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<document id="80B1C78B9B4A33AE4C5F574D4D1F676D" ID-DOI="10.1649/0010-065X-70.4.825" ID-Zenodo-Dep="5340413" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630186311507" checkinUser="diego" docAuthor="Hisamatsu, Sadatomo, Bayless, Victoria M. &amp; Carlton, Christopher E." docDate="2016" docId="F042A324FFCAFFBA0ABFE91DFCEA8840" docLanguage="en" docName="0010-065X-70.4.825.pdf" docOrigin="The Coleopterists Bulletin 70 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-70.4.825" docStyle="DocumentStyle:A0A70CFE30726088A421CCB226FF86C7.4:TheColeopBull.2010-2016.journal_article.1cover" docStyleId="A0A70CFE30726088A421CCB226FF86C7" docStyleName="TheColeopBull.2010-2016.journal_article.1cover" docStyleVersion="4" docTitle="Cyclocaccus epakros Hisamatsu 2016, new species" docType="treatment" docVersion="3" lastPageNumber="870" masterDocId="0C7BDB5CFFD3FFA30A01E84CFFFD8C1E" masterDocTitle="Revision ofCyclocaccusSharp (Coleoptera: Nitidulidae: Nitidulinae)" masterLastPageNumber="870" masterPageNumber="825" pageNumber="825" updateTime="1699179913154" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0A61AB937C92859254749BD510E079A8">Revision ofCyclocaccusSharp (Coleoptera: Nitidulidae: Nitidulinae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="DC7F232CCAB974552276114F78D1146C">Hisamatsu, Sadatomo</mods:namePart>
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<mods:namePart id="ECB57D5E995B22C5073EE966B8B1919B">Bayless, Victoria M.</mods:namePart>
</mods:name>
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<mods:namePart id="77D50E55BD26EDE275776387A711F960">Carlton, Christopher E.</mods:namePart>
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<mods:typeOfResource id="34487E40C1559E9FB830862BDA4415A8">text</mods:typeOfResource>
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<mods:titleInfo id="5439669F2A6EFEDD921F038B8762D6CE">
<mods:title id="4FD297B0FD840D3265A71AF105BDB662">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="29EEA9A06EB42A88EDC611E0173EA852">
<mods:date id="EF9671D6129BA46DAF3DCBE4C6154B0D">2016</mods:date>
<mods:detail id="4A7F974CA43B61B7831CEBE48378592A" type="pubDate">
<mods:number id="40BCE2F143295F86D0C99646CEE32CAD">2016-12-31</mods:number>
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<mods:number id="285BE6CD215B442A4CAAE1ED2F478F37">70</mods:number>
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<mods:number id="7AB47CF2D39BC0B37FE408225F42698F">4</mods:number>
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<mods:url id="9545A8354591D040169EFAA9D76138B1">http://dx.doi.org/10.1649/0010-065x-70.4.825</mods:url>
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<mods:classification id="28B815D0EECD5ACD7D35D3F44EB4DA48">journal article</mods:classification>
<mods:identifier id="E4920DB05C682422FEABABD9E4C26CB4" type="DOI">10.1649/0010-065X-70.4.825</mods:identifier>
<mods:identifier id="1DA3ECF57B62D1BE0F08D00D3D6588F6" type="Zenodo-Dep">5340413</mods:identifier>
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<treatment id="F042A324FFCAFFBA0ABFE91DFCEA8840" LSID="urn:lsid:plazi:treatment:F042A324FFCAFFBA0ABFE91DFCEA8840" httpUri="http://treatment.plazi.org/id/F042A324FFCAFFBA0ABFE91DFCEA8840" pageId="25">
<subSubSection id="30F141B9FFCAFFBA0ABFE91DFDFF8D9F" pageId="25" type="nomenclature">
<paragraph id="78541232FFCAFFBA0ABFE91DFDFF8D9F" blockId="25.[190,640,337,385]" pageId="25">
<emphasis id="4A9FCE20FFCAFFBA0ABFE91DFD7D8D78" bold="true" box="[190,640,337,358]" pageId="25">
<taxonomicName id="BFEB69B1FFCAFFBA0ABFE91DFE028D78" authority="Hisamatsu" authorityName="Hisamatsu" authorityYear="2016" box="[190,511,337,358]" class="Insecta" family="Nitidulidae" genus="Cyclocaccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" phylum="Arthropoda" rank="species" species="epakros" status="sp. nov.">
<emphasis id="4A9FCE20FFCAFFBA0ABFE91DFE718D78" bold="true" box="[190,396,337,358]" italics="true" pageId="25">Cyclocaccus epakros</emphasis>
Hisamatsu
</taxonomicName>
,
<taxonomicNameLabel id="51AC735BFFCAFFBA080DE91DFD7D8D78" box="[524,640,337,358]" pageId="25" rank="species">new species</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="E0D00EB7FFCAFFBA0B45E920FE6A8D9F" box="[324,407,364,385]" captionStart="Fig" captionStartId="7.[197,228,1638,1657]" captionTargetBox="[227,1149,194,1609]" captionTargetId="figure-0@7.[212,1153,194,1614]" captionTargetPageId="7" captionText="Fig. 1. Dorsal habitus of Cyclocaccus species in the C. laeticulus species-group. A) C. clinei, holotype, B) C. laeticulus, holotype, C) C. laeticulus, holotype, labels, D) C. laeticulus, Venezuela, E) C. speciosus, holotype, F) C. monticola, lectotype, G) C. monticola, lectotype, labels, H) C. monticola, Guatemala, I) C. monticola, Bolivia, J) C. costaricensis, holotype, K) C. maculatus, holotype, L) C. epakros, holotype. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340415" httpUri="https://zenodo.org/record/5340415/files/figure.png" pageId="25">Figs. 1L</figureCitation>
,
<figureCitation id="E0D00EB7FFCAFFBA0BA2E920FE4C8D9F" box="[419,433,364,385]" captionStart="Fig" captionStartId="15.[197,228,1686,1705]" captionTargetBox="[191,1130,204,1639]" captionTargetId="figure-1@15.[173,1193,183,1662]" captionTargetPageId="15" captionText="Fig. 9. Cyclocaccus epakros, Guerrero, Mexico. A) Tegmen, ventral view, B) Median lobe, ventral view, C) Right antenna, ventral view, D) Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340432" httpUri="https://zenodo.org/record/5340432/files/figure.png" pageId="25">9</figureCitation>
, Map 3)
</paragraph>
</subSubSection>
<subSubSection id="30F141B9FFCAFFBA0AC4E9DBFD6E8E85" pageId="25" type="materials_examined">
<paragraph id="78541232FFCAFFBA0AC4E9DBFD6E8E85" blockId="25.[173,660,406,1729]" pageId="25">
<emphasis id="4A9FCE20FFCAFFBA0AC4E9DBFEC38DB2" bold="true" box="[197,318,407,428]" pageId="25">Type Series.</emphasis>
<materialsCitation id="C883186FFFCAFFBA0B48E9DBFDE68DE2" collectingDate="1992-07-27" collectionCode="SEMC" collectorName="J. S. Ashe &amp; H. Frania" elevation="2040" location="Alvarez" pageId="25" specimenCount="1" specimenCount-male="1" stateProvince="Mexico" typeStatus="holotype">
<typeStatus id="A750AC90FFCAFFBA0B48E9DBFE5C8DB2" box="[329,417,407,428]" pageId="25" type="holotype">Holotype</typeStatus>
(
<collectionCode id="1EFA8AF7FFCAFFBA0BB1E9DBFE0B8DB2" box="[432,502,407,428]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="25" type="Museum">SEMC</collectionCode>
):
<specimenCount id="6EEDD9BBFFCAFFBA0806E9DAFDE48DB2" box="[519,537,406,428]" pageId="25" type="male"></specimenCount>
,
<collectingRegion id="BA2FDCD0FFCAFFBA082EE9DBFD6D8DB2" box="[559,656,407,428]" country="Mexico" name="Mexico" pageId="25">MEXICO</collectingRegion>
:
<collectingRegion id="BA2FDCD0FFCAFFBA0AACE9FDFEFF8DD8" box="[173,258,433,454]" country="Mexico" name="Guerrero" pageId="25">Guerrero</collectingRegion>
/ ca.
<locationDeviation id="69E60B10FFCAFFBA0B38E9FDFE148DD8" box="[313,489,433,454]" location="Atoyac" pageId="25">
<quantity id="BF13BFD7FFCAFFBA0B38E9FDFE8A8DD8" box="[313,375,433,454]" metricMagnitude="4" metricUnit="m" metricValue="7.8" pageId="25" unit="km" value="78.0">78 km</quantity>
NE Atoyac
</locationDeviation>
de /
<location id="7D3444E9FFCAFFBA081CE9FDFD978DD8" LSID="urn:lsid:plazi:treatment:F042A324FFCAFFBA0ABFE91DFCEA8840:7D3444E9FFCAFFBA081CE9FDFD978DD8" box="[541,618,433,454]" name="Alvarez" pageId="25" stateProvince="Mexico">Alvarez</location>
,
<date id="0C5534F2FFCAFFBA0874E9FDFEF18DFF" pageId="25" value="1992-07-27">
<collectingDate id="1C11CD1AFFCAFFBA0874E9FDFEF18DFF" pageId="25" value="1992-07-27">27. VII. 1992</collectingDate>
</date>
/
<quantity id="BF13BFD7FFCAFFBA0B25E980FE508DFF" box="[292,429,459,481]" metricMagnitude="3" metricUnit="m" metricValue="2.04" metricValueMax="2.12" metricValueMin="1.96" pageId="25" unit="m" value="2040.0" valueMax="2120.0" valueMin="1960.0">
<elevation id="F3C6F501FFCAFFBA0B25E980FE508DFF" box="[292,429,459,481]" metricMagnitude="3" metricUnit="m" metricValue="2.04" metricValueMax="2.12" metricValueMin="1.96" pageId="25" unit="m" value="2040.0" valueMax="2120.0" valueMin="1960.0">19602120 m</elevation>
</quantity>
,
<collectorName id="D51E77E4FFCAFFBA0BBBE980FDE28DFF" box="[442,543,460,481]" pageId="25">J. S. Ashe</collectorName>
/
<collectorName id="D51E77E4FFCAFFBA0837E980FD6F8DFF" box="[566,658,460,481]" pageId="25">H. Frania</collectorName>
#133. ex: leaf / litter/rotten wood
</materialsCitation>
.
<materialsCitation id="C883186FFFCAFFBA0831E9AAFEFD8E2E" collectingDate="1992-07-27" collectionCode="SEMC" collectorName="J. S. Ashe &amp; H. Frania" country="Mexico" elevation="2040" location="Alvarez" pageId="25" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Mexico" typeStatus="paratype">
<typeStatus id="A750AC90FFCAFFBA0831E9AAFD728DE5" box="[560,655,486,507]" pageId="25" type="paratype">Paratypes</typeStatus>
:
<emphasis id="4A9FCE20FFCAFFBA0AACEA4DFEDD8E08" bold="true" box="[173,288,513,534]" pageId="25">
[
<collectingCountry id="00FC52A2FFCAFFBA0AB3EA4DFEE78E08" box="[178,282,513,534]" name="Mexico" pageId="25">MEXICO</collectingCountry>
]
</emphasis>
<specimenCount id="6EEDD9BBFFCAFFBA0B2EEA4DFEB08E08" box="[303,333,512,534]" pageId="25" type="male">2♂</specimenCount>
<specimenCount id="6EEDD9BBFFCAFFBA0B4DEA4DFE978E08" box="[332,362,512,534]" pageId="25" type="female">1♀</specimenCount>
, same data as the holotype (
<collectionCode id="1EFA8AF7FFCAFFBA0AB2EA57FF058E2E" box="[179,248,539,560]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="25" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C883186FFFCAFFBA0B0FEA57FEFD8E78" collectingDate="1987-07-08" collectionCode="SEMC" collectorName="K. Schaffner" country="Mexico" elevation="2743" location="Guerrero" pageId="25" specimenCount="2" specimenCount-female="2" stateProvince="Guerrero" typeStatus="paratype">
<specimenCount id="6EEDD9BBFFCAFFBA0B0FEA57FED18E2F" box="[270,300,539,561]" pageId="25" type="female">2♀</specimenCount>
,
<quantity id="BF13BFD7FFCAFFBA0B3DEA57FE658E2E" box="[316,408,539,560]" metricMagnitude="4" metricUnit="m" metricValue="1.02998016" pageId="25" unit="mi" value="6.4">6.4 miles</quantity>
southwest,
<collectingRegion id="BA2FDCD0FFCAFFBA0813EA57FD978E2E" box="[530,618,539,560]" country="Mexico" name="Guerrero" pageId="25">Guerrero</collectingRegion>
, ca. alt.
<quantity id="BF13BFD7FFCAFFBA0AD8EA7AFEC98E55" box="[217,308,566,587]" metricMagnitude="3" metricUnit="m" metricValue="2.7432" pageId="25" unit="ft" value="9000.0">
<elevation id="F3C6F501FFCAFFBA0AD8EA7AFEC98E55" box="[217,308,566,587]" metricMagnitude="3" metricUnit="m" metricValue="2.7432" pageId="25" unit="ft" value="9000.0">9,000 ft.</elevation>
</quantity>
,
<date id="0C5534F2FFCAFFBA0B47EA7AFE338E55" box="[326,462,566,587]" pageId="25" value="1987-07-08">
<collectingDate id="1C11CD1AFFCAFFBA0B47EA7AFE338E55" box="[326,462,566,587]" pageId="25" value="1987-07-08">8. VII. 1987</collectingDate>
</date>
,
<collectorName id="D51E77E4FFCAFFBA0BE1EA7AFD9F8E55" box="[480,610,566,587]" pageId="25">K. Schaffner</collectorName>
leg. (
<collectionCode id="1EFA8AF7FFCAFFBA0AB2EA1DFF058E78" box="[179,248,593,614]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="25" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C883186FFFCAFFBA0B0EEA1DFD728E85" collectingDate="1991-09-18" collectionCode="CMN" collectorName="W. San Cristobal &amp; R. Anderson" country="Mexico" county="Cerro" elevation="2750" location="Pico" municipality="Huitepec" pageId="25" specimenCount="1" specimenCount-female="1" stateProvince="Chiapas" typeStatus="paratype">
<specimenCount id="6EEDD9BBFFCAFFBA0B0EEA1DFED08E78" box="[271,301,592,614]" pageId="25" type="female">1♀</specimenCount>
,
<collectingCounty id="91356ABEFFCAFFBA0B3FEA1DFE8B8E78" box="[318,374,593,614]" pageId="25">Cerro</collectingCounty>
,
<collectingMunicipality id="98308848FFCAFFBA0B84EA1DFE278E78" box="[389,474,593,614]" pageId="25">Huitepec</collectingMunicipality>
(
<location id="7D3444E9FFCAFFBA0BECEA1DFDE68E78" LSID="urn:lsid:plazi:treatment:F042A324FFCAFFBA0ABFE91DFCEA8840:7D3444E9FFCAFFBA0BECEA1DFDE68E78" box="[493,539,593,614]" country="Mexico" county="Cerro" municipality="Huitepec" name="Pico" pageId="25" stateProvince="Chiapas">Pico</location>
), ca.
<quantity id="BF13BFD7FFCAFFBA0856EA1DFD728E78" box="[599,655,593,614]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="25" unit="km" value="5.0">5 km</quantity>
,
<collectorName id="D51E77E4FFCAFFBA0AACEA27FEA98E9E" box="[173,340,619,640]" pageId="25">W. San Cristobal</collectorName>
,
<collectingRegion id="BA2FDCD0FFCAFFBA0B5EEA27FE538E9E" box="[351,430,619,640]" country="Mexico" name="Chiapas" pageId="25">Chiapas</collectingRegion>
, alt.
<quantity id="BF13BFD7FFCAFFBA0BDCEA27FDD28E9E" box="[477,559,619,640]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="25" unit="m" value="2750.0">
<elevation id="F3C6F501FFCAFFBA0BDCEA27FDD28E9E" box="[477,559,619,640]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="25" unit="m" value="2750.0">2,750 m</elevation>
</quantity>
, ex: cloud forest litter,
<date id="0C5534F2FFCAFFBA0B1FEACAFE658E85" box="[286,408,646,667]" pageId="25" value="1991-09-18">
<collectingDate id="1C11CD1AFFCAFFBA0B1FEACAFE658E85" box="[286,408,646,667]" pageId="25" value="1991-09-18">18. IX. 1991</collectingDate>
</date>
,
<collectorName id="D51E77E4FFCAFFBA0BA3EACAFDE58E85" box="[418,536,646,667]" pageId="25">R. Anderson</collectorName>
leg. (
<collectionCode id="1EFA8AF7FFCAFFBA084FEACAFD758E85" box="[590,648,646,667]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="25" type="Museum">CMN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="30F141B9FFCAFFBA0AC4EAECFDF98F1B" pageId="25" type="etymology">
<paragraph id="78541232FFCAFFBA0AC4EAECFDF98F1B" blockId="25.[173,660,406,1729]" pageId="25">
<emphasis id="4A9FCE20FFCAFFBA0AC4EAECFECA8EAB" bold="true" box="[197,311,672,693]" pageId="25">Etymology.</emphasis>
The specific epithet is derived from the Greek word “epakros” meaning “pointed at the end,” a reference to the strongly acuminate apex of the tegmen in ventral view.
</paragraph>
</subSubSection>
<subSubSection id="30F141B9FFCAFFBA0AC4EB46FE4A899D" pageId="25" type="diagnosis">
<paragraph id="78541232FFCAFFBA0AC4EB46FE4F885A" blockId="25.[173,660,406,1729]" pageId="25">
<emphasis id="4A9FCE20FFCAFFBA0AC4EB46FED18F01" bold="true" box="[197,300,778,799]" pageId="25">Diagnosis.</emphasis>
This species differs from other congeners by the following combination of characters: pronotum and elytra different of colors; pronotum pale yellow, with dark brown longitudinal maculation at midline, possessing two vague lateral markings; antennal segment XI with one oval antennal sensilla; pro- and mesotarsal segments II and III of males moderately dilated; apical margin of tegmen strongly acute in ventral view; median lobe subparallel-sided in ventral view; apical margin of median lobe widely rounded; apex of short, thick gonocoxites deeply incised.
</paragraph>
<paragraph id="78541232FFCAFFBA0AC4EC05FE4A899D" blockId="25.[173,660,406,1729]" pageId="25">
<emphasis id="4A9FCE20FFCAFFBA0AC4EC05FE9B8840" bold="true" box="[197,358,1097,1118]" pageId="25">Similar Species.</emphasis>
This new species is similar to
<taxonomicName id="BFEB69B1FFCAFFBA0AACEC2FFED38867" authorityName="Hisamatsu" authorityYear="2016" box="[173,302,1123,1145]" class="Insecta" family="Nitidulidae" genus="Cyclocaccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="4A9FCE20FFCAFFBA0AACEC2FFED38867" box="[173,302,1123,1145]" italics="true" pageId="25">C. maculatus</emphasis>
</taxonomicName>
in having one antennal sensilla and rather short and thick gonocoxites, but can be distinguished by the following combination of characters: abdominal tergite VII and legs fully reddish brown to chestnut brown; apical margin of tegmen strongly acute in ventral view. It is also similar to
<taxonomicName id="BFEB69B1FFCAFFBA0AACED4EFED58906" authorityName="Sharp" authorityYear="1891" box="[173,296,1282,1304]" class="Insecta" family="Nitidulidae" genus="Cyclocaccus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="4A9FCE20FFCAFFBA0AACED4EFF408906" box="[173,189,1282,1304]" italics="true" pageId="25">C</emphasis>
.
<emphasis id="4A9FCE20FFCAFFBA0ACAED4EFED58906" box="[203,296,1282,1304]" italics="true" pageId="25">monticola</emphasis>
</taxonomicName>
, but can be distinguished by the following combination of characters: antennal segment XI with one oval antennal sensilla; apical margin of tegmen strongly acute in ventral view; gonocoxites short and thick.
</paragraph>
</subSubSection>
<subSubSection id="30F141B9FFCAFFBA0AC4EDC4FBF88837" pageId="25" type="description">
<paragraph id="78541232FFCAFFBA0AC4EDC4FBF88837" blockId="25.[173,660,406,1729]" lastBlockId="25.[706,1194,194,1118]" pageId="25">
<emphasis id="4A9FCE20FFCAFFBA0AC4EDC4FEC28983" bold="true" box="[197,319,1416,1437]" pageId="25">Description.</emphasis>
Length 1.632.21 (2.03) mm, width 1.301.72 (1.60) mm, depth
<quantity id="BF13BFD7FFCAFFBA0807EDEFFD6E89A6" box="[518,659,1442,1464]" metricMagnitude="-4" metricUnit="m" metricValue="9.75" metricValueMax="10.2" metricValueMin="9.3" pageId="25" unit="mm" value="0.975" valueMax="1.02" valueMin="0.93">0.931.02 mm</quantity>
(0.97) (
<emphasis id="4A9FCE20FFCAFFBA0AF4EDF1FEFC89CC" box="[245,257,1469,1490]" italics="true" pageId="25">n</emphasis>
= 5).
<emphasis id="4A9FCE20FFCAFFBA0B3CEDF1FE8B89CC" bold="true" box="[317,374,1469,1490]" pageId="25">Male.</emphasis>
Body (
<figureCitation id="E0D00EB7FFCAFFBA0BBEEDF1FDFA89CC" box="[447,519,1469,1490]" captionStart="Fig" captionStartId="7.[197,228,1638,1657]" captionTargetBox="[227,1149,194,1609]" captionTargetId="figure-0@7.[212,1153,194,1614]" captionTargetPageId="7" captionText="Fig. 1. Dorsal habitus of Cyclocaccus species in the C. laeticulus species-group. A) C. clinei, holotype, B) C. laeticulus, holotype, C) C. laeticulus, holotype, labels, D) C. laeticulus, Venezuela, E) C. speciosus, holotype, F) C. monticola, lectotype, G) C. monticola, lectotype, labels, H) C. monticola, Guatemala, I) C. monticola, Bolivia, J) C. costaricensis, holotype, K) C. maculatus, holotype, L) C. epakros, holotype. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340415" httpUri="https://zenodo.org/record/5340415/files/figure.png" pageId="25">Fig. 1L</figureCitation>
) oval; dorsum and venter strongly shining; strongly convex dorsally, moderately convex ventrally; dorsum and venter glabrous. Dorsal disc of head, antennal club (except paler apical area of segment XI), and scutellum dark brown; pronotum pale yellow, with dark brown longitudinal maculation at midline, usually with vague spot on each side of the longitudinal maculation; elytra, abdominal tergites, and venter reddish brown to chestnut brown; hypomera and elytral epipleura pale yellow.
<emphasis id="4A9FCE20FFCAFFBA0E6DE88EFB558CC9" bold="true" box="[1132,1192,194,215]" pageId="25">Head:</emphasis>
Dorsal disc possessing sparse, minute punctures; interspaces reticulate. Labrum feebly notched at middle. Antenna (
<figureCitation id="E0D00EB7FFCAFFBA097AE95EFC318D39" box="[891,972,274,295]" captionStart="Fig" captionStartId="15.[197,228,1686,1705]" captionTargetBox="[191,1130,204,1639]" captionTargetId="figure-1@15.[173,1193,183,1662]" captionTargetPageId="15" captionText="Fig. 9. Cyclocaccus epakros, Guerrero, Mexico. A) Tegmen, ventral view, B) Median lobe, ventral view, C) Right antenna, ventral view, D) Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340432" httpUri="https://zenodo.org/record/5340432/files/figure.png" pageId="25">Fig. 9C</figureCitation>
) with club long and large, ACL/ASI-VIII = 0.82 (
<emphasis id="4A9FCE20FFCAFFBA09DEE960FC168D5F" box="[991,1003,300,321]" italics="true" pageId="25">n</emphasis>
= 1); antennal segment XI with an oval antennal sensilla; approximate ratio of each segment (
<emphasis id="4A9FCE20FFCAFFBA09EAE92EFC0A8D69" box="[1003,1015,354,375]" italics="true" pageId="25">n</emphasis>
= 1) 3.23: 2.24: 2.29: 1.59: 1.72: 1.10: 1.00: 1.13: 4.00: 3.24: 4.44.
<emphasis id="4A9FCE20FFCAFFBA08C2E9DBFCD28DB2" bold="true" box="[707,815,407,428]" pageId="25">Pronotum:</emphasis>
2.923.28 (3.10) times as wide as long (
<emphasis id="4A9FCE20FFCAFFBA08CBE9FDFD2B8DD8" box="[714,726,433,454]" italics="true" pageId="25">n</emphasis>
= 5); disc possessing sparse, minute punctures; interspaces reticulate.
<emphasis id="4A9FCE20FFCAFFBA099BE980FBFE8DFF" bold="true" box="[922,1027,460,481]" pageId="25">Scutellum:</emphasis>
Disc possessing few minute punctures; interspaces reticulate.
<emphasis id="4A9FCE20FFCAFFBA0E62E9AAFB558DE5" bold="true" box="[1123,1192,486,507]" pageId="25">Elytra:</emphasis>
0.971.03 (1.00) times as long as wide (
<emphasis id="4A9FCE20FFCAFFBA0E5EEA4DFB968E08" box="[1119,1131,513,534]" italics="true" pageId="25">n</emphasis>
= 5), 3.454.02 (3.77) times as long as pronotum (
<emphasis id="4A9FCE20FFCAFFBA0E69EA50FB898E2F" box="[1128,1140,540,561]" italics="true" pageId="25">n</emphasis>
= 5), widest at middle; each elytron with 9 regular rows of punctures about as large as eye facet; interstices without punctures or with few minute punctures.
<emphasis id="4A9FCE20FFCAFFBA08C3EACAFCF18E85" bold="true" box="[706,780,646,667]" pageId="25">Venter:</emphasis>
Prosternum feebly convex along midline. Metaventrite strongly convex. Abdominal sternite III without subcoxal lines.
<emphasis id="4A9FCE20FFCAFFBA09C4EAF7FC078ECE" bold="true" box="[965,1018,699,720]" pageId="25">Legs:</emphasis>
Slender; pro- and mesotarsal segments II and III moderately dilated; tarsal claws simple.
<emphasis id="4A9FCE20FFCAFFBA098BEABCFC108F1B" bold="true" box="[906,1005,752,773]" pageId="25">Genitalia:</emphasis>
Tegmen (
<figureCitation id="E0D00EB7FFCAFFBA0E54EABCFB5F8F1B" box="[1109,1186,752,773]" captionStart="Fig" captionStartId="15.[197,228,1686,1705]" captionTargetBox="[191,1130,204,1639]" captionTargetId="figure-1@15.[173,1193,183,1662]" captionTargetPageId="15" captionText="Fig. 9. Cyclocaccus epakros, Guerrero, Mexico. A) Tegmen, ventral view, B) Median lobe, ventral view, C) Right antenna, ventral view, D) Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340432" httpUri="https://zenodo.org/record/5340432/files/figure.png" pageId="25">Fig. 9A</figureCitation>
) long, TL/TW = 2.18 (
<emphasis id="4A9FCE20FFCAFFBA09A1EB47FC518F3E" box="[928,940,779,800]" italics="true" pageId="25">n</emphasis>
= 1), constricted at midlength, then strongly convergent apically; apical and lateral margins of tegmen strongly undulating in ventral view; apical margin of tegmen strongly acute in ventral view; median lobe (
<figureCitation id="E0D00EB7FFCAFFBA0E21EB39FB978F94" box="[1056,1130,885,906]" captionStart="Fig" captionStartId="15.[197,228,1686,1705]" captionTargetBox="[191,1130,204,1639]" captionTargetId="figure-1@15.[173,1193,183,1662]" captionTargetPageId="15" captionText="Fig. 9. Cyclocaccus epakros, Guerrero, Mexico. A) Tegmen, ventral view, B) Median lobe, ventral view, C) Right antenna, ventral view, D) Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340432" httpUri="https://zenodo.org/record/5340432/files/figure.png" pageId="25">Fig. 9B</figureCitation>
) long, in ventral view subparallel-sided; apical margin of median lobe widely rounded; sclerites of internal sac as in
<figureCitation id="E0D00EB7FFCAFFBA092AEB88FC878FC7" box="[811,890,964,985]" captionStart="Fig" captionStartId="15.[197,228,1686,1705]" captionTargetBox="[191,1130,204,1639]" captionTargetId="figure-1@15.[173,1193,183,1662]" captionTargetPageId="15" captionText="Fig. 9. Cyclocaccus epakros, Guerrero, Mexico. A) Tegmen, ventral view, B) Median lobe, ventral view, C) Right antenna, ventral view, D) Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340432" httpUri="https://zenodo.org/record/5340432/files/figure.png" pageId="25">Fig. 9B</figureCitation>
.
<emphasis id="4A9FCE20FFCAFFBA0988EB88FC2A8FC7" bold="true" box="[905,983,964,985]" pageId="25">Female.</emphasis>
Pro- and mesotarsal segments IIII more weakly dilated than those of males. Ovipositor (
<figureCitation id="E0D00EB7FFCAFFBA0983EBB6FC2F8811" box="[898,978,1018,1039]" captionStart="Fig" captionStartId="15.[197,228,1686,1705]" captionTargetBox="[191,1130,204,1639]" captionTargetId="figure-1@15.[173,1193,183,1662]" captionTargetPageId="15" captionText="Fig. 9. Cyclocaccus epakros, Guerrero, Mexico. A) Tegmen, ventral view, B) Median lobe, ventral view, C) Right antenna, ventral view, D) Ovipositor. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340432" httpUri="https://zenodo.org/record/5340432/files/figure.png" pageId="25">Fig. 9D</figureCitation>
) with deeply incised apices of short, thick gonocoxites.
</paragraph>
</subSubSection>
<subSubSection id="30F141B9FFCAFFBA08DBEC63FCEA8840" pageId="25" type="distribution">
<paragraph id="78541232FFCAFFBA08DBEC63FCEA8840" blockId="25.[706,1194,194,1118]" pageId="25">
<emphasis id="4A9FCE20FFCAFFBA08DBEC63FCA1885A" bold="true" box="[730,860,1071,1092]" pageId="25">Distribution.</emphasis>
<collectingCountry id="00FC52A2FFCAFFBA0969EC63FC4D885A" box="[872,944,1071,1092]" name="Mexico" pageId="25">Mexico</collectingCountry>
(
<collectingRegion id="BA2FDCD0FFCAFFBA09C2EC63FBE7885A" box="[963,1050,1071,1092]" country="Mexico" name="Guerrero" pageId="25">Guerrero</collectingRegion>
and
<collectingRegion id="BA2FDCD0FFCAFFBA0E55EC63FB5F885A" box="[1108,1186,1071,1092]" country="Mexico" name="Chiapas" pageId="25">Chiapas</collectingRegion>
) (Map 3).
</paragraph>
</subSubSection>
</treatment>
</document>