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<mods:title id="B9F3FD33E78DE7E880A073BF1F538825">News from Baltic amber: a new tribe of extinct Paleogene ironclad beetles (Coleoptera: Zopheridae: Zopherinae)</mods:title>
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<mods:namePart id="9C517698611024FBF551B037831FF8ED">Alekseev, Vitalii I.</mods:namePart>
<mods:affiliation id="005FE6B76D83223E1372D6C3868A9092">Kaliningrad Regional Amber Museum, Marshal Vasilevskii square 1, Kaliningrad 236016, Russia. &amp; Immanuel Kant Baltic Federal University, A. Nevskii str. 14, Kaliningrad 236016, Russia</mods:affiliation>
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<mods:namePart id="3EB92BF4030E631D87A331F8917F7E00">Nabozhenko, Maxim V.</mods:namePart>
<mods:affiliation id="ED577626B8A43BA71F6E54311F767758">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str. 45, Makhachkala, Republic of Dagestan, 367000, Russia. &amp; Dagestan State University, M. Gadzhiev str. 43 a, Makhachkala, Republic of Dagestan, 367000, Russia</mods:affiliation>
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<heading id="214881A0CB35771BFF4E328AFDDAFA4A" bold="true" box="[151,584,1446,1472]" fontSize="11" level="1" pageId="1" pageNumber="436" reason="1">
<emphasis id="48CBEADECB35771BFF4E328AFDDAFA4A" bold="true" box="[151,584,1446,1472]" pageId="1" pageNumber="436">
Subfamily
<taxonomicName id="BDBF4D4FCB35771BFEC6328AFDDAFA4A" authority="Solier, 1834" authorityName="Solier" authorityYear="1834" box="[287,584,1446,1472]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="subFamily" subFamily="Zopherinae">Zopherinae Solier, 1834</taxonomicName>
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<emphasis id="48CBEADECB35771BFF4E32C3FF44F983" bold="true" box="[151,214,1519,1545]" pageId="1" pageNumber="436">Note.</emphasis>
The beetle specimen in amber piece No KAM 8325/2 possesses the following set of morphological characters: (1) antennal insertions concealed under a lateral expansion of the frons and not visible in dorsal view; (2) 55 4 tarsal formula; (3) intercoxal prosternal process expanded laterally at apex; (4) antennae 11-segmented with indistinct, weak 2-segmented club; (5) abdominal ventrite 5 with paired preapical grooves; (6) mesocoxal cavities closed laterally; (7) prosternum separated from hypomera by complete notosternal sutures.
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<paragraph id="7A0036CCCB35771BFF1E318FFCF9F88F" blockId="1.[151,1437,1518,1941]" pageId="1" pageNumber="436">
The following features make it possible to distinguish the new species from
<taxonomicName id="BDBF4D4FCB35771BFBF4318FFB43F937" box="[1069,1233,1699,1725]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
: (1) outer edge of the abdominal ventrites is not formed into a groove which receives the elytral edges; (2) abdomen with ventrites separated by narrow band of sclerotized membrane, not connate.
</paragraph>
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The combination of these features in the fossil beetle allows it to be assigned to the subfamily
<taxonomicName id="BDBF4D4FCB35771BFB343023FAFFF8A3" authorityName="Solier" authorityYear="1834" box="[1261,1389,1807,1833]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="subFamily" subFamily="Zopherinae">Zopherinae</taxonomicName>
and excluded from other externally similar groups (e.g.,
<taxonomicName id="BDBF4D4FCB35771BFD36301FFC00F8C7" box="[751,914,1843,1869]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
,
<taxonomicName id="BDBF4D4FCB35771BFC46301FFB9BF8C7" authorityName="Pascoe" authorityYear="1869" box="[927,1033,1843,1869]" class="Insecta" family="Ulodidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="family">Ulodidae</taxonomicName>
,
<taxonomicName id="BDBF4D4FCB35771BFBCF301FFB04F8C7" authorityName="Erichson" authorityYear="1842" box="[1046,1174,1843,1869]" class="Insecta" family="Colydiinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="family">Colydiinae</taxonomicName>
). On the basis of these and other (see below) diagnostic characters, the specimen cannot be definitely attributed to any existing tribes of the subfamily and should be placed, in our opinion, in a separate new tribe.
</paragraph>
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