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<document id="AD3804ABD23627F559C03A84C2BCDD14" ID-DOI="10.11646/zootaxa.3973.1.4" ID-GBIF-Dataset="141343cb-d3a1-4d85-8075-6268d66c6f10" ID-ISSN="1175-5326" ID-Zenodo-Dep="231922" ID-ZooBank="688DC9B0-2EA4-43D5-926A-B184A0A9A7B2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461202010501" checkinUser="plazi" docAuthor="Cumberlidge, Neil" docDate="2015" docId="F02E87F81077FFBFFF63BFC9DFE00AE0" docLanguage="en" docName="zt03973p138.pdf" docOrigin="Zootaxa 3973 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Potamonautes langi Rathbun 1921" docType="treatment" docVersion="7" lastPageNumber="126" masterDocId="0C17FF801076FFB8FFF4BC22DC5E082C" masterDocTitle="Redescriptions of three species of freshwater crabs from the Democratic Republic of Congo, Central Africa (Brachyura: Potamoidea: Potamonautidae)" masterLastPageNumber="138" masterPageNumber="119" pageNumber="120" updateTime="1698600957567" updateUser="plazi">
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<mods:title id="0403A5F49B9DB2F94DAAD4B106DFD46F">Redescriptions of three species of freshwater crabs from the Democratic Republic of Congo, Central Africa (Brachyura: Potamoidea: Potamonautidae)</mods:title>
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<mods:namePart id="A65ECF606317E1013A26B80C0E0AE9CF">Cumberlidge, Neil</mods:namePart>
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<mods:date id="FA10F11FF172370A20636F02162E4116">2015</mods:date>
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<treatment id="F02E87F81077FFBFFF63BFC9DFE00AE0" ID-DOI="http://doi.org/10.5281/zenodo.6096908" ID-GBIF-Taxon="119629580" ID-Zenodo-Dep="6096908" LSID="urn:lsid:plazi:treatment:F02E87F81077FFBFFF63BFC9DFE00AE0" httpUri="http://treatment.plazi.org/id/F02E87F81077FFBFFF63BFC9DFE00AE0" lastPageId="7" lastPageNumber="126" pageId="1" pageNumber="120">
<subSubSection id="309D65651077FFB9FF63BFC9DD4C0C67" pageId="1" pageNumber="120" type="nomenclature">
<paragraph id="783836EE1077FFB9FF63BFC9DE050C2A" blockId="1.[151,603,1003,1099]" box="[151,603,1003,1030]" pageId="1" pageNumber="120">
<heading id="237081821077FFB9FF63BFC9DE050C2A" bold="true" box="[151,603,1003,1030]" fontSize="11" level="1" pageId="1" pageNumber="120" reason="1">
<taxonomicName id="BF874D6D1077FFB9FF63BFC9DE050C2A" ID-CoL="77VN4" authority="Rathbun, 1921" authorityName="Rathbun" authorityYear="1921" box="[151,603,1003,1030]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="120" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1077FFB9FF63BFC9DE050C2A" bold="true" box="[151,603,1003,1030]" pageId="1" pageNumber="120">
<emphasis id="4AF3EAFC1077FFB9FF63BFC9DDD60C29" bold="true" box="[151,392,1003,1029]" italics="true" pageId="1" pageNumber="120">Potamonautes langi</emphasis>
(
<bibRefCitation id="1C164B1F1077FFB9FE63BFCEDE0D0C2A" author="Rathbun" box="[407,595,1004,1030]" pageId="1" pageNumber="120" refString="Rathbun, M. J. (1921) Brachyuran Crabs of the Belgian Congo. Bulletin of the American Museum of Natural History, 43, 379 - 468." type="journal article" year="1921">Rathbun, 1921</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="783836EE1077FFB9FF63B82DDDFD0C0B" blockId="1.[151,603,1003,1099]" box="[151,419,1038,1063]" pageId="1" pageNumber="120">Langs Freshwater Crab</paragraph>
<paragraph id="783836EE1077FFB9FF63B810DD4C0C67" blockId="1.[151,603,1003,1099]" box="[151,274,1074,1099]" pageId="1" pageNumber="120">
(
<figureCitation id="E0BC2A6B1077FFB9FF6BB810DD570C67" box="[159,265,1074,1099]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1859,1881]" captionStartId-1="3.[151,250,1693,1715]" captionStartId-2="4.[151,250,1513,1535]" captionStartId-3="5.[151,250,1923,1945]" captionTargetBox-0="[227,1315,312,1773]" captionTargetBox-1="[281,1315,225,1619]" captionTargetBox-2="[386,1200,193,1490]" captionTargetBox-3="[227,1340,892,1859]" captionTargetId-0="figure@2.[208,1329,301,1778]" captionTargetId-1="figure@3.[279,1331,215,1627]" captionTargetId-2="figure@4.[386,1200,193,1492]" captionTargetId-3="figure@5.[207,1341,877,1880]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." captionText-1="FIGURE 2. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, mandible frontal view; B, mandibular palp superior view; C, third maxilliped frontal view; D, G 1 ventral view; E, G 1 dorsal view; F, superior view of terminal article; G, G 2 ventral view. Scale bar: 2.5 mm (A), 1.5 mm (B), 12.5 mm C, 4.5 mm (D G)." captionText-2="FIGURE 3. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). Whole animal A, frontal view; B, dorsal view; C, ventral view. Scale bar: 21.4 mm (A C)." captionText-3="FIGURE 4. Map of the D. R. Congo in Central Africa showing the known localities of Potamonautes langi (black circles, not shown to scale). See text for exact locality coordinates." httpUri-0="https://zenodo.org/record/231923/files/figure.png" httpUri-1="https://zenodo.org/record/231924/files/figure.png" httpUri-2="https://zenodo.org/record/231925/files/figure.png" httpUri-3="https://zenodo.org/record/231926/files/figure.png" pageId="1" pageNumber="120">Figs. 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="309D65651077FFB9FF63B859D95B0D27" pageId="1" pageNumber="120" type="reference_group">
<paragraph id="783836EE1077FFB9FF63B859DF2F0CBC" blockId="1.[151,1285,1146,1291]" box="[151,881,1146,1168]" pageId="1" pageNumber="120">
<treatmentCitationGroup id="589711C01077FFB9FF63B859DF2F0CBC" box="[151,881,1146,1168]" pageId="1" pageNumber="120">
<taxonomicName id="BF874D6D1077FFB9FF63B859DEFA0CBC" authority="Rathbun, 1921: 430" authorityName="Rathbun" authorityPageNumber="430" authorityYear="1921" box="[151,676,1146,1168]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="120" phylum="Arthropoda" rank="species" species="langi" subGenus="Potamonautes">
<emphasis id="4AF3EAFC1077FFB9FF63B859DCAD0CBC" box="[151,243,1147,1168]" italics="true" pageId="1" pageNumber="120">Potamon</emphasis>
(
<emphasis id="4AF3EAFC1077FFB9FEF5B859DDCC0CBC" box="[257,402,1147,1168]" italics="true" pageId="1" pageNumber="120">Potamonautes</emphasis>
)
<emphasis id="4AF3EAFC1077FFB9FE54B858DD8A0CBC" box="[416,468,1146,1168]" italics="true" pageId="1" pageNumber="120">langi</emphasis>
<treatmentCitation id="F92610FF1077FFB9FE2FB858DEFA0CBC" author="Rathbun" box="[475,676,1146,1168]" page="430" pageId="1" pageNumber="120" year="1921">
<bibRefCitation id="1C164B1F1077FFB9FE2FB858DE2D0CBC" author="Rathbun" box="[475,627,1146,1168]" pageId="1" pageNumber="120" refString="Rathbun, M. J. (1921) Brachyuran Crabs of the Belgian Congo. Bulletin of the American Museum of Natural History, 43, 379 - 468." type="journal article" year="1921">Rathbun, 1921</bibRefCitation>
: 430
</treatmentCitation>
</taxonomicName>
433, pl. 32, fig. 14.
</treatmentCitationGroup>
</paragraph>
<paragraph id="783836EE1077FFB9FF63B8B8DEFA0C83" blockId="1.[151,1285,1146,1291]" box="[151,676,1177,1199]" pageId="1" pageNumber="120">
<treatmentCitationGroup id="589711C01077FFB9FF63B8B8DEFA0C83" box="[151,676,1177,1199]" pageId="1" pageNumber="120">
<emphasis id="4AF3EAFC1077FFB9FF63B8B8DD260C83" box="[151,376,1177,1199]" italics="true" pageId="1" pageNumber="120">
<taxonomicName id="BF874D6D1077FFB9FF63B8B8DD760C83" ID-CoL="7PVHW" box="[151,296,1178,1199]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="120" phylum="Arthropoda" rank="genus">Potamonautes</taxonomicName>
langi—
</emphasis>
<treatmentCitation id="F92610FF1077FFB9FE8CB8BBDE400C83" author="Balss" box="[376,542,1177,1199]" page="189" pageId="1" pageNumber="120" year="1936">
<bibRefCitation id="1C164B1F1077FFB9FE8CB8BBDDB20C83" author="Balss" box="[376,492,1177,1199]" pageId="1" pageNumber="120" refString="Balss, H. (1936) Beitrage zur Kenntnis der Potamidae (Suβwasserkrabben) des Kongogebeites. Revue Zoologique et Botanique d ' Afrique, 2 (2), 165 - 204." type="journal article" year="1936">Balss 1936</bibRefCitation>
: 189
</treatmentCitation>
, 190, fig. 25.
</treatmentCitationGroup>
</paragraph>
<paragraph id="783836EE1077FFB9FF63B89BDDAB0CE2" blockId="1.[151,1285,1146,1291]" box="[151,501,1208,1230]" pageId="1" pageNumber="120">
<treatmentCitationGroup id="589711C01077FFB9FF63B89BDDAB0CE2" box="[151,501,1208,1230]" pageId="1" pageNumber="120">
<emphasis id="4AF3EAFC1077FFB9FF63B89BDD1D0CE2" box="[151,323,1208,1230]" italics="true" pageId="1" pageNumber="120">
<taxonomicName id="BF874D6D1077FFB9FF63B89BDCAD0CE2" ID-CoL="7PTXN" box="[151,243,1209,1230]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="120" phylum="Arthropoda" rank="genus">Potamon</taxonomicName>
langi—
</emphasis>
<treatmentCitation id="F92610FF1077FFB9FEB7B89ADDAF0CE2" author="Chace" box="[323,497,1208,1230]" page="215" pageId="1" pageNumber="120" year="1942">
<bibRefCitation id="1C164B1F1077FFB9FEB7B89ADDE00CE2" author="Chace" box="[323,446,1208,1230]" pageId="1" pageNumber="120" refString="Chace, F. A. (1942) Scientific results of a fourth expedition to forested areas in eastern Africa, III: Decapod Crustacea. Bulletin of the Museum of Comparative Zoology at Harvard College, 91 (3), 185 - 233." type="journal article" year="1942">Chace 1942</bibRefCitation>
: 215
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="783836EE1077FFB9FF63B8FAD95B0D27" blockId="1.[151,1285,1146,1291]" pageId="1" pageNumber="120">
<taxonomicName id="BF874D6D1077FFB9FF63B8FADE530CC0" ID-CoL="7PVHW" authority="(Obesopotamonautes)" baseAuthorityName="Obesopotamonautes" box="[151,525,1238,1261]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="4AF3EAFC1077FFB9FF63B8FADD760CC1" box="[151,296,1240,1261]" italics="true" pageId="1" pageNumber="120">Potamonautes</emphasis>
(
<emphasis id="4AF3EAFC1077FFB9FEC2B8F4DE5B0CC0" box="[310,517,1238,1260]" italics="true" pageId="1" pageNumber="120">Obesopotamonautes</emphasis>
)
</taxonomicName>
<emphasis id="4AF3EAFC1077FFB9FDE0B8F4DE010CC0" box="[532,607,1238,1260]" italics="true" pageId="1" pageNumber="120">langi—</emphasis>
<bibRefCitation id="1C164B1F1077FFB9FDAAB8F5DE980CC0" author="Bott" box="[606,710,1238,1260]" pageId="1" pageNumber="120" refString="Bott, R. (1955) Die Suβwasserkrabben von Afrika und ihre Stammesgeschichte. Annales du Musee Royal du Congo Belge, 1 (3), 209 - 352." type="journal article" year="1955">Bott 1955</bibRefCitation>
: 256257, figs. 17, 18, 79a, b, pl. 12, figs. lad.
<taxonomicName id="BF874D6D1077FFB9FF63B8D4DD3C0D27" ID-CoL="77VN4" box="[151,354,1269,1291]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="120" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1077FFB9FF63B8D4DD3C0D27" box="[151,354,1269,1291]" italics="true" pageId="1" pageNumber="120">Potamonautes langi</emphasis>
</taxonomicName>
<bibRefCitation id="1C164B1F1077FFB9FE9BB8D7DE710D27" author="Cumberlidge" box="[367,559,1269,1291]" pageId="1" pageNumber="120" refString="Cumberlidge, N. (1998) The African and Madagascan freshwater crabs in the Zoologische Staatssammlung, Munich (Crustacea: Decapoda: Brachyura: Potamoidea), Spixiana, 21 (3), 193 - 214." type="journal article" year="1998">Cumberlidge 1998</bibRefCitation>
: 200; Ng
<emphasis id="4AF3EAFC1077FFB9FD66B8D4DE9B0D27" box="[658,709,1269,1291]" italics="true" pageId="1" pageNumber="120">et al.</emphasis>
2008: 171; Cumberlidge
<emphasis id="4AF3EAFC1077FFB9FC3CB8D4DFA20D27" box="[968,1020,1269,1291]" italics="true" pageId="1" pageNumber="120">et al.</emphasis>
2009: appendix item 942.
</paragraph>
</subSubSection>
<subSubSection id="309D65651077FFBAFF63B915D90308D4" lastPageId="2" lastPageNumber="121" pageId="1" pageNumber="120" type="materials_examined">
<paragraph id="783836EE1077FFBAFF63B915D90308D4" blockId="1.[151,1437,1335,2009]" lastBlockId="2.[151,1436,151,248]" lastPageId="2" lastPageNumber="121" pageId="1" pageNumber="120">
<emphasis id="4AF3EAFC1077FFB9FF63B915DDE00D7C" bold="true" box="[151,446,1335,1360]" pageId="1" pageNumber="120">
<typeStatus id="A73C884C1077FFB9FF63B915DC8C0D7C" box="[151,210,1335,1360]" pageId="1" pageNumber="120">Type</typeStatus>
material examined.
</emphasis>
D.R.
<collectingCountry id="0090767E1077FFB9FDF4B91ADE390D7C" box="[512,615,1336,1360]" name="Republic of the Congo" pageId="1" pageNumber="120">CONGO</collectingCountry>
.
<collectingCountry id="0090767E1077FFB9FD85B91ADEE20D7C" box="[625,700,1336,1360]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani (
<geoCoordinate id="1DB350291077FFB9FC56B915DFA00D63" box="[930,1022,1335,1360]" direction="north" orientation="latitude" pageId="1" pageNumber="120" precision="555" value="0.52">0.52° N</geoCoordinate>
,
<geoCoordinate id="1DB350291077FFB9FBFDB915D8330D7C" box="[1033,1133,1335,1360]" direction="east" orientation="longitude" pageId="1" pageNumber="120" precision="555" value="25.2">25.20° E</geoCoordinate>
), adult male
<typeStatus id="A73C884C1077FFB9FB09B915D9010D7C" box="[1277,1375,1335,1360]" pageId="1" pageNumber="120" type="holotype">holotype</typeStatus>
(CW 45, CL 34.5,
<collectingCountry id="0090767E1077FFB9FED9B97EDD0A0D58" box="[301,340,1372,1396]" name="Switzerland" pageId="1" pageNumber="120">CH</collectingCountry>
16, FW 15),
<date id="0C39102E1077FFB9FE19B97EDEDF0D59" box="[493,641,1372,1397]" pageId="1" pageNumber="120" value="1915-04-21">21 Apr. 1915</date>
(J. Chapin and H. Lang) (
<collectionCode id="1E96AE2B1077FFB9FC5EB97ED85C0D58" LSID="urn:lsid:biocol.org:col:34925" box="[938,1026,1372,1396]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="120">AMNH</collectionCode>
3353).
<collectingCountry id="0090767E1077FFB9FBADB97ED8FA0D58" box="[1113,1188,1372,1396]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FA7AB97EDC8E0DB4" pageId="1" pageNumber="120" type="male">1 male</specimenCount>
,
<specimenCount id="6E81FD671077FFB9FF14B9A2DD010DB4" box="[224,351,1407,1432]" pageId="1" pageNumber="120" type="female">10 females</specimenCount>
(4 ovigerous)
<typeStatus id="A73C884C1077FFB9FDE5B9A2DE220DB4" box="[529,636,1408,1432]" pageId="1" pageNumber="120" type="paratype">paratypes</typeStatus>
(
<collectionCode id="1E96AE2B1077FFB9FD64B9A2DEB90DB4" LSID="urn:lsid:biocol.org:col:34925" box="[656,743,1408,1432]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="120">AMNH</collectionCode>
3336);
<specimenCount id="6E81FD671077FFB9FCB3B9A2DFE60DB4" box="[839,952,1407,1432]" pageId="1" pageNumber="120" type="female">3 females</specimenCount>
(2 ovigerous),
<typeStatus id="A73C884C1077FFB9FB85B9A2D8820DB4" box="[1137,1244,1408,1432]" pageId="1" pageNumber="120" type="paratype">paratypes</typeStatus>
(
<collectionCode id="1E96AE2B1077FFB9FB04B9A2D9190DB4" LSID="urn:lsid:biocol.org:col:34925" box="[1264,1351,1408,1432]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="120">AMNH</collectionCode>
3469);
<collectingCountry id="0090767E1077FFB9FF63B986DCBC0D90" box="[151,226,1444,1468]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FE3DB986DE7C0D91" box="[457,546,1444,1469]" pageId="1" pageNumber="120" type="male">3 males</specimenCount>
,
<specimenCount id="6E81FD671077FFB9FDD9B986DEC70D91" box="[557,665,1444,1469]" pageId="1" pageNumber="120" type="female">8 females</specimenCount>
(5 ovigerous),
<specimenCount id="6E81FD671077FFB9FCB1B986DFEA0D91" box="[837,948,1444,1469]" pageId="1" pageNumber="120" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="1E96AE2B1077FFB9FC37B986D8450D90" LSID="urn:lsid:biocol.org:col:34925" box="[963,1051,1444,1468]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="120">AMNH</collectionCode>
3363);
<collectingCountry id="0090767E1077FFB9FB85B986D8E20D90" box="[1137,1212,1444,1468]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FF63B9EADCB60DCC" box="[151,232,1479,1504]" pageId="1" pageNumber="120" type="male">1 male</specimenCount>
,
<specimenCount id="6E81FD671077FFB9FF02B9EADD380DCC" box="[246,358,1479,1504]" pageId="1" pageNumber="120" type="female">7 females</specimenCount>
(4 ovigerous) (
<specimenCount id="6E81FD671077FFB9FDE8B9E5DE270DCC" box="[540,633,1479,1504]" pageId="1" pageNumber="120" type="male">5 males</specimenCount>
,
<specimenCount id="6E81FD671077FFB9FD73B9EADF5A0DCC" box="[647,772,1479,1504]" pageId="1" pageNumber="120" type="female">10 females</specimenCount>
(2 ovigerous) sent to
<collectionCode id="1E96AE2B1077FFB9FBFDB9EAD8000DCC" LSID="urn:lsid:biocol.org:col:34871" box="[1033,1118,1480,1504]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="120">USNM</collectionCode>
),
<typeStatus id="A73C884C1077FFB9FB81B9EAD8BE0DCC" box="[1141,1248,1480,1504]" pageId="1" pageNumber="120" type="paratype">paratypes</typeStatus>
(
<collectionCode id="1E96AE2B1077FFB9FB06B9EAD9170DCC" LSID="urn:lsid:biocol.org:col:34925" box="[1266,1353,1480,1504]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="120">AMNH</collectionCode>
3332);
<collectingCountry id="0090767E1077FFB9FF63B9CEDCBC0E28" box="[151,226,1516,1540]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FE3CB9CEDE7F0E29" box="[456,545,1516,1541]" pageId="1" pageNumber="120" type="male">2 males</specimenCount>
,
<specimenCount id="6E81FD671077FFB9FDD9B9CEDEF80E29" box="[557,678,1516,1541]" pageId="1" pageNumber="120" type="female">11 females</specimenCount>
(5 ovigerous)
<typeStatus id="A73C884C1077FFB9FCBDB9CFDFEA0E29" box="[841,948,1517,1541]" pageId="1" pageNumber="120" type="paratype">paratypes</typeStatus>
(
<collectionCode id="1E96AE2B1077FFB9FC37B9CED8450E28" LSID="urn:lsid:biocol.org:col:34925" box="[963,1051,1516,1540]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="120">AMNH</collectionCode>
3362);
<collectingCountry id="0090767E1077FFB9FB85B9CED8E20E28" box="[1137,1212,1516,1540]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FF63BA32DCAC0E04" box="[151,242,1551,1576]" pageId="1" pageNumber="120" type="male">3 males</specimenCount>
,
<specimenCount id="6E81FD671077FFB9FEF4BA2DDD220E04" box="[256,380,1551,1576]" pageId="1" pageNumber="120" type="female">16 females</specimenCount>
(8 ovigerous),
<specimenCount id="6E81FD671077FFB9FDDABA32DEF30E04" box="[558,685,1551,1576]" pageId="1" pageNumber="120" type="juvenile">3 juveniles</specimenCount>
,
<typeStatus id="A73C884C1077FFB9FD4FBA32DF780E04" box="[699,806,1552,1576]" pageId="1" pageNumber="120" type="paratype">paratypes</typeStatus>
(
<collectionCode id="1E96AE2B1077FFB9FCC3BA32DFD10E04" LSID="urn:lsid:biocol.org:col:34925" box="[823,911,1552,1576]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="120">AMNH</collectionCode>
3343);
<collectingCountry id="0090767E1077FFB9FC1EBA32D86B0E04" box="[1002,1077,1552,1576]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FAD1BA32D9DE0E04" box="[1317,1408,1551,1576]" pageId="1" pageNumber="120" type="male">3 males</specimenCount>
,
<specimenCount id="6E81FD671077FFB9FA7ABA2DDCB00E61" pageId="1" pageNumber="120" type="female">5 females</specimenCount>
(3 ovigerous),
<typeStatus id="A73C884C1077FFB9FE6DBA17DE560E61" box="[409,520,1589,1613]" pageId="1" pageNumber="120" type="paratype">paratypes</typeStatus>
;
<collectingCountry id="0090767E1077FFB9FDE0BA16DE010E60" box="[532,607,1588,1612]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FCB2BA16DFEC0E61" box="[838,946,1588,1613]" pageId="1" pageNumber="120" type="female">2 females</specimenCount>
(both ovigerous)
<typeStatus id="A73C884C1077FFB9FB8EBA17D8B80E61" box="[1146,1254,1589,1613]" pageId="1" pageNumber="120" type="paratype">paratypes</typeStatus>
(
<collectionCode id="1E96AE2B1077FFB9FB00BA16D9150E60" LSID="urn:lsid:biocol.org:col:34925" box="[1268,1355,1588,1612]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="120">AMNH</collectionCode>
3418);
<specimenCount id="6E81FD671077FFB9FF63BA7ADD5D0E5C" box="[151,259,1623,1648]" pageId="1" pageNumber="120" type="female">2 females</specimenCount>
(both ovigerous)
<typeStatus id="A73C884C1077FFB9FE38BA7ADE690E5C" box="[460,567,1624,1648]" pageId="1" pageNumber="120" type="paratype">paratypes</typeStatus>
(
<collectionCode id="1E96AE2B1077FFB9FDB1BA7ADEC30E5C" LSID="urn:lsid:biocol.org:col:34925" box="[581,669,1624,1648]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="120">AMNH</collectionCode>
3393);
<collectingCountry id="0090767E1077FFB9FD00BA7ADF610E5C" box="[756,831,1624,1648]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FBD3BA7AD8D50E5C" box="[1063,1163,1623,1648]" pageId="1" pageNumber="120" type="female">1 female</specimenCount>
,
<typeStatus id="A73C884C1077FFB9FB62BA7AD8A70E5C" box="[1174,1273,1624,1648]" pageId="1" pageNumber="120" type="paratype">paratype</typeStatus>
, caught on the shore; coll.
<date id="0C39102E1077FFB9FED5BA5EDD9B0EB8" box="[289,453,1660,1684]" pageId="1" pageNumber="120" value="1909-08-14">14 Aug. 1909</date>
;
<collectingCountry id="0090767E1077FFB9FE20BA5EDE410EB8" box="[468,543,1660,1684]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FCE0BA5EDF250EB9" box="[788,891,1660,1685]" pageId="1" pageNumber="120" type="female">1 female</specimenCount>
,
<typeStatus id="A73C884C1077FFB9FC7EBA5FDFB00EB9" box="[906,1006,1661,1685]" pageId="1" pageNumber="120" type="paratype">paratype</typeStatus>
, coll.
<date id="0C39102E1077FFB9FBC3BA5ED8820EB8" box="[1079,1244,1660,1685]" pageId="1" pageNumber="120" value="1909-08-15">15 Aug. 1909</date>
;
<collectingCountry id="0090767E1077FFB9FB1EBA5ED96B0EB8" box="[1258,1333,1660,1684]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FEE2BA82DDDB0E94" box="[278,389,1695,1720]" pageId="1" pageNumber="120" type="female">4 females</specimenCount>
,
<typeStatus id="A73C884C1077FFB9FE64BA82DDA00E94" box="[400,510,1696,1720]" pageId="1" pageNumber="120" type="paratype">paratypes</typeStatus>
, common under logs in the water, coll.
<date id="0C39102E1077FFB9FC42BA82D80D0E94" box="[950,1107,1696,1720]" pageId="1" pageNumber="120" value="1909-08-17">17 Aug. 1909</date>
;
<collectingCountry id="0090767E1077FFB9FBA9BA82D8F60E94" box="[1117,1192,1696,1720]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FA7ABA82DCB80EF1" pageId="1" pageNumber="120" type="female">1 female</specimenCount>
,
<typeStatus id="A73C884C1077FFB9FF00BAE7DD3C0EF1" box="[244,354,1733,1757]" pageId="1" pageNumber="120" type="paratype">paratypes</typeStatus>
, coll.
<date id="0C39102E1077FFB9FE5EBAE6DE120EF0" box="[426,588,1732,1756]" pageId="1" pageNumber="120" value="1909-08-18">18 Aug. 1909</date>
;
<collectingCountry id="0090767E1077FFB9FDAEBAE6DEFB0EF0" box="[602,677,1732,1756]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FC63BAE6DFB60EF1" box="[919,1000,1732,1757]" pageId="1" pageNumber="120" type="male">1 male</specimenCount>
,
<typeStatus id="A73C884C1077FFB9FC02BAE7D8070EF1" box="[1014,1113,1733,1757]" pageId="1" pageNumber="120" type="paratype">paratype</typeStatus>
, coll.
<date id="0C39102E1077FFB9FB56BAE6D91A0EF0" box="[1186,1348,1732,1756]" pageId="1" pageNumber="120" value="1909-08-20">20 Aug. 1909</date>
;
<collectingCountry id="0090767E1077FFB9FAA6BAE6D9C30EF0" box="[1362,1437,1732,1756]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FE72BACADDB10F2C" box="[390,495,1767,1792]" pageId="1" pageNumber="120" type="female">1 female</specimenCount>
,
<typeStatus id="A73C884C1077FFB9FE0BBACADE3C0F2C" box="[511,610,1768,1792]" pageId="1" pageNumber="120" type="paratype">paratype</typeStatus>
, ovigerous, coll.
<date id="0C39102E1077FFB9FCC5BACADF860F2C" box="[817,984,1768,1792]" pageId="1" pageNumber="120" value="1909-08-21">21 Aug. 1909</date>
;
<collectingCountry id="0090767E1077FFB9FC1DBACAD86A0F2C" box="[1001,1076,1768,1792]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FADBBACAD9C60F2C" box="[1327,1432,1767,1792]" pageId="1" pageNumber="120" type="female">1 female</specimenCount>
,
<typeStatus id="A73C884C1077FFB9FF63BB2FDCA40F09" box="[151,250,1805,1829]" pageId="1" pageNumber="120" type="paratype">paratype</typeStatus>
, coll.
<date id="0C39102E1077FFB9FECABB2EDD820F08" box="[318,476,1804,1828]" pageId="1" pageNumber="120" value="1909-08-28">28 Aug. 1909</date>
;
<collectingCountry id="0090767E1077FFB9FE13BB2EDE6C0F08" box="[487,562,1804,1828]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FCEFBB2EDFD40F09" box="[795,906,1804,1829]" pageId="1" pageNumber="120" type="female">2 females</specimenCount>
,
<typeStatus id="A73C884C1077FFB9FC62BB2FD85A0F09" box="[918,1028,1805,1829]" pageId="1" pageNumber="120" type="paratype">paratypes</typeStatus>
,
<specimenCount id="6E81FD671077FFB9FBE4BB2ED82A0F09" box="[1040,1140,1804,1829]" pageId="1" pageNumber="120" type="female">1 female</specimenCount>
,
<typeStatus id="A73C884C1077FFB9FB74BB2FD8BD0F09" box="[1152,1251,1805,1829]" pageId="1" pageNumber="120" type="paratype">paratype</typeStatus>
, ovigerous, coll.
<date id="0C39102E1077FFB9FF63BB0DDD550F64" box="[151,267,1839,1864]" pageId="1" pageNumber="120" value="1915-02">Feb. 1915</date>
;
<collectingCountry id="0090767E1077FFB9FEECBB12DD3D0F64" box="[280,355,1840,1864]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani,
<specimenCount id="6E81FD671077FFB9FDA4BB12DEFE0F64" box="[592,672,1839,1864]" pageId="1" pageNumber="120" type="male">1 male</specimenCount>
,
<specimenCount id="6E81FD671077FFB9FD58BB12DF760F64" box="[684,808,1839,1864]" pageId="1" pageNumber="120" type="female">13 females</specimenCount>
(8 ovigerous),
<typeStatus id="A73C884C1077FFB9FC23BB12D81B0F64" box="[983,1093,1840,1864]" pageId="1" pageNumber="120" type="paratype">paratypes</typeStatus>
, coll.
<date id="0C39102E1077FFB9FB78BB12D95E0F64" box="[1164,1280,1839,1864]" pageId="1" pageNumber="120" value="1915-04">Apr. 1915</date>
; Kisangani, 4
<typeStatus id="A73C884C1077FFB9FF63BB77DD5B0F41" box="[151,261,1877,1901]" pageId="1" pageNumber="120" type="paratype">paratypes</typeStatus>
, (
<date id="0C39102E1077FFB9FEEBBB76DD820F41" box="[287,476,1876,1901]" pageId="1" pageNumber="120" value="1915-04-01" valueMax="1915-04-30" valueMin="1915-04-01">130 Apr. 1915</date>
) (J. Chapin and H. Lang) (
<collectionCode id="1E96AE2B1077FFB9FCDABB76DF330F40" LSID="urn:lsid:biocol.org:col:33791" box="[814,877,1876,1900]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="1" pageNumber="120">MCZ</collectionCode>
<collectionCode id="1E96AE2B1077FFB9FC8EBB76DF9E0F41" box="[890,960,1876,1901]" pageId="1" pageNumber="120">CRU-</collectionCode>
10614).
<emphasis id="4AF3EAFC1077FFB9FBD5BB76D8BC0F41" bold="true" box="[1057,1250,1876,1901]" pageId="1" pageNumber="120">Other material.</emphasis>
D.R.
<collectingCountry id="0090767E1077FFB9FAC5BB76D9C60F40" box="[1329,1432,1876,1900]" name="Republic of the Congo" pageId="1" pageNumber="120">CONGO</collectingCountry>
. Bukama, Katanga (
<geoCoordinate id="1DB350291077FFB9FE85BB55DD960FBC" box="[369,456,1911,1936]" direction="south" orientation="latitude" pageId="1" pageNumber="120" precision="555" value="-9.15">9.15° S</geoCoordinate>
,
<geoCoordinate id="1DB350291077FFB9FE22BB55DE630FBC" box="[470,573,1911,1936]" direction="east" orientation="longitude" pageId="1" pageNumber="120" precision="555" value="25.83">25.83° E</geoCoordinate>
),
<specimenCount id="6E81FD671077FFB9FDA7BB5ADEF30FBC" box="[595,685,1911,1936]" pageId="1" pageNumber="120" type="male">2 males</specimenCount>
,
<specimenCount id="6E81FD671077FFB9FD4EBB5ADF660FBC" box="[698,824,1911,1936]" pageId="1" pageNumber="120" type="female">12 females</specimenCount>
,
<date id="0C39102E1077FFB9FCB1BB5BDFED0FBC" box="[837,947,1911,1936]" pageId="1" pageNumber="120" value="1935-06">Jun. 1935</date>
(P. Brien) (
<collectionCode id="1E96AE2B1077FFB9FBCCBB5AD8D20FBC" box="[1080,1164,1912,1936]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="1" pageNumber="120">MRAC</collectionCode>
23.937-23.953);
<collectingCountry id="0090767E1077FFB9FAA5BB5AD9C20FBC" box="[1361,1436,1912,1936]" name="Republic of the Congo" pageId="1" pageNumber="120">Congo</collectingCountry>
River at Kisangani (
<geoCoordinate id="1DB350291077FFB9FE83BBBEDD990F98" box="[375,455,1948,1973]" direction="north" orientation="latitude" pageId="1" pageNumber="120" precision="555" value="0.52">0.52 N</geoCoordinate>
,
<geoCoordinate id="1DB350291077FFB9FE26BBBEDE740F98" box="[466,554,1948,1973]" direction="east" orientation="longitude" pageId="1" pageNumber="120" precision="555" value="25.2">25.20 E</geoCoordinate>
) (
<collectionCode id="1E96AE2B1077FFB9FDB5BBBEDECB0F98" box="[577,661,1948,1972]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="1" pageNumber="120">MRAC</collectionCode>
32.872-32.875, 32.879-32.883, 32.895-32.897); Kaleke, Lake Kisale (
<geoCoordinate id="1DB350291077FFB9FF6BBBE2DCA80FF4" box="[159,246,1984,2009]" direction="south" orientation="latitude" pageId="1" pageNumber="120" precision="555" value="-8.26">8.26° S</geoCoordinate>
,
<geoCoordinate id="1DB350291077FFB9FEF0BBE2DD340FF4" box="[260,362,1984,2009]" direction="east" orientation="longitude" pageId="1" pageNumber="120" precision="555" value="26.49">26.49° E</geoCoordinate>
),
<specimenCount id="6E81FD671077FFB9FE74BBE2DD850FF5" box="[384,475,1984,2009]" pageId="1" pageNumber="120" type="male">2 males</specimenCount>
,
<specimenCount id="6E81FD671077FFB9FE13BBE2DE140FF5" box="[487,586,1984,2009]" pageId="1" pageNumber="120" type="female">1 female</specimenCount>
(
<collectionCode id="1E96AE2B1077FFB9FDAEBBE2DEF00FF4" box="[602,686,1984,2008]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="1" pageNumber="120">MRAC</collectionCode>
32.010-32.012); Lualaba, Kindu, Maniema (
<geoCoordinate id="1DB350291077FFB9FB59BBE2D95A0FF4" box="[1197,1284,1984,2008]" direction="south" orientation="latitude" pageId="1" pageNumber="120" precision="555" value="-2.93">2.93° S</geoCoordinate>
,
<geoCoordinate id="1DB350291077FFB9FAE6BBE2D9260FF4" box="[1298,1400,1984,2009]" direction="east" orientation="longitude" pageId="1" pageNumber="120" precision="555" value="25.94">25.94° E</geoCoordinate>
),
<specimenCount id="6E81FD671077FFBAFA7ABBE2DC8E089C" lastPageId="2" lastPageNumber="121" pageId="1" pageNumber="120" type="male">1 male</specimenCount>
,
<specimenCount id="6E81FD671074FFBAFF2FBCBADD65089C" box="[219,315,151,176]" pageId="2" pageNumber="121" type="female">1 female</specimenCount>
(
<collectionCode id="1E96AE2B1074FFBAFEBDBCBADDC3089C" box="[329,413,152,176]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="2" pageNumber="121">MRAC</collectionCode>
32.898-32.900);
<specimenCount id="6E81FD671074FFBAFDABBCBADE93089C" box="[607,717,151,176]" pageId="2" pageNumber="121" type="female">9 females</specimenCount>
,
<specimenCount id="6E81FD671074FFBAFD2CBCBADF18089C" box="[728,838,151,176]" pageId="2" pageNumber="121" type="juvenile">1 juvenile</specimenCount>
male; Mobi River,
<quantity id="BF7F9B0B1074FFBAFBEBBCBAD838089C" box="[1055,1126,151,176]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="121" unit="km" value="30.0">30 km</quantity>
south of Kisangani (
<geoCoordinate id="1DB350291074FFBAFAB8BCBAD9C9089C" box="[1356,1431,152,176]" direction="south" orientation="latitude" pageId="2" pageNumber="121" precision="555" value="-0.22">0.22 S</geoCoordinate>
,
<geoCoordinate id="1DB350291074FFBAFF63BC9EDCAE08F8" box="[151,240,188,213]" direction="east" orientation="longitude" pageId="2" pageNumber="121" precision="555" value="25.32">25.32 E</geoCoordinate>
),
<specimenCount id="6E81FD671074FFBAFEF0BC9EDD2E08F9" box="[260,368,188,213]" pageId="2" pageNumber="121" type="female">3 females</specimenCount>
(
<collectionCode id="1E96AE2B1074FFBAFE8ABC9EDD8C08F8" box="[382,466,188,212]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="2" pageNumber="121">MRAC</collectionCode>
32.901-32.903); Lake Upemba, Haut-Lomami District (
<geoCoordinate id="1DB350291074FFBAFBB3BC9ED8C208F8" box="[1095,1180,188,213]" direction="south" orientation="latitude" pageId="2" pageNumber="121" precision="555" value="-8.59">8.59° S</geoCoordinate>
,
<geoCoordinate id="1DB350291074FFBAFB5DBC9ED95308F8" box="[1193,1293,188,213]" direction="east" orientation="longitude" pageId="2" pageNumber="121" precision="555" value="26.51">26.51° E</geoCoordinate>
),
<specimenCount id="6E81FD671074FFBAFAD5BC9ED9D408F9" box="[1313,1418,188,213]" pageId="2" pageNumber="121" type="adult">19 adults</specimenCount>
, l female subadult, CW
<date id="0C39102E1074FFBAFE7FBCC2DE6B08D4" box="[395,565,224,248]" pageId="2" pageNumber="121" value="1937-05-01" valueMax="1937-05-07" valueMin="1937-05-01">37, 0107 May</date>
1925 (G. F. de Witte) (
<collectionCode id="1E96AE2B1074FFBAFCC1BCC2DF2E08D4" box="[821,880,224,248]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="121">ZSM</collectionCode>
1193/1, formerly
<collectionCode id="1E96AE2B1074FFBAFBCFBCC2D82B08D4" box="[1083,1141,224,248]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="121">ZSM</collectionCode>
1935, don.
<collectionCode id="1E96AE2B1074FFBAFB0EBCC2D90C08D4" box="[1274,1362,224,248]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="2" pageNumber="121">MRAC</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="2CF866661074FFBAFF63BB61DF0C0F99" httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="2" pageNumber="121" targetBox="[227,1315,312,1773]" targetPageId="2">
<paragraph id="783836EE1074FFBAFF63BB61DF0C0F99" blockId="2.[151,1436,1859,1973]" pageId="2" pageNumber="121">
<emphasis id="4AF3EAFC1074FFBAFF63BB61DD4C0F74" bold="true" box="[151,274,1859,1881]" pageId="2" pageNumber="121">FIGURE 1.</emphasis>
<taxonomicName id="BF874D6D1074FFBAFEECBB66DECC0F75" authority="Rathbun, 1921" authorityName="Rathbun" authorityYear="1921" box="[280,658,1859,1881]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1074FFBAFEECBB66DDBD0F75" box="[280,483,1859,1881]" italics="true" pageId="2" pageNumber="121">Potamonautes langi</emphasis>
(Rathbun, 1921)
</taxonomicName>
. Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D.R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (AI).
</paragraph>
</caption>
<caption id="2CF866661075FFBBFF63BABFDDF50F23" httpUri="https://zenodo.org/record/231924/files/figure.png" pageId="3" pageNumber="122" targetBox="[281,1315,225,1619]" targetPageId="3">
<paragraph id="783836EE1075FFBBFF63BABFDDF50F23" blockId="3.[151,1436,1692,1807]" pageId="3" pageNumber="122">
<emphasis id="4AF3EAFC1075FFBBFF63BABFDD4C0E9E" bold="true" box="[151,274,1693,1715]" pageId="3" pageNumber="122">FIGURE 2.</emphasis>
<taxonomicName id="BF874D6D1075FFBBFEECBABCDECC0E9E" authority="Rathbun, 1921" authorityName="Rathbun" authorityYear="1921" box="[280,658,1692,1715]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1075FFBBFEECBABCDDBD0E9E" box="[280,483,1692,1715]" italics="true" pageId="3" pageNumber="122">Potamonautes langi</emphasis>
(Rathbun, 1921)
</taxonomicName>
. Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D.R. Congo (AMNH 3353). A, mandible frontal view; B, mandibular palp superior view; C, third maxilliped frontal view; D, G1 ventral view; E, G1 dorsal view; F, superior view of terminal article; G, G2 ventral view. Scale bar: 2.5 mm (A), 1.5 mm (B), 12.5 mm C, 4.5mm (DG).
</paragraph>
</caption>
<subSubSection id="309D65651075FFBBFF33BB14DF8A0FF3" pageId="3" pageNumber="122" type="diagnosis">
<paragraph id="783836EE1075FFBBFF33BB14DF8A0FF3" blockId="3.[151,1436,1846,2015]" pageId="3" pageNumber="122">
<emphasis id="4AF3EAFC1075FFBBFF33BB14DD1F0F63" bold="true" box="[199,321,1846,1871]" pageId="3" pageNumber="122">Diagnosis.</emphasis>
Postfrontal crest complete; exorbital, epibranchial teeth both large forward-pointing spines; anterolateral margin of carapace behind epibranchial tooth with 3 long, pointed teeth (
<figureCitation id="E0BC2A6B1075FFBBFB80BB78D8EB0F5E" box="[1140,1205,1882,1907]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="3" pageNumber="122">Fig. 1</figureCitation>
B); ischium of third maxilliped lacking visible sulcus (
<figureCitation id="E0BC2A6B1075FFBBFDE2BB5CDE070FBB" box="[534,601,1918,1943]" captionStart="FIGURE 2" captionStartId="3.[151,250,1693,1715]" captionTargetBox="[281,1315,225,1619]" captionTargetId="figure@3.[279,1331,215,1627]" captionTargetPageId="3" captionText="FIGURE 2. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, mandible frontal view; B, mandibular palp superior view; C, third maxilliped frontal view; D, G 1 ventral view; E, G 1 dorsal view; F, superior view of terminal article; G, G 2 ventral view. Scale bar: 2.5 mm (A), 1.5 mm (B), 12.5 mm C, 4.5 mm (D G)." httpUri="https://zenodo.org/record/231924/files/figure.png" pageId="3" pageNumber="122">Fig. 2</figureCitation>
C); thoracic sternal sulcus s3/s4 incomplete, deep at sides absent in middle (
<figureCitation id="E0BC2A6B1075FFBBFF6BBB80DCBE0F96" box="[159,224,1954,1979]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="3" pageNumber="122">Fig. 1</figureCitation>
C); carpus of cheliped with 2 subequal long, pointed teeth (
<figureCitation id="E0BC2A6B1075FFBBFC65BB80DF8B0F96" box="[913,981,1954,1979]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="3" pageNumber="122">Fig. 1</figureCitation>
F,G); G1 terminal article slim, directed outward at a 60° angle to vertical, tapering to up-curved tip (
<figureCitation id="E0BC2A6B1075FFBBFCCABBE4DFDF0FF3" box="[830,897,1990,2015]" captionStart="FIGURE 2" captionStartId="3.[151,250,1693,1715]" captionTargetBox="[281,1315,225,1619]" captionTargetId="figure@3.[279,1331,215,1627]" captionTargetPageId="3" captionText="FIGURE 2. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, mandible frontal view; B, mandibular palp superior view; C, third maxilliped frontal view; D, G 1 ventral view; E, G 1 dorsal view; F, superior view of terminal article; G, G 2 ventral view. Scale bar: 2.5 mm (A), 1.5 mm (B), 12.5 mm C, 4.5 mm (D G)." httpUri="https://zenodo.org/record/231924/files/figure.png" pageId="3" pageNumber="122">Fig. 2</figureCitation>
D,E,F).
</paragraph>
</subSubSection>
<caption id="2CF866661072FFBCFF63B9CBD9440E32" httpUri="https://zenodo.org/record/231925/files/figure.png" pageId="4" pageNumber="123" targetBox="[386,1200,193,1490]" targetPageId="4">
<paragraph id="783836EE1072FFBCFF63B9CBD9440E32" blockId="4.[151,1436,1513,1566]" pageId="4" pageNumber="123">
<emphasis id="4AF3EAFC1072FFBCFF63B9CBDD4C0DD2" bold="true" box="[151,274,1513,1535]" pageId="4" pageNumber="123">FIGURE 3.</emphasis>
<taxonomicName id="BF874D6D1072FFBCFEECB9C8DECC0DD3" authority="Rathbun, 1921" authorityName="Rathbun" authorityYear="1921" box="[280,658,1513,1535]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="123" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1072FFBCFEECB9C8DDBD0DD3" box="[280,483,1513,1535]" italics="true" pageId="4" pageNumber="123">Potamonautes langi</emphasis>
(Rathbun, 1921)
</taxonomicName>
. Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D.R. Congo (AMNH 3353). Whole animal A, frontal view; B, dorsal view; C, ventral view. Scale bar: 21.4 mm (AC).
</paragraph>
</caption>
<subSubSection id="309D65651072FFBDFF33BA67DDA30AA9" lastPageId="5" lastPageNumber="124" pageId="4" pageNumber="123" type="description">
<paragraph id="783836EE1072FFBDFF33BA67DEC309D8" blockId="4.[151,1437,1605,2026]" lastBlockId="5.[151,1436,151,824]" lastPageId="5" lastPageNumber="124" pageId="4" pageNumber="123">
<emphasis id="4AF3EAFC1072FFBCFF33BA67DD2A0E72" bold="true" box="[199,372,1605,1630]" pageId="4" pageNumber="123">Redescription.</emphasis>
Postfrontal crest complete, epigastric lobes continuous with postorbital crests, lateral ends of postorbital crests meeting anterolateral margins. Exorbital, epibranchial teeth both large forward-pointing spines; anterolateral margin between exorbital, epibranchial teeth smooth, curving slightly outward, lacking intermediate tooth; anterolateral margin posterior to epibranchial tooth with 3 long, pointed teeth (
<figureCitation id="E0BC2A6B1072FFBCFB95BA93D8FC0EE5" box="[1121,1186,1713,1738]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="4" pageNumber="123">Fig. 1</figureCitation>
B). Suborbital margin granular (
<figureCitation id="E0BC2A6B1072FFBCFEF1BAF7DD180EC2" box="[261,326,1749,1774]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="4" pageNumber="123">Fig. 1</figureCitation>
A). Carapace flat (
<collectingCountry id="0090767E1072FFBCFDEFBAF4DE1C0EC2" box="[539,578,1750,1774]" name="Switzerland" pageId="4" pageNumber="123">CH</collectingCountry>
/FW 1.07); front broad, measuring one-third CW (FW/CW 0.33) (
<figureCitation id="E0BC2A6B1072FFBCFAC0BAF7D92B0EC2" box="[1332,1397,1749,1774]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="4" pageNumber="123">Fig. 1</figureCitation>
A). Semi-circular, urogastric, cardiac, posterior, cervical carapace grooves all distinct. Epigastric crests clear, median sulcus between crests short, forked posteriorly (
<figureCitation id="E0BC2A6B1072FFBCFD40BB3FDEAB0F1A" box="[692,757,1821,1846]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="4" pageNumber="123">Fig. 1</figureCitation>
B). Carapace sidewall with distinct granular vertical sulcus, ending at longitudinal sulcus, dividing sidewall into 3 parts (
<figureCitation id="E0BC2A6B1072FFBCFC9EBB63DFF00F75" box="[874,942,1857,1882]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="4" pageNumber="123">Fig. 1</figureCitation>
A). Exopod of third maxilliped with long flagellum, third maxilliped ischium lacking visible sulcus (
<figureCitation id="E0BC2A6B1072FFBCFCDABB47DF2C0F52" box="[814,882,1893,1918]" captionStart="FIGURE 2" captionStartId="3.[151,250,1693,1715]" captionTargetBox="[281,1315,225,1619]" captionTargetId="figure@3.[279,1331,215,1627]" captionTargetPageId="3" captionText="FIGURE 2. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, mandible frontal view; B, mandibular palp superior view; C, third maxilliped frontal view; D, G 1 ventral view; E, G 1 dorsal view; F, superior view of terminal article; G, G 2 ventral view. Scale bar: 2.5 mm (A), 1.5 mm (B), 12.5 mm C, 4.5 mm (D G)." httpUri="https://zenodo.org/record/231924/files/figure.png" pageId="4" pageNumber="123">Fig. 2</figureCitation>
C). Epistomial tooth large, triangular, edges lined by large, round granules (
<figureCitation id="E0BC2A6B1072FFBCFE39BBABDE4D0F8D" box="[461,531,1929,1954]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="4" pageNumber="123">Fig. 1</figureCitation>
A). Mandibular palp two-segmented; terminal segment simple (
<figureCitation id="E0BC2A6B1072FFBCFAFDBBABD90F0F8D" box="[1289,1361,1929,1954]" captionStart="FIGURE 2" captionStartId="3.[151,250,1693,1715]" captionTargetBox="[281,1315,225,1619]" captionTargetId="figure@3.[279,1331,215,1627]" captionTargetPageId="3" captionText="FIGURE 2. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, mandible frontal view; B, mandibular palp superior view; C, third maxilliped frontal view; D, G 1 ventral view; E, G 1 dorsal view; F, superior view of terminal article; G, G 2 ventral view. Scale bar: 2.5 mm (A), 1.5 mm (B), 12.5 mm C, 4.5 mm (D G)." httpUri="https://zenodo.org/record/231924/files/figure.png" pageId="4" pageNumber="123">Fig. 2</figureCitation>
A, B). Thoracic sternal sulcus s2/s3 deep, running horizontally across sternum; thoracic sternal sulcus s3/s4 incomplete, deep at sides absent in middle, sides slanted inward &amp; backward toward anterior margin of sternoabdominal cavity (
<figureCitation id="E0BC2A6B1073FFBDFF6BBCB5DC81089C" box="[159,223,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="5" pageNumber="124">Fig. 1</figureCitation>
C). Thoracic episternal sulci s4/e4, s5/e5, s6/e6, s7/e7 all visible grooves (
<figureCitation id="E0BC2A6B1073FFBDFBD7BCB5D83D089C" box="[1059,1123,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="5" pageNumber="124">Fig. 1</figureCitation>
C). Chelipeds of adult male unequal; fingers of major (right) cheliped long, slender, widely gaping, teeth irregular, those in proximal half bigger than distal half; fingers of
<taxonomicName id="BF874D6D1073FFBDFDFEBCFDDE1308D4" box="[522,589,223,248]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="minor">minor</taxonomicName>
cheliped long, slender, almost meeting. Propodus of major right cheliped of adult male with swollen palm, fixed finger long, straight; movable finger (dactylus) long, curved
<figureCitation id="E0BC2A6B1073FFBDFB22BD26D94B0930" box="[1238,1301,260,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="5" pageNumber="124">Fig. 1</figureCitation>
D, E). First, second carpal teeth of inner margin of carpus of cheliped subequal long, pointed teeth. Lateral, medial inferior margins of merus of cheliped granular, with single large pointed distal meral tooth at distal end; superior surface of merus granular
<figureCitation id="E0BC2A6B1073FFBDFEBCBD4DDDD809A4" box="[328,390,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="5" pageNumber="124">Fig. 1</figureCitation>
F, G). Abdomen slim, triangular, edges straight (
<figureCitation id="E0BC2A6B1073FFBDFC5ABD4DDFB109A4" box="[942,1007,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="5" pageNumber="124">
Fig.
<date id="0C39102E1073FFBDFC17BD52DFB109A4" box="[995,1007,368,392]" pageId="5" pageNumber="124">1</date>
</figureCitation>
I). G1 terminal article about one-third as long as subterminal segment, longitudinal groove visible on dorsal, superior sides (but not on ventral side); G1 terminal article slim, directed outward at a 60° angle to vertical, tapering to up-curved tip (
<figureCitation id="E0BC2A6B1073FFBDFB3EBD95D94F09FC" box="[1226,1297,439,464]" captionStart="FIGURE 2" captionStartId="3.[151,250,1693,1715]" captionTargetBox="[281,1315,225,1619]" captionTargetId="figure@3.[279,1331,215,1627]" captionTargetPageId="3" captionText="FIGURE 2. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, mandible frontal view; B, mandibular palp superior view; C, third maxilliped frontal view; D, G 1 ventral view; E, G 1 dorsal view; F, superior view of terminal article; G, G 2 ventral view. Scale bar: 2.5 mm (A), 1.5 mm (B), 12.5 mm C, 4.5 mm (D G)." httpUri="https://zenodo.org/record/231924/files/figure.png" pageId="5" pageNumber="124">Fig. 2</figureCitation>
D,E,F). G2 terminal article long, flagellum-like (
<figureCitation id="E0BC2A6B1073FFBDFDC2BDFEDE2709D8" box="[566,633,476,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1693,1715]" captionTargetBox="[281,1315,225,1619]" captionTargetId="figure@3.[279,1331,215,1627]" captionTargetPageId="3" captionText="FIGURE 2. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, mandible frontal view; B, mandibular palp superior view; C, third maxilliped frontal view; D, G 1 ventral view; E, G 1 dorsal view; F, superior view of terminal article; G, G 2 ventral view. Scale bar: 2.5 mm (A), 1.5 mm (B), 12.5 mm C, 4.5 mm (D G)." httpUri="https://zenodo.org/record/231924/files/figure.png" pageId="5" pageNumber="124">Fig. 2</figureCitation>
G).
</paragraph>
<paragraph id="783836EE1073FFBDFF33BDDDDF780A34" blockId="5.[151,1436,151,824]" box="[199,806,511,536]" pageId="5" pageNumber="124">
<emphasis id="4AF3EAFC1073FFBDFF33BDDDDCA00A34" bold="true" box="[199,254,511,536]" pageId="5" pageNumber="124">Size.</emphasis>
Large-size species, adult CW
<quantity id="BF7F9B0B1073FFBDFDA0BDDDDEFD0A3B" box="[596,675,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="124" unit="mm" value="45.0">45 mm</quantity>
and higher.
</paragraph>
<paragraph id="783836EE1073FFBDFF33BE06DDA30AA9" blockId="5.[151,1436,151,824]" pageId="5" pageNumber="124">
<emphasis id="4AF3EAFC1073FFBDFF33BE06DD4E0A11" bold="true" box="[199,272,548,573]" pageId="5" pageNumber="124">Color.</emphasis>
Dorsal carapace varying from dark brown/green to dark brown/yellow tinged with red, carapace of newly molted animals much lighter, sternum gray/yellow, abdomen white/grey, chelipeds brown/purple. Cryptic color pattern may be protective.
</paragraph>
</subSubSection>
<subSubSection id="309D65651073FFBEFF33BEADD8530933" lastPageId="6" lastPageNumber="125" pageId="5" pageNumber="124" type="materials_examined">
<paragraph id="783836EE1073FFBDFF33BEADDF720A84" blockId="5.[151,1436,151,824]" box="[199,812,655,680]" pageId="5" pageNumber="124">
<emphasis id="4AF3EAFC1073FFBDFF33BEADDD3B0A84" bold="true" box="[199,357,655,680]" pageId="5" pageNumber="124">
<typeStatus id="A73C884C1073FFBDFF33BEADDD5C0A84" box="[199,258,655,680]" pageId="5" pageNumber="124">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="0090767E1073FFBDFE98BEB2DDE90A84" box="[364,439,656,680]" name="Republic of the Congo" pageId="5" pageNumber="124">Congo</collectingCountry>
River at Kisangani, D.R.
<collectingCountry id="0090767E1073FFBDFD2DBEB2DF760A84" box="[729,808,656,680]" name="Republic of the Congo" pageId="5" pageNumber="124">Congo</collectingCountry>
.
</paragraph>
<paragraph id="783836EE1073FFBDFF33BE96D9110B14" blockId="5.[151,1436,151,824]" pageId="5" pageNumber="124">
<emphasis id="4AF3EAFC1073FFBDFF33BE96DD740AE1" bold="true" box="[199,298,692,717]" pageId="5" pageNumber="124">Habitat.</emphasis>
This is a strictly aquatic species that prefers the quieter parts of the
<collectingCountry id="0090767E1073FFBDFBA3BE96D8FC0AE0" box="[1111,1186,692,716]" name="Republic of the Congo" pageId="5" pageNumber="124">Congo</collectingCountry>
River where floating material accumulates and sunken logs offer protection.
<taxonomicName id="BF874D6D1073FFBDFCE0BEFBDFA90ADC" box="[788,1015,728,752]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1073FFBDFCE0BEFBDFA90ADC" box="[788,1015,728,752]" italics="true" pageId="5" pageNumber="124">Potamonautes langi</emphasis>
</taxonomicName>
rapidly seeks cover when disturbed, including digging into mud. This species was captured in traditional traps baited with putrid or baked manioc set in deep water with a muddy substratum. These traps also catch fishes, including small silurids and mormyrids.
</paragraph>
<caption id="2CF866661073FFBDFF63BBA1DE980F9B" httpUri="https://zenodo.org/record/231926/files/figure.png" pageId="5" pageNumber="124" targetBox="[227,1340,892,1859]" targetPageId="5">
<paragraph id="783836EE1073FFBDFF63BBA1DE980F9B" blockId="5.[151,1435,1923,1975]" pageId="5" pageNumber="124">
<emphasis id="4AF3EAFC1073FFBDFF63BBA1DD4D0FB4" bold="true" box="[151,275,1923,1945]" pageId="5" pageNumber="124">FIGURE 4.</emphasis>
Map of the D.R. Congo in Central Africa showing the known localities of
<taxonomicName id="BF874D6D1073FFBDFBE7BBA6D8810FB5" box="[1043,1247,1923,1945]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="124" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1073FFBDFBE7BBA6D8810FB5" box="[1043,1247,1923,1945]" italics="true" pageId="5" pageNumber="124">Potamonautes langi</emphasis>
</taxonomicName>
(black circles, not shown to scale). See text for exact locality coordinates.
</paragraph>
</caption>
<paragraph id="783836EE1070FFBEFF33BCB5D8530933" blockId="6.[151,1437,151,2015]" pageId="6" pageNumber="125">
<emphasis id="4AF3EAFC1070FFBEFF33BCB5DDE5089C" bold="true" box="[199,443,151,176]" pageId="6" pageNumber="125">Conservation status.</emphasis>
<taxonomicName id="BF874D6D1070FFBEFE36BCBBDEFD089C" box="[450,675,152,176]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFE36BCBBDEFD089C" box="[450,675,152,176]" italics="true" pageId="6" pageNumber="125">Potamonautes langi</emphasis>
</taxonomicName>
was listed as Least Concern (
<bibRefCitation id="1C164B1F1070FFBEFC1BBCBAD829089C" author="IUCN" box="[1007,1143,152,176]" pageId="6" pageNumber="125" refString="IUCN (2003) Guidelines for Application of IUCN Red List Criteria at Regional Levels: version 3.0. IUCN Species Survival Commission. Gland, Switzerland and Cambridge, UK: IUCN." type="book" year="2003">
<collectionCode id="1E96AE2B1070FFBEFC1BBCBAD86B089C" box="[1007,1077,152,176]" pageId="6" pageNumber="125">IUCN</collectionCode>
2003
</bibRefCitation>
; Cumberlidge
<emphasis id="4AF3EAFC1070FFBEFAE8BCBBD915089C" box="[1308,1355,152,176]" italics="true" pageId="6" pageNumber="125">et al</emphasis>
. 2009) because it is known to occur in six localities and has a wide distributional range of about
<quantity id="BF7F9B0B1070FFBEFAF4BC9DD9D308FB" box="[1280,1421,190,215]" metricMagnitude="8" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="125" unit="km" value="140000.0">140,000 km</quantity>
2. Furthermore, there is no evidence of a decline in the extent and quality of its habitat and there are no known longterm threats to its habitat from disturbance and pollution (
<bibRefCitation id="1C164B1F1070FFBEFCE9BD24D85F0933" author="Cumberlidge" box="[797,1025,262,287]" pageId="6" pageNumber="125" refString="Cumberlidge, N. (2008 a) Potamonautes langi. The IUCN Red List of Threatened Species. Version 2014.3. Available from: http: // www. iucnredlist. org (accessed 27 December 2014)" type="journal article" year="2008" yearSuffix="a">Cumberlidge 2008a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="309D65651070FFBEFF33BD08DD6F0A87" pageId="6" pageNumber="125" type="distribution">
<paragraph id="783836EE1070FFBEFF33BD08DD6F0A87" blockId="6.[151,1437,151,2015]" pageId="6" pageNumber="125">
<emphasis id="4AF3EAFC1070FFBEFF33BD08DD3E096F" bold="true" box="[199,352,298,323]" pageId="6" pageNumber="125">Distribution.</emphasis>
This species is found in the tributaries of the Upper
<collectingCountry id="0090767E1070FFBEFC5FBD09DFA8096F" box="[939,1014,299,323]" name="Republic of the Congo" pageId="6" pageNumber="125">Congo</collectingCountry>
River at Kisangani and south into the Katanga district of the D.R.
<collectingCountry id="0090767E1070FFBEFE22BD6DDE7F094B" box="[470,545,335,359]" name="Republic of the Congo" pageId="6" pageNumber="125">Congo</collectingCountry>
(
<figureCitation id="E0BC2A6B1070FFBEFDC4BD6CDE28094B" box="[560,630,334,359]" captionStart="FIGURE 4" captionStartId="5.[151,250,1923,1945]" captionTargetBox="[227,1340,892,1859]" captionTargetId="figure@5.[207,1341,877,1880]" captionTargetPageId="5" captionText="FIGURE 4. Map of the D. R. Congo in Central Africa showing the known localities of Potamonautes langi (black circles, not shown to scale). See text for exact locality coordinates." httpUri="https://zenodo.org/record/231926/files/figure.png" pageId="6" pageNumber="125">Fig. 4</figureCitation>
). Most of Botts (1955) material listed under
<taxonomicName id="BF874D6D1070FFBEFB81BD72D895094B" box="[1141,1227,335,359]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFB81BD72D895094B" box="[1141,1227,335,359]" italics="true" pageId="6" pageNumber="125">P. langi</emphasis>
</taxonomicName>
has been included here, despite the fact that he relied on a non-type adult male specimen of
<emphasis id="4AF3EAFC1070FFBEFC00BD51D81609A6" box="[1012,1096,371,395]" italics="true" pageId="6" pageNumber="125">P. l angi</emphasis>
, which he photographed and illustrated (CW
<quantity id="BF7F9B0B1070FFBEFEA4BDB5DDFA0983" box="[336,420,407,431]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="125" unit="mm" value="42.0">42 mm</quantity>
, MRAC 23.937, plate XII, fig. 1ad).
<bibRefCitation id="1C164B1F1070FFBEFC97BDBADFB90983" author="Bott" box="[867,999,406,431]" pageId="6" pageNumber="125" refString="Bott, R. (1955) Die Suβwasserkrabben von Afrika und ihre Stammesgeschichte. Annales du Musee Royal du Congo Belge, 1 (3), 209 - 352." type="journal article" year="1955">Bott (1955)</bibRefCitation>
incorrectly recorded this specimen as from “Bukoba, Victoria See” (which is in
<collectingCountry id="0090767E1070FFBEFD8CBD98DEBF09FF" box="[632,737,442,467]" name="Tanzania" pageId="6" pageNumber="125">Tanzania</collectingCountry>
), but the museum label actually reads “Bukama” a locality in the Katanga region of the D.R.
<collectingCountry id="0090767E1070FFBEFE02BDFDDE1A09DB" box="[502,580,479,503]" name="Republic of the Congo" pageId="6" pageNumber="125">Congo</collectingCountry>
. There are also doubts as to the identity of two other geographically isolated specimens from the lower
<collectingCountry id="0090767E1070FFBEFE32BE21DE4F0A37" box="[454,529,515,539]" name="Republic of the Congo" pageId="6" pageNumber="125">Congo</collectingCountry>
basin and these have not been included here.
<collectingCountry id="0090767E1070FFBEFBEBBE21D8600A37" box="[1055,1086,515,539]" name="American Samoa" pageId="6" pageNumber="125">As</collectingCountry>
an example, Rathbuns (1921) record of
<taxonomicName id="BF874D6D1070FFBEFEF3BE0ADD000A13" box="[263,350,551,575]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFEF3BE0ADD000A13" box="[263,350,551,575]" italics="true" pageId="6" pageNumber="125">P. langi</emphasis>
</taxonomicName>
from Kinshasa (formerly Léopoldville) in the Lower
<collectingCountry id="0090767E1070FFBEFC30BE05D8510A13" box="[964,1039,551,575]" name="Republic of the Congo" pageId="6" pageNumber="125">Congo</collectingCountry>
basin based on a female specimen collected on
<date id="0C39102E1070FFBEFEDEBE69DD9E0A4F" box="[298,448,587,611]" pageId="6" pageNumber="125" value="1909-08-11">11 Aug 1909</date>
is doubtful due to the lack of male gonopod, cheliped, and sternum characters. Botts (1955) identification of specimens from the
<collectingCountry id="0090767E1070FFBEFD78BE4DDE890AAB" box="[652,727,623,647]" name="Republic of the Congo" pageId="6" pageNumber="125">Congo</collectingCountry>
River at Matadi in the D.R.
<collectingCountry id="0090767E1070FFBEFBEABE4DD8370AAB" box="[1054,1129,623,647]" name="Republic of the Congo" pageId="6" pageNumber="125">Congo</collectingCountry>
(MRAC 21.829-21.831) is also doubtful.
</paragraph>
</subSubSection>
<subSubSection id="309D65651070FFBEFF33BE94D8190E03" pageId="6" pageNumber="125" type="discussion">
<paragraph id="783836EE1070FFBEFF33BE94DE9A0C77" blockId="6.[151,1437,151,2015]" pageId="6" pageNumber="125">
<emphasis id="4AF3EAFC1070FFBEFF33BE94DD650AE3" bold="true" box="[199,315,694,719]" pageId="6" pageNumber="125">Remarks.</emphasis>
This redescription is necessary because Botts (1955) redescription was not based on
<typeStatus id="A73C884C1070FFBEFAF6BE9AD96C0AFC" box="[1282,1330,696,720]" pageId="6" pageNumber="125">type</typeStatus>
material, and instead relied on a series of previously unidentified museum specimens (see above). Moreover, Botts (1955) classification of the large African genus
<taxonomicName id="BF874D6D1070FFBEFD81BF22DF4B0B3B" box="[629,789,768,791]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="4AF3EAFC1070FFBEFD81BF22DF4B0B3B" box="[629,789,768,791]" italics="true" pageId="6" pageNumber="125">Potamonautes</emphasis>
</taxonomicName>
is also questionable, because he placed
<taxonomicName id="BF874D6D1070FFBEFB00BF22D9100B3B" box="[1268,1358,767,791]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFB00BF22D9100B3B" box="[1268,1358,767,791]" italics="true" pageId="6" pageNumber="125">P. langi</emphasis>
</taxonomicName>
in the subgenus
<emphasis id="4AF3EAFC1070FFBEFEF3BF01DD440B16" box="[263,282,803,826]" italics="true" pageId="6" pageNumber="125">P.</emphasis>
(
<taxonomicName id="BF874D6D1070FFBEFEDABF01DE4C0B17" box="[302,530,803,827]" class="Malacostraca" family="Potamonautidae" genus="Obesopotamonautes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="4AF3EAFC1070FFBEFEDABF01DE4C0B17" box="[302,530,803,827]" italics="true" pageId="6" pageNumber="125">Obesopotamonautes</emphasis>
</taxonomicName>
),
<bibRefCitation id="1C164B1F1070FFBEFDDFBF01DEF80B17" author="Bott" box="[555,678,802,827]" pageId="6" pageNumber="125" refString="Bott, R. (1955) Die Suβwasserkrabben von Afrika und ihre Stammesgeschichte. Annales du Musee Royal du Congo Belge, 1 (3), 209 - 352." type="journal article" year="1955">Bott, 1955</bibRefCitation>
, as the
<typeStatus id="A73C884C1070FFBEFD09BF01DF730B17" box="[765,813,803,827]" pageId="6" pageNumber="125">type</typeStatus>
species of a subgenus that also included
<taxonomicName id="BF874D6D1070FFBEFAF1BF01DDC00B73" authority="A. Milne-Edwards, 1868" authorityName="A. Milne-Edwards" authorityYear="1868" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="4AF3EAFC1070FFBEFAF1BF01D9300B17" box="[1285,1390,803,827]" italics="true" pageId="6" pageNumber="125">P. obesus</emphasis>
(A. Milne-Edwards, 1868)
</taxonomicName>
, and
<taxonomicName id="BF874D6D1070FFBEFE1DBF6ADF150B73" authority="Gordon, 1929" authorityName="Gordon" authorityYear="1929" box="[489,843,838,863]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="calcaratus">
<emphasis id="4AF3EAFC1070FFBEFE1DBF6ADEDD0B73" box="[489,643,839,863]" italics="true" pageId="6" pageNumber="125">P. calcaratus</emphasis>
(Gordon, 1929)
</taxonomicName>
. However, the redescription of
<emphasis id="4AF3EAFC1070FFBEFB26BF6AD9630B73" box="[1234,1341,839,863]" italics="true" pageId="6" pageNumber="125">P. ob e s u s</emphasis>
and
<taxonomicName id="BF874D6D1070FFBEFA73BF6ADE090BAF" authority="Reed &amp; Cumberlidge 2004" authorityName="Reed &amp; Cumberlidge" authorityYear="2004" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="calcaratus">
<emphasis id="4AF3EAFC1070FFBEFA73BF6ADD530BAF" italics="true" pageId="6" pageNumber="125">P. calcaratus</emphasis>
(Reed &amp; Cumberlidge 2004)
</taxonomicName>
casts doubt on a close relationship between
<taxonomicName id="BF874D6D1070FFBEFBB8BF49D8FC0BAF" box="[1100,1186,875,899]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFBB8BF49D8FC0BAF" box="[1100,1186,875,899]" italics="true" pageId="6" pageNumber="125">P. langi</emphasis>
</taxonomicName>
and these two species given the significant morphological differences in the gonopods and somatic characters of these two taxa and
<taxonomicName id="BF874D6D1070FFBEFA73BFB2DC8F0BE7" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFA73BFB2DC8F0BE7" italics="true" pageId="6" pageNumber="125">P. langi</emphasis>
</taxonomicName>
. The most recent molecular phylogeny of the Afrotropical potamonautids (
<bibRefCitation id="1C164B1F1070FFBEFBC9BF90D9400BE7" author="Daniels" box="[1085,1310,946,971]" pageId="6" pageNumber="125" refString="Daniels, S. R., Phiri, E., Klaus, S., Albrecht, C. &amp; Cumberlidge, N. (2015) Multi-locus phylogeny of the Afrotropical freshwater crab fauna reveals a Eocene cladogenesis and the impact of paleodrainage rearrangements. Systematic Biology. [in press] http: // dx. doi. org / 10.1093 / sysbio / syv 011" type="book" year="2015">
Daniels
<emphasis id="4AF3EAFC1070FFBEFB6BBF91D88C0BE7" box="[1183,1234,947,971]" italics="true" pageId="6" pageNumber="125">et al</emphasis>
. 2015
</bibRefCitation>
) groups
<taxonomicName id="BF874D6D1070FFBEFA73BF91DCBD0BC3" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="4AF3EAFC1070FFBEFA73BF91DCBD0BC3" italics="true" pageId="6" pageNumber="125">P. obesus</emphasis>
</taxonomicName>
and
<taxonomicName id="BF874D6D1070FFBEFEEBBFFADDEA0BC3" box="[287,436,983,1007]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="calcaratus">
<emphasis id="4AF3EAFC1070FFBEFEEBBFFADDEA0BC3" box="[287,436,983,1007]" italics="true" pageId="6" pageNumber="125">P. calcaratus</emphasis>
</taxonomicName>
together in a separate clade from
<taxonomicName id="BF874D6D1070FFBEFCB1BFFADFC00BC3" box="[837,926,983,1007]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFCB1BFFADFC00BC3" box="[837,926,983,1007]" italics="true" pageId="6" pageNumber="125">P. langi</emphasis>
</taxonomicName>
and does not support the recognition of the subgenus
<emphasis id="4AF3EAFC1070FFBEFEF2BFD9DD470C3E" box="[262,281,1019,1042]" italics="true" pageId="6" pageNumber="125">P.</emphasis>
(
<taxonomicName id="BF874D6D1070FFBEFED8BFD9DE4E0C3F" box="[300,528,1019,1043]" class="Malacostraca" family="Potamonautidae" genus="Obesopotamonautes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="4AF3EAFC1070FFBEFED8BFD9DE4E0C3F" box="[300,528,1019,1043]" italics="true" pageId="6" pageNumber="125">Obesopotamonautes</emphasis>
</taxonomicName>
).
<bibRefCitation id="1C164B1F1070FFBEFDDCBFD8DF580C3F" author="Cumberlidge" box="[552,774,1018,1043]" pageId="6" pageNumber="125" refString="Cumberlidge, N. (1997) The African and Madagascan freshwater crabs in the Museum of Natural History, Vienna (Crustacea: Decapoda: Brachyura: Potamoidea). Annalen des Naturhistorischen Museums in Wien, 99 B, 571 - 589." type="journal article" year="1997">Cumberlidge (1997</bibRefCitation>
,
<bibRefCitation id="1C164B1F1070FFBEFCE6BFD9DF130C3F" author="Cumberlidge" box="[786,845,1019,1043]" pageId="6" pageNumber="125" refString="Cumberlidge, N. (1998) The African and Madagascan freshwater crabs in the Zoologische Staatssammlung, Munich (Crustacea: Decapoda: Brachyura: Potamoidea), Spixiana, 21 (3), 193 - 214." type="journal article" year="1998">1998</bibRefCitation>
), Ng
<emphasis id="4AF3EAFC1070FFBEFC7FBFD9DF9A0C3F" box="[907,964,1019,1043]" italics="true" pageId="6" pageNumber="125">et al.</emphasis>
(2008), and Cumberlidge
<emphasis id="4AF3EAFC1070FFBEFB1ABFD9D9410C3F" box="[1262,1311,1019,1043]" italics="true" pageId="6" pageNumber="125">et al</emphasis>
. (2009) all recognized
<taxonomicName id="BF874D6D1070FFBEFEECB802DD300C1B" box="[280,366,1055,1079]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFEECB802DD300C1B" box="[280,366,1055,1079]" italics="true" pageId="6" pageNumber="125">P. langi</emphasis>
</taxonomicName>
as a valid species of
<taxonomicName id="BF874D6D1070FFBEFDAFB802DEA50C1B" box="[603,763,1056,1079]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="4AF3EAFC1070FFBEFDAFB802DEA50C1B" box="[603,763,1056,1079]" italics="true" pageId="6" pageNumber="125">Potamonautes</emphasis>
</taxonomicName>
and did not recognize the subgenus assignment, but they all made this taxonomic change without explanation.
</paragraph>
<paragraph id="783836EE1070FFBEFF33B844DC890D23" blockId="6.[151,1437,151,2015]" pageId="6" pageNumber="125">
<emphasis id="4AF3EAFC1070FFBEFF33B844DD320C53" bold="true" box="[199,364,1126,1151]" pageId="6" pageNumber="125">Comparisons.</emphasis>
<taxonomicName id="BF874D6D1070FFBEFE87B84ADE0A0C53" box="[371,596,1127,1151]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFE87B84ADE0A0C53" box="[371,596,1127,1151]" italics="true" pageId="6" pageNumber="125">Potamonautes langi</emphasis>
</taxonomicName>
can be recognized by the characters listed above in its diagnosis and by the illustrations and photographs of the
<typeStatus id="A73C884C1070FFBEFDDBB8A8DECF0C8F" box="[559,657,1162,1187]" pageId="6" pageNumber="125" type="holotype">holotype</typeStatus>
(
<figureCitation id="E0BC2A6B1070FFBEFD56B8A8DF530C8F" box="[674,781,1162,1187]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1859,1881]" captionStartId-1="3.[151,250,1693,1715]" captionStartId-2="4.[151,250,1513,1535]" captionTargetBox-0="[227,1315,312,1773]" captionTargetBox-1="[281,1315,225,1619]" captionTargetBox-2="[386,1200,193,1490]" captionTargetId-0="figure@2.[208,1329,301,1778]" captionTargetId-1="figure@3.[279,1331,215,1627]" captionTargetId-2="figure@4.[386,1200,193,1492]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." captionText-1="FIGURE 2. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, mandible frontal view; B, mandibular palp superior view; C, third maxilliped frontal view; D, G 1 ventral view; E, G 1 dorsal view; F, superior view of terminal article; G, G 2 ventral view. Scale bar: 2.5 mm (A), 1.5 mm (B), 12.5 mm C, 4.5 mm (D G)." captionText-2="FIGURE 3. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). Whole animal A, frontal view; B, dorsal view; C, ventral view. Scale bar: 21.4 mm (A C)." httpUri-0="https://zenodo.org/record/231923/files/figure.png" httpUri-1="https://zenodo.org/record/231924/files/figure.png" httpUri-2="https://zenodo.org/record/231925/files/figure.png" pageId="6" pageNumber="125">Figs. 13</figureCitation>
), including those by
<bibRefCitation id="1C164B1F1070FFBEFC09B8A8DCE30CEB" author="Rathbun" pageId="6" pageNumber="125" refString="Rathbun, M. J. (1921) Brachyuran Crabs of the Belgian Congo. Bulletin of the American Museum of Natural History, 43, 379 - 468." type="journal article" year="1921">Rathbun (1921: 430433, pl. 32, fig. 14)</bibRefCitation>
as
<taxonomicName id="BF874D6D1070FFBEFF12B892DEFB0CEB" box="[230,677,1199,1223]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi" subGenus="Acanthopotamonautes">
<emphasis id="4AF3EAFC1070FFBEFF12B892DD150CEB" box="[230,331,1200,1223]" italics="true" pageId="6" pageNumber="125">Potamon</emphasis>
(
<emphasis id="4AF3EAFC1070FFBEFEAAB88DDE070CEB" box="[350,601,1199,1223]" italics="true" pageId="6" pageNumber="125">Acanthopotamonautes</emphasis>
)
<emphasis id="4AF3EAFC1070FFBEFD9FB88DDEFB0CEB" box="[619,677,1199,1223]" italics="true" pageId="6" pageNumber="125">langi</emphasis>
</taxonomicName>
. Illustrations and photographs of another specimen from the D.R.
<collectingCountry id="0090767E1070FFBEFF63B8F1DCBC0CC7" box="[151,226,1235,1259]" name="Republic of the Congo" pageId="6" pageNumber="125">Congo</collectingCountry>
can be found in
<bibRefCitation id="1C164B1F1070FFBEFE5EB8F1DE770CC7" author="Bott" box="[426,553,1234,1259]" pageId="6" pageNumber="125" refString="Bott, R. (1955) Die Suβwasserkrabben von Afrika und ihre Stammesgeschichte. Annales du Musee Royal du Congo Belge, 1 (3), 209 - 352." type="journal article" year="1955">Bott (1955</bibRefCitation>
: pl. XII, fig. 1 ad; fig. 17, 18, 79a) as
<taxonomicName id="BF874D6D1070FFBEFC0FB8F1DC8F0D23" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi" subGenus="Obesopotamonautes">
<emphasis id="4AF3EAFC1070FFBEFC0FB8F1D8C50CC6" box="[1019,1179,1235,1258]" italics="true" pageId="6" pageNumber="125">Potamonautes</emphasis>
(
<emphasis id="4AF3EAFC1070FFBEFB5AB8F1D9CD0CC7" box="[1198,1427,1235,1259]" italics="true" pageId="6" pageNumber="125">Obesopotamonautes</emphasis>
)
<emphasis id="4AF3EAFC1070FFBEFF63B8D5DC8F0D23" box="[151,209,1271,1295]" italics="true" pageId="6" pageNumber="125">langi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="783836EE1070FFBEFF33B938D8190E03" blockId="6.[151,1437,151,2015]" pageId="6" pageNumber="125">
This species is superficially similar to several other species of
<taxonomicName id="BF874D6D1070FFBEFC70B939D87A0D1E" box="[900,1060,1307,1330]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="4AF3EAFC1070FFBEFC70B939D87A0D1E" box="[900,1060,1307,1330]" italics="true" pageId="6" pageNumber="125">Potamonautes</emphasis>
</taxonomicName>
with which it shares a number of taxonomic characters. These include a strong and complete postfrontal crest; strong, sharp exorbital and epibranchial teeth; several distinct spines on the anterolateral margins immediately behind each epibranchial tooth (
<figureCitation id="E0BC2A6B1070FFBEFF6BB9A4DC810DB3" box="[159,223,1414,1439]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="6" pageNumber="125">Fig. 1</figureCitation>
B); a third maxilliped ischium that lacks a visible sulcus (
<figureCitation id="E0BC2A6B1070FFBEFC87B9A4DFE90DB3" box="[883,951,1414,1439]" captionStart="FIGURE 2" captionStartId="3.[151,250,1693,1715]" captionTargetBox="[281,1315,225,1619]" captionTargetId="figure@3.[279,1331,215,1627]" captionTargetPageId="3" captionText="FIGURE 2. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, mandible frontal view; B, mandibular palp superior view; C, third maxilliped frontal view; D, G 1 ventral view; E, G 1 dorsal view; F, superior view of terminal article; G, G 2 ventral view. Scale bar: 2.5 mm (A), 1.5 mm (B), 12.5 mm C, 4.5 mm (D G)." httpUri="https://zenodo.org/record/231924/files/figure.png" pageId="6" pageNumber="125">Fig. 2</figureCitation>
C); a first carpal tooth on the carpus of the cheliped that is strong and pointed; and a strong and pointed distal meral tooth (
<figureCitation id="E0BC2A6B1070FFBEFBACB988D8C10DEF" box="[1112,1183,1450,1475]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="6" pageNumber="125">Fig. 1</figureCitation>
F, G). Other African freshwater crabs with this suite of characters include
<taxonomicName id="BF874D6D1070FFBEFD00B9F2D8FA0DCB" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" box="[756,1188,1486,1511]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="4AF3EAFC1070FFBEFD00B9F2DF2F0DCB" box="[756,881,1487,1511]" italics="true" pageId="6" pageNumber="125">P. niloticus</emphasis>
(H. Milne Edwards, 1837)
</taxonomicName>
(Nile River basin),
<taxonomicName id="BF874D6D1070FFBEFA73B9F2DE710E27" authority="A. Milne-Edwards, 1886" authorityName="A. Milne-Edwards" authorityYear="1886" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="paecilei">
<emphasis id="4AF3EAFC1070FFBEFA73B9F2DCAE0E27" italics="true" pageId="6" pageNumber="125">P. paecilei</emphasis>
(A. Milne-Edwards, 1886)
</taxonomicName>
(Lower
<collectingCountry id="0090767E1070FFBEFD6EB9D1DEBB0E27" box="[666,741,1523,1547]" name="Republic of the Congo" pageId="6" pageNumber="125">Congo</collectingCountry>
River basin),
<emphasis id="4AF3EAFC1070FFBEFC60B9D1D8560E26" box="[916,1032,1523,1546]" italics="true" pageId="6" pageNumber="125">P. w ar re n i</emphasis>
(Calman, 1918), and
<taxonomicName id="BF874D6D1070FFBEFAE1B9D1DD800E03" authority="Stewart, Coke &amp; Cook, 1995" authorityName="Stewart, Coke &amp; Cook" authorityYear="1995" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="4AF3EAFC1070FFBEFAE1B9D1D9C20E27" box="[1301,1436,1523,1547]" italics="true" pageId="6" pageNumber="125">P. dentatus,</emphasis>
<bibRefCitation id="1C164B1F1070FFBEFF63BA35DD800E03" author="Stewart" box="[151,478,1558,1583]" pageId="6" pageNumber="125" refString="Stewart, B. A., Coke, M. &amp; Cook, P. A. (1995) Potamonautes dentatus, new species, a fresh-water crab (Brachyura: Potamoidea: Potamonautidae) from KwaZulu-Natal, South Africa. Journal of Crustacean Biology, 15, 558 - 568. http: // dx. doi. org / 10.2307 / 1548776" type="journal article" year="1995">Stewart, Coke &amp; Cook, 1995</bibRefCitation>
</taxonomicName>
. The latter are found in river basins in southern Africa.
</paragraph>
</subSubSection>
<subSubSection id="309D65651070FFBFFF33BA19DFE00AE0" lastPageId="7" lastPageNumber="126" pageId="6" pageNumber="125" type="reference_group">
<paragraph id="783836EE1070FFBEFF33BA19DDE10F5F" blockId="6.[151,1437,151,2015]" pageId="6" pageNumber="125">
<taxonomicName id="BF874D6D1070FFBEFF33BA19DDF60E7F" ID-CoL="77VN4" box="[199,424,1595,1619]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFF33BA19DDF60E7F" box="[199,424,1595,1619]" italics="true" pageId="6" pageNumber="125">Potamonautes langi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="BF874D6D1070FFBEFDC9BA19DEE40E7F" ID-CoL="4M4JJ" box="[573,698,1595,1619]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="4AF3EAFC1070FFBEFDC9BA19DEE40E7F" box="[573,698,1595,1619]" italics="true" pageId="6" pageNumber="125">P. niloticus</emphasis>
</taxonomicName>
in that the second carpal tooth on the carpus of the cheliped is as large as the first carpal tooth in
<taxonomicName id="BF874D6D1070FFBEFDF0BA42DE050E5B" ID-CoL="77VN4" box="[516,603,1631,1655]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFDF0BA42DE050E5B" box="[516,603,1631,1655]" italics="true" pageId="6" pageNumber="125">P. langi</emphasis>
</taxonomicName>
(
<figureCitation id="E0BC2A6B1070FFBEFD9FBA7CDEF10E5B" box="[619,687,1630,1655]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="6" pageNumber="125">Fig. 1</figureCitation>
F,G) but weak, low and blunt in
<emphasis id="4AF3EAFC1070FFBEFBD8BA42D8EE0E5B" box="[1068,1200,1631,1655]" italics="true" pageId="6" pageNumber="125">
<taxonomicName id="BF874D6D1070FFBEFBD8BA42D8F20E5B" ID-CoL="4M4JJ" box="[1068,1196,1631,1655]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="niloticus">P. niloticus</taxonomicName>
,
</emphasis>
and the G1 terminal article of
<taxonomicName id="BF874D6D1070FFBEFEF1BAA1DD020EB7" ID-CoL="77VN4" box="[261,348,1667,1691]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFEF1BAA1DD020EB7" box="[261,348,1667,1691]" italics="true" pageId="6" pageNumber="125">P. langi</emphasis>
</taxonomicName>
is clearly directed outward at 45° to the longitudinal axis of G1 (
<figureCitation id="E0BC2A6B1070FFBEFBBCBAA0D8D30EB7" box="[1096,1165,1666,1691]" captionStart="FIGURE 2" captionStartId="3.[151,250,1693,1715]" captionTargetBox="[281,1315,225,1619]" captionTargetId="figure@3.[279,1331,215,1627]" captionTargetPageId="3" captionText="FIGURE 2. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, mandible frontal view; B, mandibular palp superior view; C, third maxilliped frontal view; D, G 1 ventral view; E, G 1 dorsal view; F, superior view of terminal article; G, G 2 ventral view. Scale bar: 2.5 mm (A), 1.5 mm (B), 12.5 mm C, 4.5 mm (D G)." httpUri="https://zenodo.org/record/231924/files/figure.png" pageId="6" pageNumber="125">Fig. 2</figureCitation>
D,E), whereas that of
<taxonomicName id="BF874D6D1070FFBEFA73BAA1DCA60E93" ID-CoL="4M4JJ" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="4AF3EAFC1070FFBEFA73BAA1DCA60E93" italics="true" pageId="6" pageNumber="125">P. niloticus</emphasis>
</taxonomicName>
is only slightly angled outward at 25° to the longitudinal axis of G1 (
<bibRefCitation id="1C164B1F1070FFBEFBCCBA85D8F10E93" author="Bott" box="[1080,1199,1702,1727]" pageId="6" pageNumber="125" refString="Bott, R. (1955) Die Suβwasserkrabben von Afrika und ihre Stammesgeschichte. Annales du Musee Royal du Congo Belge, 1 (3), 209 - 352." type="journal article" year="1955">Bott 1955</bibRefCitation>
; Cumberlidge 2009;
<bibRefCitation id="1C164B1F1070FFBEFF63BAE8DD8B0ECF" author="Cumberlidge" box="[151,469,1738,1763]" pageId="6" pageNumber="125" refString="Cumberlidge, N. &amp; Clark, P. F. (2010) Two new species of freshwater crabs from Mt. Elgon, East Africa (Brachyura: Potamoidea: Potamonautidae) with a redescription of Potamonautes niloticus (A. Milne-Edwards, 1887) and notes on their natural history. Journal of Natural History, 44, 1807 - 1829. http: // dx. doi. org / 10.1080 / 00222931003678792" type="journal article" year="2010">Cumberlidge &amp; Clark 2010</bibRefCitation>
).
<taxonomicName id="BF874D6D1070FFBEFE1EBAE9DE930ECF" ID-CoL="77VN4" box="[490,717,1739,1763]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFE1EBAE9DE930ECF" box="[490,717,1739,1763]" italics="true" pageId="6" pageNumber="125">Potamonautes langi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="BF874D6D1070FFBEFC93BAE9DF830ECF" ID-CoL="4M4JM" box="[871,989,1739,1763]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="paecilei">
<emphasis id="4AF3EAFC1070FFBEFC93BAE9DF830ECF" box="[871,989,1739,1763]" italics="true" pageId="6" pageNumber="125">P. paecilei</emphasis>
</taxonomicName>
in that
<emphasis id="4AF3EAFC1070FFBEFBCCBAE9D8D50ECE" box="[1080,1163,1739,1763]" italics="true" pageId="6" pageNumber="125">P. l a ng i</emphasis>
has three large pointed teeth on the anterolateral margin behind the epibranchial tooth (
<figureCitation id="E0BC2A6B1070FFBEFC85BACCDFEC0F2B" box="[881,946,1774,1799]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="6" pageNumber="125">Fig. 1</figureCitation>
B), whereas
<taxonomicName id="BF874D6D1070FFBEFBB1BAD2D8E20F2B" ID-CoL="4M4JM" box="[1093,1212,1775,1799]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="paecilei">
<emphasis id="4AF3EAFC1070FFBEFBB1BAD2D8E20F2B" box="[1093,1212,1775,1799]" italics="true" pageId="6" pageNumber="125">P. paecilei</emphasis>
</taxonomicName>
has two large teeth behind the epibranchial tooth on the anterolateral margin, and the sternal sulcus s3/s4 of
<emphasis id="4AF3EAFC1070FFBEFB85BB31D89F0F06" box="[1137,1217,1811,1835]" italics="true" pageId="6" pageNumber="125">P. l a ng i</emphasis>
is deep at the sides and absent in the middle (
<figureCitation id="E0BC2A6B1070FFBEFE32BB14DE570F63" box="[454,521,1846,1871]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="6" pageNumber="125">Fig. 1</figureCitation>
C), whereas the sternal sulcus s3/s4 of
<taxonomicName id="BF874D6D1070FFBEFC27BB1AD8170F63" ID-CoL="4M4JM" box="[979,1097,1847,1871]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="paecilei">
<emphasis id="4AF3EAFC1070FFBEFC27BB1AD8170F63" box="[979,1097,1847,1871]" italics="true" pageId="6" pageNumber="125">P. paecilei</emphasis>
</taxonomicName>
is completely absent (
<bibRefCitation id="1C164B1F1070FFBEFAA4BB15DC8D0F5F" author="Capart" pageId="6" pageNumber="125" refString="Capart, A. (1954) Revision des types des especes de Potamonidae de l'Afrique Tropicale conserves au Museum d'Histoire Naturelle de Paris. Volume Jubilaire de Victor Van Strallen, Directeur de l'Institut Royal des Sciences Naturelles de Belgique, 1925 - 1934 (II), 819 - 847." type="book chapter" year="1954">Capart 1954</bibRefCitation>
; Cumberlidge 1999).
</paragraph>
<paragraph id="783836EE1070FFBFFF33BBA2DF580948" blockId="6.[151,1437,151,2015]" lastBlockId="7.[151,1436,151,717]" lastPageId="7" lastPageNumber="126" pageId="6" pageNumber="125">
<taxonomicName id="BF874D6D1070FFBEFF33BBA2DDF90FBB" ID-CoL="77VN4" box="[199,423,1919,1943]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFF33BBA2DDF90FBB" box="[199,423,1919,1943]" italics="true" pageId="6" pageNumber="125">Potamonautes langi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="BF874D6D1070FFBEFDCDBBA2DEED0FBB" ID-CoL="4M4H4" box="[569,691,1919,1943]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="4AF3EAFC1070FFBEFDCDBBA2DEED0FBB" box="[569,691,1919,1943]" italics="true" pageId="6" pageNumber="125">P. dentatus</emphasis>
</taxonomicName>
in that the carapace margin between the exorbital and epibranchial teeth is straight in
<taxonomicName id="BF874D6D1070FFBEFE77BB81DD800F97" ID-CoL="77VN4" box="[387,478,1955,1979]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1070FFBEFE77BB81DD800F97" box="[387,478,1955,1979]" italics="true" pageId="6" pageNumber="125">P. langi</emphasis>
</taxonomicName>
and there are three large pointed teeth on the anterolateral margin behind the epibranchial tooth (
<figureCitation id="E0BC2A6B1070FFBEFE81BBE4DDE90FF3" box="[373,439,1990,2015]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="6" pageNumber="125">Fig. 1</figureCitation>
B), whereas in
<taxonomicName id="BF874D6D1070FFBEFD9DBBEADEB80FF3" ID-CoL="4M4H4" box="[617,742,1991,2015]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="4AF3EAFC1070FFBEFD9DBBEADEB80FF3" box="[617,742,1991,2015]" italics="true" pageId="6" pageNumber="125">P. dentatus</emphasis>
</taxonomicName>
there is a deep sinus posterior to the exorbital tooth and the pointed teeth on the anterolateral margin behind the epibranchial tooth are all small. The sternal sulcus s3/s4 of
<taxonomicName id="BF874D6D1071FFBFFA73BCBBDC8F08F8" ID-CoL="77VN4" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1071FFBFFA73BCBBDC8F08F8" italics="true" pageId="7" pageNumber="126">P. langi</emphasis>
</taxonomicName>
is deep at the sides and absent in the middle (
<figureCitation id="E0BC2A6B1071FFBFFD20BC9EDF4D08F8" box="[724,787,188,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1859,1881]" captionTargetBox="[227,1315,312,1773]" captionTargetId="figure@2.[208,1329,301,1778]" captionTargetPageId="2" captionText="FIGURE 1. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, carapace, frontal view; B, dorsal view; C, anterior thoracic sternum; D, right cheliped, frontal view; E, left cheliped, frontal view; F, carpus and merus of right cheliped, dorsal view; G, carpus and merus of right cheliped, ventral view; H, pereiopod 3; I, pleon (abdomen). Scale bar: 10.8 mm (A I)." httpUri="https://zenodo.org/record/231923/files/figure.png" pageId="7" pageNumber="126">Fig. 1</figureCitation>
C) whereas sternal sulcus s3/s4 of
<taxonomicName id="BF874D6D1071FFBFFB68BC9FD94808F8" ID-CoL="4M4H4" box="[1180,1302,188,212]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="4AF3EAFC1071FFBFFB68BC9FD94808F8" box="[1180,1302,188,212]" italics="true" pageId="7" pageNumber="126">P. dentatus</emphasis>
</taxonomicName>
is deep and complete and forms a distinct V-shape with the bottom of the V meeting the sternoabdominal cavity (
<bibRefCitation id="1C164B1F1071FFBFFAF2BCC2DC8D0931" author="Stewart" pageId="7" pageNumber="126" refString="Stewart, B. A., Coke, M. &amp; Cook, P. A. (1995) Potamonautes dentatus, new species, a fresh-water crab (Brachyura: Potamoidea: Potamonautidae) from KwaZulu-Natal, South Africa. Journal of Crustacean Biology, 15, 558 - 568. http: // dx. doi. org / 10.2307 / 1548776" type="journal article" year="1995">
Stewart
<emphasis id="4AF3EAFC1071FFBFFA90BCC3D9CB08D4" box="[1380,1429,224,248]" italics="true" pageId="7" pageNumber="126">et al</emphasis>
. 1995
</bibRefCitation>
).
<taxonomicName id="BF874D6D1071FFBFFF1CBD27DD950930" ID-CoL="77VN4" box="[232,459,260,284]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="langi">
<emphasis id="4AF3EAFC1071FFBFFF1CBD27DD950930" box="[232,459,260,284]" italics="true" pageId="7" pageNumber="126">Potamonautes langi</emphasis>
</taxonomicName>
differs from
<emphasis id="4AF3EAFC1071FFBFFD90BD27DE890930" box="[612,727,261,284]" italics="true" pageId="7" pageNumber="126">P. w arreni</emphasis>
in that
<emphasis id="4AF3EAFC1071FFBFFCC0BD27DFD90930" box="[820,903,260,284]" italics="true" pageId="7" pageNumber="126">P. l a ng i</emphasis>
has a slim G1 terminal article that is strongly directed outward at a 45° angle (
<figureCitation id="E0BC2A6B1071FFBFFDF3BD05DE15096C" box="[519,587,295,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1693,1715]" captionTargetBox="[281,1315,225,1619]" captionTargetId="figure@3.[279,1331,215,1627]" captionTargetPageId="3" captionText="FIGURE 2. Potamonautes langi (Rathbun, 1921). Holotype, adult male, CW 45 mm, from the Congo River at Kisangani, D. R. Congo (AMNH 3353). A, mandible frontal view; B, mandibular palp superior view; C, third maxilliped frontal view; D, G 1 ventral view; E, G 1 dorsal view; F, superior view of terminal article; G, G 2 ventral view. Scale bar: 2.5 mm (A), 1.5 mm (B), 12.5 mm C, 4.5 mm (D G)." httpUri="https://zenodo.org/record/231924/files/figure.png" pageId="7" pageNumber="126">Fig. 2</figureCitation>
D,E), whereas the G1 terminal article of
<emphasis id="4AF3EAFC1071FFBFFBE3BD0BD8D90913" box="[1047,1159,296,320]" italics="true" pageId="7" pageNumber="126">P. w a r ren i</emphasis>
is only slightly directed outward at a 30° angle (
<bibRefCitation id="1C164B1F1071FFBFFE56BD6EDE480949" author="Bott" box="[418,534,332,357]" pageId="7" pageNumber="126" refString="Bott, R. (1955) Die Suβwasserkrabben von Afrika und ihre Stammesgeschichte. Annales du Musee Royal du Congo Belge, 1 (3), 209 - 352." type="journal article" year="1955">Bott 1955</bibRefCitation>
;
<bibRefCitation id="1C164B1F1071FFBFFDD5BD6EDEA70948" author="Daniels" box="[545,761,332,357]" pageId="7" pageNumber="126" refString="Daniels, S. R., Stewart, B. A., Ridgway, T. M. &amp; Florence, W. (2001) Carapace dentition patterns, morphometrics and allozyme differentiation amongst two toothed freshwater crab species (Potamonautes warreni and P. unispinus) (Decapoda: Brachyura: Potamonautidae) from river systems in South Africa. Journal of Zoology, 255, 389 - 404. http: // dx. doi. org / 10.1017 / S 0952836901001480" type="journal article" year="2001">
Daniels
<emphasis id="4AF3EAFC1071FFBFFD8ABD6FDEF10948" box="[638,687,332,356]" italics="true" pageId="7" pageNumber="126">et al</emphasis>
. 2001
</bibRefCitation>
).
</paragraph>
<paragraph id="783836EE1071FFBFFF33BD52DFE00AE0" blockId="7.[151,1436,151,717]" pageId="7" pageNumber="126">
Four other Afrotropical genera include species that have strong, sharp exorbital and epibranchial teeth and several pointed teeth on the anterolateral margin immediately behind the epibranchial tooth. These are
<taxonomicName id="BF874D6D1071FFBFFF63BD9ADE8809FC" ID-CoL="4JZJQ" authority="A. Milne-Edwards, 1887" authorityName="A. Milne-Edwards" authorityYear="1887" box="[151,726,439,464]" class="Malacostraca" family="Potamonautidae" genus="Platythelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="armata">
<emphasis id="4AF3EAFC1071FFBFFF63BD9ADDC509FC" box="[151,411,440,464]" italics="true" pageId="7" pageNumber="126">Platythelphusa armata</emphasis>
(A. Milne-Edwards, 1887)
</taxonomicName>
(Lake Tanganyika),
<taxonomicName id="BF874D6D1071FFBFFC3EBD9ADC8609D9" ID-CoL="3B29P" authority="A. Milne-Edwards, 1886" authorityName="A. Milne-Edwards" authorityYear="1886" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="brazzae">
<emphasis id="4AF3EAFC1071FFBFFC3EBD9AD8F209FC" box="[970,1196,440,464]" italics="true" pageId="7" pageNumber="126">Erimetopus brazzae</emphasis>
(A. Milne-Edwards, 1886)
</taxonomicName>
(Lower
<collectingCountry id="0090767E1071FFBFFEBBBDFEDDC409D8" box="[335,410,476,500]" name="Republic of the Congo" pageId="7" pageNumber="126">Congo</collectingCountry>
River basin),
<taxonomicName id="BF874D6D1071FFBFFDADBDFED8FB09D8" ID-CoL="6Q944" authority="A. Milne-Edwards, 1887" authorityName="A. Milne-Edwards" authorityYear="1887" box="[601,1189,476,501]" class="Malacostraca" family="Potamonautidae" genus="Liberonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="chaperi">
<emphasis id="4AF3EAFC1071FFBFFDADBDFEDF0D09D8" box="[601,851,476,500]" italics="true" pageId="7" pageNumber="126">Liberonautes chaperi</emphasis>
(A. Milne-Edwards, 1887)
</taxonomicName>
(West Africa), and
<taxonomicName id="BF874D6D1071FFBFFF63BE22DEDB0A34" ID-CoL="7B2Z7" authority="Rathbun, 1921" authorityName="Rathbun" authorityYear="1921" box="[151,645,511,536]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="faradjensis">
<emphasis id="4AF3EAFC1071FFBFFF63BE22DDE30A34" box="[151,445,512,536]" italics="true" pageId="7" pageNumber="126">Sudanonautes faradjensis</emphasis>
(
<bibRefCitation id="1C164B1F1071FFBFFE24BDDDDE230A34" author="Rathbun" box="[464,637,511,536]" pageId="7" pageNumber="126" refString="Rathbun, M. J. (1921) Brachyuran Crabs of the Belgian Congo. Bulletin of the American Museum of Natural History, 43, 379 - 468." type="journal article" year="1921">Rathbun, 1921</bibRefCitation>
)
</taxonomicName>
(Central Africa) but they can all be distinguished from species of
<taxonomicName id="BF874D6D1071FFBFFF63BE07DD690A10" ID-CoL="7PVHW" box="[151,311,549,572]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="4AF3EAFC1071FFBFFF63BE07DD690A10" box="[151,311,549,572]" italics="true" pageId="7" pageNumber="126">Potamonautes</emphasis>
</taxonomicName>
at the genus level (
<bibRefCitation id="1C164B1F1071FFBFFDE1BE06DEB20A11" author="Cumberlidge" box="[533,748,548,573]" pageId="7" pageNumber="126" refString="Cumberlidge, N. (1985) Redescription of Liberonautes chaperi (A. Milne-Edwards, 1887) n. comb., a fresh-water crab from Ivory Coast (Brachyura, Potamonautidae). Canadian Journal of Zoology, 63, 2704 - 2707. http: // dx. doi. org / 10.1139 / z 85 - 404" type="journal article" year="1985">Cumberlidge 1985</bibRefCitation>
,
<bibRefCitation id="1C164B1F1071FFBFFD0CBE06DF6D0A11" author="Cumberlidge" box="[760,819,548,573]" pageId="7" pageNumber="126" refString="Cumberlidge, N. (1995) Redescription of the Central African fresh-water crab Sudanonautes faradjensis (Rathbun, 1921) (Brachyura: Potamoidea: Potamonautidae). Proceedings of the Biological Society of Washington, 108 (4), 629 - 636." type="journal article" year="1995">1995</bibRefCitation>
, 1999; Cumberlidge
<emphasis id="4AF3EAFC1071FFBFFBD5BE07D80D0A10" box="[1057,1107,548,572]" italics="true" pageId="7" pageNumber="126">et al</emphasis>
. 1999; Cumberlidge &amp; Reed 2004). In contrast to
<emphasis id="4AF3EAFC1071FFBFFE8ABE6BDEE10A4C" box="[382,703,584,608]" italics="true" pageId="7" pageNumber="126">
<taxonomicName id="BF874D6D1071FFBFFE8ABE6BDE7F0A4C" ID-CoL="7PVHW" box="[382,545,585,608]" class="Malacostraca" family="Potamonautidae" genus="Potamonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="126" phylum="Arthropoda" rank="genus">Potamonautes</taxonomicName>
,
<taxonomicName id="BF874D6D1071FFBFFDD8BE6ADEE10A4C" ID-CoL="7PMS4" box="[556,703,584,608]" class="Malacostraca" family="Potamonautidae" genus="Liberonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="126" phylum="Arthropoda" rank="genus">Liberonautes</taxonomicName>
</emphasis>
and
<taxonomicName id="BF874D6D1071FFBFFD00BE6ADFCF0A4C" ID-CoL="7Q3BJ" box="[756,913,584,608]" class="Malacostraca" family="Potamonautidae" genus="Sudanonautes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="4AF3EAFC1071FFBFFD00BE6ADFCF0A4C" box="[756,913,584,608]" italics="true" pageId="7" pageNumber="126">Sudanonautes</emphasis>
</taxonomicName>
have an intermediate tooth on the anterolateral margin between the exorbital and epibranchial teeth (Cumberlidge 1999);
<taxonomicName id="BF874D6D1071FFBFFC22BE4FD8090AA8" ID-CoL="7PDSH" box="[982,1111,621,644]" class="Malacostraca" family="Potamonautidae" genus="Erimetopus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="4AF3EAFC1071FFBFFC22BE4FD8090AA8" box="[982,1111,621,644]" italics="true" pageId="7" pageNumber="126">Erimetopus</emphasis>
</taxonomicName>
has spines on the margins of the merus of P2-P5 (Cumberlidge &amp; Reed 2004); and
<taxonomicName id="BF874D6D1071FFBFFD05BEB2DFC70A84" ID-CoL="7PTVW" box="[753,921,656,680]" class="Malacostraca" family="Potamonautidae" genus="Platythelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="4AF3EAFC1071FFBFFD05BEB2DFC70A84" box="[753,921,656,680]" italics="true" pageId="7" pageNumber="126">Platythelphusa</emphasis>
</taxonomicName>
has a frontal margin that projects forward and is not downturned over the antennular fossae (Cumberlidge
<emphasis id="4AF3EAFC1071FFBFFCC2BE97DF390AE0" box="[822,871,692,716]" italics="true" pageId="7" pageNumber="126">et al</emphasis>
. 1999).
</paragraph>
</subSubSection>
</treatment>
</document>