ocean." volume="77" year="1980">Mandal and Bhattacharya (1980)</bibRefCitation>
, who considered
<taxonomicName id="137AFABC3AE4B1B49ABB8B6452A86724" class="Insecta" family="Crambidae" genus="Sylepta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="incomitata" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="incomitata">
<emphasis id="E969568893C51CF19CBCBB85A078AFD3" italics="true" pageId="0" pageNumber="143">incomitata</emphasis>
</taxonomicName>
a species of
<taxonomicName id="36544B8176F874829B815EC7DC9FF525" authorityName="Lederer" authorityYear="1863" class="Insecta" family="Crambidae" genus="Nosophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nosophora" order="Lepidoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="F59A05F5A496F9262E8CC8CF001617A7" italics="true" pageId="0" pageNumber="143">Nosophora</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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