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<document id="2EEDDD303741006E4745ED37D25B041E" ID-CLB-Dataset="302074" ID-DOI="10.3897/phytokeys.246.129476" ID-GBIF-Dataset="b381880a-e0bb-4b02-ad0f-2f90a0b6c995" ID-publisher-id="129476" URI-arpha="7631F16E-4CD8-5A76-9F28-45E5F398CE20" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1725298343909" checkinUser="pensoft" docAuthor="Meerts, Pierre J. &amp; Paton, Alan J." docDate="2024" docId="F171D4DB6E3B55D48BADBC0AB3933BC4" docLanguage="en" docName="PhytoKeys 246: 71-178" docOrigin="PhytoKeys 246" docSource="https://phytokeys.pensoft.net/article/129476/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Coleus esculentus var. densus Meerts &amp; A. J. Paton 2024, comb. et stat. nov." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="178" masterDocId="7631F16E4CD85A769F2845E5F398CE20" masterDocTitle="The genus Coleus (Lamiaceae) in Central Africa (Democratic Republic of the Congo, Rwanda, Burundi), with the description of 15 new species" masterLastPageNumber="178" masterPageNumber="71" pageNumber="71" updateTime="1725322260938" updateUser="ExternalLinkService">
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<mods:title id="E3F78E18E2931FD12E14EFE27174210C">The genus Coleus (Lamiaceae) in Central Africa (Democratic Republic of the Congo, Rwanda, Burundi), with the description of 15 new species</mods:title>
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<mods:namePart id="4C5169BF54B294CFD81F99C150BDB894">Meerts, Pierre J.</mods:namePart>
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<mods:affiliation id="B9BD458DC4052F3BFD8C7F48C2FA1357">Meise Botanic Garden, Nieuwelaan 38, BE- 1860 Meise, Belgium &amp; Université Libre de Bruxelles, Av. F. D. Roosevelt 50 CP 244, BE- 1050 Brussels, Belgium &amp; Fédération Wallonie-Bruxelles, Service général de lEnseignement supérieur et de la Recherche scientifique, Brussels, Belgium</mods:affiliation>
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<mods:namePart id="D9AABC3A4E7586DA238B1630470E8E62">Paton, Alan J.</mods:namePart>
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<mods:affiliation id="0D87EEE5A5F8B123BF69348EA495A8B6">The Herbarium, Royal Botanic Gardens, Kew, Richmond TW 9 3 AB, UK</mods:affiliation>
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<mods:title id="00BF1D0C4BA94A3025D00F1AC816CB70">PhytoKeys</mods:title>
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<mods:date id="92F72F9B5F08F1C60E315DA9F6D4C85D">2024</mods:date>
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<treatment id="F171D4DB6E3B55D48BADBC0AB3933BC4" ID-DOI="http://doi.org/10.5281/zenodo.13629791" ID-GBIF-Taxon="240061422" ID-Zenodo-Dep="13629791" ID-arpha="F171D4DB-6E3B-55D4-8BAD-BC0AB3933BC4" ID-ipni="urn:lsid:ipni.org:names:77347690-1" LSID="urn:lsid:plazi:treatment:F171D4DB6E3B55D48BADBC0AB3933BC4" LSID-IPNI="urn:lsid:ipni.org:names:77347690-1" httpUri="http://treatment.plazi.org/id/F171D4DB6E3B55D48BADBC0AB3933BC4">
<subSubSection id="F0FF666B9002FCDCFDBE2BC81E68815B" type="nomenclature">
<paragraph id="41A6ED8EF83B499E4DAB59DBBFF3BB57">
<taxonomicName id="8608B435FC480B59D37DC92DBE01CEA6" ID-CoL="8XY4T" authority="(N. E. Br.) Meerts &amp; A. J. Paton" authorityName="Meerts &amp; A. J. Paton" authorityYear="2024" baseAuthorityName="Meerts &amp; A. J. Paton" baseAuthorityYear="2024" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="variety" species="esculentus" status="comb. et stat. nov." variety="densus">
<emphasis id="A9E5CDEA672FF0F059A29FE96EE57F0D" italics="true">Coleus esculentus var. densus</emphasis>
(N. E. Br.) Meerts &amp; A. J. Paton
</taxonomicName>
,
<taxonomicNameLabel id="16C5AE6955301D7D5102D8034E3C949C">comb. et stat. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="645DA60BD71CE75263E9F3EDA4428F71" type="reference_group">
<paragraph id="3133A7A8C594DAF55F62AFCBAAB36D0C">
<treatmentCitationGroup id="C5EA4E30CB220040A7FCCA038A09FFDE">
<treatmentCitation id="D5F8C6E04C69B7084EDDBF1FD17C79EA" author="N. E. Brown" issue="85" journal="Bull. Misc. Inform., Kew" page="12" score="2.5435888525040795" volume="1894" year="1894">
<taxonomicName id="FB143FF3134F14798BC4CCD0BBEC4C3E" ID-CoL="4K478" authority="N. E. Brown" authorityName="N. E. Brown" class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="densus">
<emphasis id="27DC9BAEA161EDE843C11F4C84F76546" italics="true">Plectranthus densus</emphasis>
N. E. Brown
</taxonomicName>
in
<bibCitation id="EFB99F10DA8FAA380F45C95E89F3D0E1" issue="85" journal="Bull. Misc. Inform., Kew" page="12" score="2.5435888525040795" volume="1894" year="1894">Bull. Misc. Inform., Kew 1894 (85): 12. 1894</bibCitation>
</treatmentCitation>
. Type:
<collectingCountry id="784AECB56415157065F1CB5F3C732989" name="Tanzania">Tanzania</collectingCountry>
, N of Lake
<collectingCountry id="620C53042511C4B6F83B13B86A325C65" name="Malawi">Malawi</collectingCountry>
(Nyassa),
<date id="41FE4DA73C43A87189228C8139B43294" value="1880-10">Oct 1880</date>
,
<emphasis id="216D20F68969A650B66B94D327123869" italics="true">Thomson s. n</emphasis>
. (
<typeStatus id="69110FC802B7D6F7EC255541DE972676" type="holotype">holotype</typeStatus>
<collectionCode id="E183720F728CCADD243F1C90D8BF2DBC" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
[
<specimenCode id="CE68ACBD8499824294E4876543C4BB77">
<collectionCode id="2BFAB5829C6A6C843A7E59DB0CC73DE1" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
000431965
</specimenCode>
]). (Basionym)
</treatmentCitationGroup>
</paragraph>
<paragraph id="AC5EECBEC3C1DC28254595D456FBEA5D">
<treatmentCitationGroup id="A12430523816530D15C73353562E05D9">
<treatmentCitation id="F0A2A16B89EDA115B1C6BA0746AF093C" author="A. J. Paton" httpUri="http://treatment.plazi.org/id/7A7C9B98C00653639F74EEA8F02506D1" journal="Phytokeys" page="39" score="9.729594737742866" volume="129" year="2019">
<taxonomicName id="A8EC2A87639F1B9405A0C44840FE05BC" ID-CoL="8XY4T" authority="(N. E. Br.) A. J. Paton" authorityName="A. J. Paton" baseAuthorityName="N. E. Br." class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="densus">
<emphasis id="964AC80B0F5461C4AF759C1DFF93F290" italics="true">Coleus densus</emphasis>
(N. E. Br.) A. J. Paton
</taxonomicName>
, Phytokeys 129: 39. 2019
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="E64A806829CEE42D2FB21D37B0A1A8CF">
<treatmentCitationGroup id="74D775E69982887C366354257350B36B">
=
<treatmentCitation id="F3FD5ED6E93785E725DD8D7DBE3FB530" author="" journal="De Wild. Ann. Mus. Congo Belge, Bot." page="135" score="54.002150658740284" volume="2" year="1913">
<taxonomicName id="6BAD294491439CEEEA5932525CFE98CC" ID-CoL="4K475" authority="De Wild." authorityName="De Wild." class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="dekindtianus">
<emphasis id="513FC05AE1936FF1A2ADC930DF083B39" italics="true">Plectranthus dekindtianus</emphasis>
De Wild.
</taxonomicName>
Ann. Mus.
<bibCitation id="3683228A43672574B26CA5D805998BC5" author="Congo Belge" journal="Bot." page="135" score="0.23648317473498365" volume="2" year="1913">
<collectingCountry id="94BC5A3CA824DB6CA31C02C2F6A0FFCA" name="Democratic Republic of the Congo">Congo Belge</collectingCountry>
, Bot., sér. 4, 2: 135. 1913
</bibCitation>
</treatmentCitation>
. Type:
<collectingCountry id="671B6D6D1A6330912F3EF965BF17C62A">DR. Congo</collectingCountry>
, Bugege (?),
<date id="3983C050B9B341E6FA6590C1E43A6205" value="1911-09">Sep 1911</date>
.
<emphasis id="54B043A535FCA28C1EB505C70D17E2AA" italics="true">A. Hock s. n</emphasis>
. (
<typeStatus id="BB1CB40AC0FBE22520D738117C353CF4" type="holotype">holotype</typeStatus>
<collectionCode id="638EBECE2C305E33D056ECA2AD2786E0" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
[
<specimenCode id="87D5FADEC0585DA8D0C5FCC9AC26FCAE">
<collectionCode id="B8CAF2B7930E62B9894533B294B8AD25" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
0000009824928
</specimenCode>
]), syn. nov.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWUFK" type="description">
<paragraph id="2C2A09EC54C2682CEE8172976D0E4A6E">
<heading id="7D6E642A6E2D1C73AA0D9C8A733145BB" reason="title">Description.</heading>
</paragraph>
<paragraph id="D3834C935054ACC79586ACDA85DC8438">
<bibRefCitation id="0D0C6D99F170CDC0E17985D61C456796" author="Paton AJ &amp; Bramley G &amp; Ryding O &amp; Polhill RM &amp; Harvey YB &amp; Iwarsson M &amp; Willis F &amp; Phillipson PB &amp; Balkwill K &amp; Lukhoba CW &amp; Otieno D &amp; Harley RM" editor="Beentje HJ &amp; Ghazanfar SA &amp; Polhill RM" etAl="et al." firstAuthor="Paton" journalOrPublisher="Royal Botanic Gardens, Kew, Richmond" page="292" pagination="1-430" refId="B20" refString="Paton AJ, Bramley G, Ryding O, Polhill RM, Harvey YB, Iwarsson M, Willis F, Phillipson PB, Balkwill K, Lukhoba CW, Otieno D, Harley RM (2009) Lamiaceae (Labiatae). In: Beentje HJ, Ghazanfar SA, Polhill RM (Eds) Flora of Tropical East Africa. Royal Botanic Gardens, Kew, Richmond, 1430." title="Lamiaceae (Labiatae)." volumeTitle="Flora of Tropical East Africa." year="2009">Paton et al. (2009: 292)</bibRefCitation>
,
<bibRefCitation id="6F4B76EBF65B71D367823D60E1F2B6C0" author="Paton AJ &amp; Bramley G &amp; Ryding O &amp; Polhill RM &amp; Harvey YB &amp; Iwarsson M &amp; Willis F &amp; Phillipson PB &amp; Balkwill K &amp; Lukhoba CW &amp; Otieno D &amp; Harley RM" editor="Timberlake J" etAl="et al." firstAuthor="Paton" journalOrPublisher="Royal Botanic Gardens, Kew, Richmond" page="241" pagination="1-346" refId="B21" refString="Paton AJ, Bramley G, Ryding O, Polhill RM, Harvey YB, Iwarsson M, Willis F, Phillipson PB, Balkwill K, Lukhoba CW, Otieno D, Harley RM (2013) Lamiaceae. In: Timberlake J (Ed.) Flora Zambesiaca 8 (8). Royal Botanic Gardens, Kew, Richmond, 1346." title="Lamiaceae." volumeTitle="Flora Zambesiaca 8 (8)." year="2013">Paton et al. (2013: 241)</bibRefCitation>
, as
<taxonomicName id="E1A35AAF3B14E5C92C8CA06E37F44A6B" authority="N. E. Br." authorityName="N. E. Br." class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="densus">
<emphasis id="ACF0C338C623838E39DF20B0EC6C8184" italics="true">Plectranthus densus</emphasis>
N. E. Br.
</taxonomicName>
, restricted to specimens with inflorescence lax.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPVFK" type="distribution">
<paragraph id="B0F9BA87EE150AF7A5646A610739C02E">
<heading id="0B5498D04152AE079527AF17D05B8EB1" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="A5857F40A45659D72ED7BEE4BCCC10EC">
<collectionCode id="AB78B96F76E880130F7994FC68B5992C">SW</collectionCode>
<collectingCountry id="79157FE205E5B34C43FDFF7441BD5EE0" name="Tanzania">Tanzania</collectingCountry>
to S Tropical Africa.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUVFK" type="habitat">
<paragraph id="8EB9486E3A30C5B5C8234EE42DBD3467">
<heading id="86C2810D34613181834DD9A533DD3151" reason="title">Habitat and ecology.</heading>
</paragraph>
<paragraph id="2ED8E2C75CC3F61A47470A2E3C2086A3">
Dry woodlands, savannah,
<quantity id="1576CAA4A8A08FCD816E74D5D345A92F" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="2.0" metricValueMin="1.3" unit="m" value="1650.0" valueMax="2000.0" valueMin="1300.0">13002000 m</quantity>
elev.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZVFK" type="Additional specimens">
<paragraph id="C8CFED42B810341A06B0EBE191D2ED6C">
<heading id="3C1289B5C01623B4DFF329E72672FA2D" reason="title">Additional specimens.</heading>
</paragraph>
<paragraph id="3EF300C55FE1062A3B0B1F09323891E7">
<collectingCountry id="6CC1C10457D92638EF6932A187400CEC">
DR.
<collectingRegion id="B1B740AD439E63F08DB0E3FEAEAB6B61" country="Republic of the Congo" name="Bas-Congo">Congo</collectingRegion>
</collectingCountry>
,
<emphasis id="4CB767155E10E0E63388BB14621EC4A7" bold="true" italics="true">
<collectingRegion id="AE221CE23E5A1A91D80136F03A2D40FC" country="Republic of the Congo" name="Kasai-Oriental">Kasaï</collectingRegion>
</emphasis>
, Kahemba, route vers lAngola,
<date id="5AD0533B0F5FF251790E54C691EA366E" value="1948-05-31">31 May 1948</date>
,
<emphasis id="658BB4CE6419CBFA39D677A28047498E" italics="true">P. Duvigneaud 955</emphasis>
(
<collectionCode id="2DEC7B20A79529248C62FA998B642C19" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14449" lsid="urn:lsid:biocol.org:col:14449" name="Université Libre de Bruxelles" type="Herbarium">BRLU</collectionCode>
);
<emphasis id="57F3D9913FC11CCFDC3882306B01F695" bold="true" italics="true">
<collectingRegion id="6F4B2135428949EAB777853A46262C42">Haut-Katanga</collectingRegion>
</emphasis>
, Kambove,
<date id="8AC1A12E95445F091358EE6B9465EC39" value="1979-08-17">17 Aug 1979</date>
,
<emphasis id="19789B6B992887BEB057199378EFA47F" italics="true">
<collectionCode id="9C50991535842ACC3A0B9CD6B167B936" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" type="Herbarium">H</collectionCode>
. Breyne 3735
</emphasis>
(
<collectingRegion id="1807B59A9C5FCEFA9A8E959CBD19641C" country="Serbia" name="Bor">BR</collectingRegion>
); Lubumbashi, Kasapa,
<date id="EC85563C67AEEBB8992CFC436B68BD4E" value="1972-09-15">15 Sep 1972</date>
,
<emphasis id="AA0D1E204940863F5D2755AA7C1816BE" italics="true">J. Bulaimu 533</emphasis>
(
<collectingRegion id="B1A07C627FA14F64ADB2B3D17FF5A131" country="Serbia" name="Bor">BR</collectingRegion>
); Upemba, piste vers Mitwaba,
<date id="4BC18B14BF1DFF4B93DAD181395A50FA" value="1947-04-12">12 Apr 1947</date>
,
<emphasis id="F18B339ABBCEA9FE65843C930428DC0D" italics="true">
<collectionCode id="020C07D69C556B70611DAAB0F1F4B366" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. F. de Witte 2488
</emphasis>
(
<collectingRegion id="DEB5F48D57EA24873590B8797D6DE8F4" country="Serbia" name="Bor">BR</collectingRegion>
);
<quantity id="CB6747D35F879C3EFA8DA5688B1B6410" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
<collectionCode id="E6314C01E191395572BF53E94A7B36ED" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" type="Herbarium">W</collectionCode>
of Mindingi,
<date id="6D9E8647BFF5F566CC32B47449D30C6A" value="1956-07-21">21 Jul 1956</date>
,
<emphasis id="0313B990E11F0EED58AF220D04E0C470" italics="true">P. Duvigneaud &amp; J. Timperman 2067</emphasis>
(
<collectionCode id="4E8CBA2A68486C81F8C6683C5734FD6F" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14449" lsid="urn:lsid:biocol.org:col:14449" name="Université Libre de Bruxelles" type="Herbarium">BRLU</collectionCode>
); Upemba, entre Masombwe et Lusinga,
<date id="D95EEFF4F029551CD68CAE91B9DDB599" value="1948-09-21">21 Sep 1948</date>
,
<emphasis id="0B1BB89BB600483C6C01A301CE36FB98" italics="true">J. Lebrun 3644</emphasis>
(
<collectingRegion id="8957B4196A771FC506A4D2047E9065CE" country="Serbia" name="Bor">BR</collectingRegion>
); Luiswishi,
<date id="77C878B9E36A843E855872FE7AC78D6F" value="1974-10-02">2 Oct 1974</date>
,
<emphasis id="1DF3D59E3F8A57E6914D8B77B823D4C0" italics="true">F. Malaisse 7965</emphasis>
(
<collectingRegion id="4C5796B76CCBB9331F46F48C43C45754" country="Serbia" name="Bor">BR</collectingRegion>
);
<collectingRegion id="4C611F504B7AF62DC9FF191ED525D33E" country="Republic of the Congo" name="Katanga">Katanga</collectingRegion>
, s. l., 21 Sep 58,
<emphasis id="CC9A69729E4D7ED7D03554137AD6F5CB" italics="true">J. Plancke 119 / 1734</emphasis>
(
<collectionCode id="6E33DBBDF1994BC43B8779B811F78F33" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14449" lsid="urn:lsid:biocol.org:col:14449" name="Université Libre de Bruxelles" type="Herbarium">BRLU</collectionCode>
); Upemba,
<date id="4DCE55481DD45465D54FD588A2D4186F" value="1949-08-25">25 Aug 1949</date>
,
<emphasis id="642DD80D02D237CA56B2381DDD78D429" italics="true">L. van Meel</emphasis>
in
<emphasis id="12C3CBC70B56B273D437A3CF27F77030" italics="true">
<collectionCode id="1EE066B4097C5F552BE0EF4BDD7BB6FE" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
. F. de Witte 7555
</emphasis>
(
<collectingRegion id="723D9665B36F49FF1A7B4812973C10E7" country="Serbia" name="Bor">BR</collectingRegion>
).
</paragraph>
<paragraph id="6420D74D05D367C4E1CECB128B7E1CCE">
<collectingCountry id="37A7CD47AB4F29E3BA4968EE78640065" name="Burundi">Burundi</collectingCountry>
.
<collectingRegion id="CE23DE0D8F48433F9E33E52069A8E630" country="Burundi" name="Bururi">Bururi</collectingRegion>
,
<date id="128C8391FC84491F7A6860BA7B4F8F84" value="1974-09-21">21 Sep 1974</date>
,
<emphasis id="6FA90822EAD1ED918F004230C5EC2950" italics="true">J. Rammeloo 4690</emphasis>
(
<collectionCode id="D8BCC0FD82E207BD5BC445EA2DBD3D9F" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJXFK" type="notes">
<paragraph id="82DC7EC2B7E11C8FE24AD6B193888F05">
<heading id="513FCE96D89B4564270EFA8888A677E8" reason="title">Note.</heading>
</paragraph>
<paragraph id="B9D89293F02BA00F98A6B9A6B0538FC5">
<taxonomicName id="AF85C43791E044A5ED511875E5AFEDAE" authority="De Wild." authorityName="De Wild." class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="dekindtianus">
<emphasis id="D3FAB943C84EA36E54A07737EAF27BEA" italics="true">Plectranthus dekindtianus</emphasis>
De Wild.
</taxonomicName>
was synonymised with
<taxonomicName id="01C524F407B2B2DF89C5C91021C98EE6" authority="Gurke" authorityName="Gurke" class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="tetragonus">
<emphasis id="DDED53FFEA0CEFB8F19CE17580D93A38" italics="true">Plectranthus tetragonus</emphasis>
Gürke
</taxonomicName>
by
<bibRefCitation id="EC284CDC4BDF11BBF85571B0F467A44F" author="Paton AJ &amp; Bramley G &amp; Ryding O &amp; Polhill RM &amp; Harvey YB &amp; Iwarsson M &amp; Willis F &amp; Phillipson PB &amp; Balkwill K &amp; Lukhoba CW &amp; Otieno D &amp; Harley RM" editor="Beentje HJ &amp; Ghazanfar SA &amp; Polhill RM" etAl="et al." firstAuthor="Paton" journalOrPublisher="Royal Botanic Gardens, Kew, Richmond" pagination="1-430" refId="B20" refString="Paton AJ, Bramley G, Ryding O, Polhill RM, Harvey YB, Iwarsson M, Willis F, Phillipson PB, Balkwill K, Lukhoba CW, Otieno D, Harley RM (2009) Lamiaceae (Labiatae). In: Beentje HJ, Ghazanfar SA, Polhill RM (Eds) Flora of Tropical East Africa. Royal Botanic Gardens, Kew, Richmond, 1430." title="Lamiaceae (Labiatae)." volumeTitle="Flora of Tropical East Africa." year="2009">Paton et al. (2009)</bibRefCitation>
; however, the type specimen lacks bristles on the stem and is better placed here
</paragraph>
</subSubSection>
</treatment>
</document>