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<document id="3B30F60B82A6C134375E62723AFEE8B1" ID-CLB-Dataset="21548" ID-DOI="10.11646/zootaxa.4809.2.6" ID-GBIF-Dataset="d53c2985-17f2-40f0-a70b-d6d1f0ba9d9a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3934208" ID-ZooBank="ECB2E640-6102-4230-8CF3-730AD1038AC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594126239908" checkinUser="plazi" docAuthor="Lopes, Peterson Lásaro, Gasca-Álvarez, Héctor Jaime &amp; Skelley, Paul E." docDate="2020" docId="F029879CC22C114BFF4CF949FECB3E01" docLanguage="en" docName="zootaxa.4809.2.6.pdf" docOrigin="Zootaxa 4809 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bacis nigropictus Crotch 1876" docType="treatment" docVersion="7" lastPageNumber="353" masterDocId="0C10FFE4C22D114DFFDBFFF7FFFD3F21" masterDocTitle="Redescriptions, new synonymy, new combination and new records of Bacis Dejean, 1836 and Oligocorynus Dejean, 1876 (Coleoptera: Erotylidae: Erotylinae) for Colombia" masterLastPageNumber="362" masterPageNumber="349" pageNumber="350" updateTime="1698843165593" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D814ACC32596582DC733B6A58288FFD4">Redescriptions, new synonymy, new combination and new records of Bacis Dejean, 1836 and Oligocorynus Dejean, 1876 (Coleoptera: Erotylidae: Erotylinae) for Colombia</mods:title>
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<mods:namePart id="8743B0FB71218478351038862904A8AF">Lopes, Peterson Lásaro</mods:namePart>
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<mods:namePart id="A277AC8C3B71BC009FF24148834D30BD">Gasca-Álvarez, Héctor Jaime</mods:namePart>
<mods:affiliation id="C1A4F2684ECA9A024D4A395654DBD88F">Research Program, Corporación Sentido Natural, Bogotá, Colombia.</mods:affiliation>
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<mods:namePart id="193F67F5CED288189AF021A987F2E1EF">Skelley, Paul E.</mods:namePart>
<mods:affiliation id="A1A1FFA91205B2B77E42C72525BD79C4">Florida State Collection of Arthropods, Florida Department of Agriculture and Consumer Services - DPI, P. O. Box 147100, Gainesville, FL 32614 - 7100 USA.</mods:affiliation>
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<treatment id="F029879CC22C114BFF4CF949FECB3E01" ID-DOI="http://doi.org/10.5281/zenodo.4323977" ID-GBIF-Taxon="164971378" ID-Zenodo-Dep="4323977" LSID="urn:lsid:plazi:treatment:F029879CC22C114BFF4CF949FECB3E01" httpUri="http://treatment.plazi.org/id/F029879CC22C114BFF4CF949FECB3E01" lastPageId="6" lastPageNumber="353" pageId="1" pageNumber="350">
<subSubSection id="309A6501C22C114CFF4CF949FDEE39F8" box="[151,531,1726,1753]" pageId="1" pageNumber="350" type="nomenclature">
<paragraph id="783F368AC22C114CFF4CF949FDEE39F8" blockId="1.[151,531,1726,1789]" box="[151,531,1726,1753]" pageId="1" pageNumber="350">
<heading id="237781E6C22C114CFF4CF949FDEE39F8" bold="true" box="[151,531,1726,1753]" fontSize="11" level="1" pageId="1" pageNumber="350" reason="1">
<taxonomicName id="BF804D09C22C114CFF4CF949FDEE39F8" authority="Crotch, 1876" authorityName="Crotch" authorityYear="1876" box="[151,531,1726,1753]" class="Insecta" family="Erotylidae" genus="Bacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="4AF4EA98C22C114CFF4CF949FDEE39F8" bold="true" box="[151,531,1726,1753]" pageId="1" pageNumber="350">
<emphasis id="4AF4EA98C22C114CFF4CF949FE9739F9" bold="true" box="[151,362,1726,1752]" italics="true" pageId="1" pageNumber="350">Bacis nigropictus</emphasis>
<bibRefCitation id="1C114B7BC22C114CFEA9F949FDEE39F8" author="Crotch, G. R." box="[370,531,1726,1753]" pageId="1" pageNumber="350" pagination="377 - 572" refId="ref5170" refString="Crotch, G. R. (1876) A revision of the coleopterous family Erotylidae. Cistula Entomologica, 1, 377 - 572." type="journal article" year="1876">Crotch, 1876</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="309A6501C22C114CFF4CF915FE8939DC" box="[151,372,1762,1789]" pageId="1" pageNumber="350" type="description">
<paragraph id="783F368AC22C114CFF4CF915FE8939DC" blockId="1.[151,531,1726,1789]" box="[151,372,1762,1789]" pageId="1" pageNumber="350">
(
<figureCitation id="E0BB2A0FC22C114CFF44F915FEE439DC" box="[159,281,1762,1789]" captionStart-0="FIGURES 18" captionStart-1="FIGURES 916" captionStart-2="FIGURES 1725" captionStart-3="FIGURES 2629" captionStartId-0="2.[151,264,1946,1970]" captionStartId-1="3.[151,264,1918,1942]" captionStartId-2="5.[151,264,1912,1936]" captionStartId-3="6.[151,264,1515,1539]" captionTargetBox-0="[164,1427,184,1906]" captionTargetBox-1="[178,1402,212,1826]" captionTargetBox-2="[262,1300,195,1849]" captionTargetBox-3="[216,1296,329,1451]" captionTargetId-0="figure@2.[153,1434,179,1917]" captionTargetId-1="figure@3.[166,1422,178,1892]" captionTargetId-2="figure@5.[201,1387,181,1886]" captionTargetId-3="figure@6.[151,1436,306,1492]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURES 18. Bacis nigropictus Crotch, 1876, specimen from Brazil. Habitus: (12) dorsal, (3) lateral, and (4) ventral view (scale bars for 14 = 3 mm); (5) head (scale bar = 0.3 mm); (6) left antenna (scale bar = 1 mm); (7) labrum (scale bar = 0.3 mm); (8) epipharynx (scale bar = 0.3 mm; arrow: torma)." captionText-1="FIGURES 916. Bacis nigropictus Crotch, 1876. (910) Mandibles, dorsal view (arrow: proximal incisor); (1112) mandibles, ventral view (arrow: proximal incisor); (13) maxilla, dorsal view, (14) maxilla, ventral view, (15) hypopharynx, dorsal view (arrow: hypopharynx arm); (16) labium, dorsal view. Scale bars = 0.3 mm." captionText-2="FIGURES 1725. Bacis nigropictus Crotch, 1876. (17) Right metathoracic wing (scale bar = 3 mm); (18) eighth abdominal tergite; (19) eighth abdominal sternite; (20) tenth abdominal tergite; (21) ninth abdominal sternite (scale bars for 1821 = 0.3 mm); (2224) aedeagus, dorsal, ventral, and lateral view, respectively (scale bar = 0.5 mm); (25) head of flagellum (scale bar = 0.1 mm)." captionText-3="FIGURES 2629. Bacis nigropictus Crotch, 1876. (26) Eighth abdominal tergite; (27) eighth abdominal sternite; (28) ovipositor, dorsal view (arrow: proctiger lobe); (29) ovipositor, ventral view. Scale bars = 0.3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3934210" figureDoi-1="http://doi.org/10.5281/zenodo.3934212" figureDoi-2="http://doi.org/10.5281/zenodo.3934214" figureDoi-3="http://doi.org/10.5281/zenodo.3934216" httpUri-0="https://zenodo.org/record/3934210/files/figure.png" httpUri-1="https://zenodo.org/record/3934212/files/figure.png" httpUri-2="https://zenodo.org/record/3934214/files/figure.png" httpUri-3="https://zenodo.org/record/3934216/files/figure.png" pageId="1" pageNumber="350">Figs. 129</figureCitation>
,
<figureCitation id="E0BB2A0FC22C114CFEFEF915FE9039DC" box="[293,365,1762,1789]" captionStart="FIGURES 5760" captionStartId="12.[151,264,1673,1697]" captionTargetBox="[158,1431,185,1636]" captionTargetId="figure@12.[151,1436,181,1647]" captionTargetPageId="12" captionText="FIGURES 5760. Type material of Zonarius limbatus Kuhnt, 1911. (57) Lectotype; (5860), paralectotypes. Photographs provided by Bernd Jaeger." figureDoi="http://doi.org/10.5281/zenodo.3934226" httpUri="https://zenodo.org/record/3934226/files/figure.png" pageId="1" pageNumber="350">5760</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="309A6501C22C114CFF4CF8D0FCE938D6" pageId="1" pageNumber="350" type="reference_group">
<paragraph id="783F368AC22C114CFF4CF8D0FC8E387C" blockId="1.[151,1437,1830,2039]" pageId="1" pageNumber="350">
<treatmentCitationGroup id="589011A4C22C114CFF4CF8D0FC8E387C" pageId="1" pageNumber="350">
<treatmentCitation id="F921109BC22C114CFF4CF8D0FDF5381E" author="Crotch, G. R." box="[151,520,1830,1855]" page="557" pageId="1" pageNumber="350" year="1876">
<taxonomicName id="BF804D09C22C114CFF4CF8D0FDF5381E" authority="Crotch 1876: 557" authorityName="Crotch" authorityPageNumber="557" authorityYear="1876" box="[151,520,1830,1855]" class="Insecta" family="Erotylidae" genus="Bacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="4AF4EA98C22C114CFF4CF8D0FEB4381F" box="[151,329,1831,1854]" italics="true" pageId="1" pageNumber="350">Bacis nigropictus</emphasis>
<bibRefCitation id="1C114B7BC22C114CFE8AF8D0FDF5381E" author="Crotch, G. R." box="[337,520,1830,1855]" pageId="1" pageNumber="350" pagination="377 - 572" refId="ref5170" refString="Crotch, G. R. (1876) A revision of the coleopterous family Erotylidae. Cistula Entomologica, 1, 377 - 572." type="journal article" year="1876">Crotch 1876: 557</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="F921109BC22C114CFDCFF8D0FD1B381E" author="Fleutiaux, E." box="[532,742,1831,1855]" page="224" pageId="1" pageNumber="350" year="1886">
<bibRefCitation id="1C114B7BC22C114CFDCFF8D0FD1B381E" author="Fleutiaux, E." box="[532,742,1831,1855]" pageId="1" pageNumber="350" pagination="216 - 224" refId="ref5299" refString="Fleutiaux, E. (1886) Supplement au catalogue des Coleopteres de M. M. Gemminger et de Harold (Languriides et Erotylides). Annales de la Societe Entomologique de Belgique, 30, 216 - 224." type="journal article" year="1886">Fleutiaux 1886: 224</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F921109BC22C114CFD2AF8D0FC6B381E" author="Kuhnt, P." box="[753,918,1831,1855]" page="48" pageId="1" pageNumber="350" year="1909">
<bibRefCitation id="1C114B7BC22C114CFD2AF8D0FC6B381E" author="Kuhnt, P." box="[753,918,1831,1855]" pageId="1" pageNumber="350" pagination="1 - 139" refId="ref5405" refString="Kuhnt, P. (1909) Coleoptera, fam. Erotylidae, subfam. Erotylinae. In: Wytsman, P. (Ed.), Genera Insectorum. Fasc. 88. V. Verteneuil &amp; L. Desmet, Brussels, pp. 1 - 139, 4 pls." type="book chapter" year="1909">Kuhnt 1909: 48</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="F921109BC22C114CFC7AF8D0FC00381E" author="Kuhnt, P." box="[929,1021,1831,1855]" page="34" pageId="1" pageNumber="350" year="1911">
<bibRefCitation id="1C114B7BC22C114CFC7AF8D0FC00381E" author="Kuhnt, P." box="[929,1021,1831,1855]" pageId="1" pageNumber="350" pagination="1 - 103" refId="ref5492" refString="Kuhnt, P. (1911) Family Erotylidae. In: Junk, W. (Ed.), Coleopterorum catalogus, Pars 34 (15), pp. 1 - 103." type="journal article" year="1911">1911: 34</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F921109BC22C114CFBD2F8D0FB07381E" author="Blackwelder, R. E." box="[1033,1274,1830,1855]" page="463" pageId="1" pageNumber="350" year="1945">
<bibRefCitation id="1C114B7BC22C114CFBD2F8D0FB07381E" author="Blackwelder, R. E." box="[1033,1274,1830,1855]" pageId="1" pageNumber="350" pagination="343 - 550" refId="ref5075" refString="Blackwelder, R. E. (1945) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Bulletin of the United States National Museum, 185 (3), 343 - 550. https: // doi. org / 10.5479 / si. 03629236.185.3" type="journal article" year="1945">Blackwelder 1945: 463</bibRefCitation>
</treatmentCitation>
, list;
<treatmentCitation id="F921109BC22C114CFAE8F8D0FED3387C" author="Alvarenga, M." page="118" pageId="1" pageNumber="350" year="1994">
<bibRefCitation id="1C114B7BC22C114CFAE8F8D0FED3387C" author="Alvarenga, M." pageId="1" pageNumber="350" pagination="1 - 175" refId="ref5033" refString="Alvarenga, M. (1994) Catalogo dos Erotylidae (Coleoptera) Neotropicais. Revista Brasileira de Zoologia, 11 (1), 1 - 175. https: // doi. org / 10.1590 / S 0101 - 81751994000100001" type="journal article" year="1994">Alvarenga 1994: 118</bibRefCitation>
</treatmentCitation>
, catalog.
<typeStatus id="A73B8828C22C114CFE55F8B1FE3D387F" box="[398,448,1862,1886]" pageId="1" pageNumber="350">Type</typeStatus>
series:
<collectingCountry id="0097761AC22C114CFDD6F8B2FD88387C" box="[525,629,1861,1885]" name="Venezuela" pageId="1" pageNumber="350">Venezuela</collectingCountry>
(UMZC, not examined).
</treatmentCitationGroup>
</paragraph>
<paragraph id="783F368AC22C114CFF4CF892FEC438F9" blockId="1.[151,1437,1830,2039]" pageId="1" pageNumber="350">
<treatmentCitationGroup id="589011A4C22C114CFF4CF892FEC438F9" pageId="1" pageNumber="350">
<treatmentCitation id="F921109BC22C114CFF4CF892FDF3385D" author="Kuhnt, P." box="[151,526,1892,1916]" page="249" pageId="1" pageNumber="350" year="1910">
<taxonomicName id="BF804D09C22C114CFF4CF892FDF3385D" authority="Kuhnt 1910: 249" authorityName="Kuhnt" authorityPageNumber="249" authorityYear="1910" box="[151,526,1892,1916]" class="Insecta" family="Erotylidae" genus="Zonarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="4AF4EA98C22C114CFF4CF892FEAE385D" box="[151,339,1892,1916]" italics="true" pageId="1" pageNumber="350">Zonarius limbatus</emphasis>
<bibRefCitation id="1C114B7BC22C114CFE87F893FDF3385D" author="Kuhnt, P." box="[348,526,1892,1916]" pageId="1" pageNumber="350" pagination="219 - 270" refId="ref5459" refString="Kuhnt, P. (1910) Neue Erotylidae (Col.). Mitteilungen aus dem Museum fur Naturkunde in Berlin: Deutsche Entomologische Zeitschrift, 28, 219 - 270." type="journal article" year="1910">Kuhnt 1910: 249</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 253, fig. 14;
<treatmentCitation id="F921109BC22C114CFD7EF893FCB6385D" author="Kuhnt, P." box="[677,843,1892,1916]" page="30" pageId="1" pageNumber="350" year="1911">
<bibRefCitation id="1C114B7BC22C114CFD7EF893FCB6385D" author="Kuhnt, P." box="[677,843,1892,1916]" pageId="1" pageNumber="350" pagination="1 - 103" refId="ref5492" refString="Kuhnt, P. (1911) Family Erotylidae. In: Junk, W. (Ed.), Coleopterorum catalogus, Pars 34 (15), pp. 1 - 103." type="journal article" year="1911">Kuhnt 1911: 30</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F921109BC22C114CFC82F893FBB1385D" author="Blackwelder, R. E." box="[857,1100,1891,1916]" page="462" pageId="1" pageNumber="350" year="1945">
<bibRefCitation id="1C114B7BC22C114CFC82F893FBB1385D" author="Blackwelder, R. E." box="[857,1100,1891,1916]" pageId="1" pageNumber="350" pagination="343 - 550" refId="ref5075" refString="Blackwelder, R. E. (1945) Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Bulletin of the United States National Museum, 185 (3), 343 - 550. https: // doi. org / 10.5479 / si. 03629236.185.3" type="journal article" year="1945">Blackwelder 1945: 462</bibRefCitation>
</treatmentCitation>
, list.
<emphasis id="4AF4EA98C22C114CFB53F893FADC385D" bold="true" box="[1160,1313,1892,1916]" pageId="1" pageNumber="350">New synonym</emphasis>
.
<typeStatus id="A73B8828C22C114CFAEAF893FA64385D" box="[1329,1433,1892,1916]" pageId="1" pageNumber="350" type="lectotype">Lectotype</typeStatus>
, here designated:
<collectingCountry id="0097761AC22C114CFEAAF874FE2138BA" box="[369,476,1923,1947]" name="Venezuela" pageId="1" pageNumber="350">Venezuela</collectingCountry>
,
<collectingRegion id="BA44F868C22C114CFE3EF874FDCF38BA" box="[485,562,1923,1947]" country="Venezuela" name="Merida" pageId="1" pageNumber="350">Mérida</collectingRegion>
:
<collectingRegion id="BA44F868C22C114CFDE1F874FD7E38BA" box="[570,643,1923,1947]" country="Venezuela" name="Merida" pageId="1" pageNumber="350">Mérida</collectingRegion>
(
<geoCoordinate id="1DB4504DC22C114CFD4AF875FCFB38BA" box="[657,774,1922,1947]" degrees="8" direction="north" minutes="35" orientation="latitude" pageId="1" pageNumber="350" precision="15" seconds="53" value="8.598055">8°3553N</geoCoordinate>
,
<geoCoordinate id="1DB4504DC22C114CFCCBF874FC7638BA" box="[784,907,1923,1947]" degrees="71" direction="west" minutes="8" orientation="longitude" pageId="1" pageNumber="350" precision="15" seconds="41" value="-71.14472">71°841W</geoCoordinate>
), 104093, †999 (ZMHB, examined).
<typeStatus id="A73B8828C22C114CFADAF874FA6438BA" box="[1281,1433,1923,1947]" pageId="1" pageNumber="350" type="paralectotype">Paralectotypes</typeStatus>
: Colombia—Starke col.,
<specimenCount id="6E86FD03C22C114CFE64F855FE293898" box="[447,468,1954,1977]" pageId="1" pageNumber="350" type="male"></specimenCount>
, 21678, †971 (ZMHB, examined); †995 (ZMHB, examined); Kuhnt det., 4678, †997 (ZMHB, examined).
</treatmentCitationGroup>
</paragraph>
<paragraph id="783F368AC22C114CFF4CF828FCE938D6" blockId="1.[151,1437,1830,2039]" box="[151,788,2015,2039]" pageId="1" pageNumber="350">
<treatmentCitationGroup id="589011A4C22C114CFF4CF828FCE938D6" box="[151,788,2015,2039]" pageId="1" pageNumber="350">
<taxonomicName id="BF804D09C22C114CFF4CF828FE2538D6" authority="(Kuhnt)" baseAuthorityName="Kuhnt" box="[151,472,2015,2039]" class="Insecta" family="Erotylidae" genus="Oligocorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="4AF4EA98C22C114CFF4CF828FE8338D6" box="[151,382,2015,2039]" italics="true" pageId="1" pageNumber="350">Oligocorynus limbatus</emphasis>
(Kuhnt)
</taxonomicName>
;
<treatmentCitation id="F921109BC22C114CFE3AF828FD4438D6" author="Alvarenga, M." box="[481,697,2015,2039]" page="109" pageId="1" pageNumber="350" year="1994">
<bibRefCitation id="1C114B7BC22C114CFE3AF828FD4438D6" author="Alvarenga, M." box="[481,697,2015,2039]" pageId="1" pageNumber="350" pagination="1 - 175" refId="ref5033" refString="Alvarenga, M. (1994) Catalogo dos Erotylidae (Coleoptera) Neotropicais. Revista Brasileira de Zoologia, 11 (1), 1 - 175. https: // doi. org / 10.1590 / S 0101 - 81751994000100001" type="journal article" year="1994">Alvarenga 1994: 109</bibRefCitation>
</treatmentCitation>
, catalog.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="2CFF6602C22F114FFF4CF86DFD6338DB" ID-DOI="http://doi.org/10.5281/zenodo.3934210" ID-Zenodo-Dep="3934210" httpUri="https://zenodo.org/record/3934210/files/figure.png" pageId="2" pageNumber="351" startId="2.[151,264,1946,1970]" targetBox="[164,1427,184,1906]" targetPageId="2">
<paragraph id="783F368AC22F114FFF4CF86DFD6338DB" blockId="2.[151,1436,1946,2042]" pageId="2" pageNumber="351">
<emphasis id="4AF4EA98C22F114FFF4CF86DFEC63893" bold="true" box="[151,315,1946,1970]" pageId="2" pageNumber="351">FIGURES 18.</emphasis>
<taxonomicName id="BF804D09C22F114FFE99F86DFD7F3893" authority="Crotch, 1876" authorityName="Crotch" authorityYear="1876" box="[322,642,1946,1970]" class="Insecta" family="Erotylidae" genus="Bacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="4AF4EA98C22F114FFE99F86DFE0E3890" box="[322,499,1946,1969]" italics="true" pageId="2" pageNumber="351">Bacis nigropictus</emphasis>
<bibRefCitation id="1C114B7BC22F114FFE20F86DFD7F3893" author="Crotch, G. R." box="[507,642,1946,1970]" pageId="2" pageNumber="351" pagination="377 - 572" refId="ref5170" refString="Crotch, G. R. (1876) A revision of the coleopterous family Erotylidae. Cistula Entomologica, 1, 377 - 572." type="journal article" year="1876">Crotch, 1876</bibRefCitation>
</taxonomicName>
, specimen from Brazil. Habitus: (
<emphasis id="4AF4EA98C22F114FFC07F86DFBFF3893" bold="true" box="[988,1026,1946,1970]" pageId="2" pageNumber="351">12</emphasis>
) dorsal, (
<emphasis id="4AF4EA98C22F114FFBBEF86DFB8F3893" bold="true" box="[1125,1138,1946,1970]" pageId="2" pageNumber="351">3</emphasis>
) lateral, and (
<emphasis id="4AF4EA98C22F114FFAD9F86DFAF23893" bold="true" box="[1282,1295,1946,1970]" pageId="2" pageNumber="351">4</emphasis>
) ventral view (scale bars for 14 = 3 mm); (
<emphasis id="4AF4EA98C22F114FFE1AF849FE3338F7" bold="true" box="[449,462,1982,2006]" pageId="2" pageNumber="351">5</emphasis>
) head (scale bar = 0.3 mm); (
<emphasis id="4AF4EA98C22F114FFD2DF849FCFE38F7" bold="true" box="[758,771,1982,2006]" pageId="2" pageNumber="351">6</emphasis>
) left antenna (scale bar = 1 mm); (
<emphasis id="4AF4EA98C22F114FFB85F849FB9638F7" bold="true" box="[1118,1131,1982,2006]" pageId="2" pageNumber="351">7</emphasis>
) labrum (scale bar = 0.3 mm); (
<emphasis id="4AF4EA98C22F114FFF7BF815FF5038DB" bold="true" box="[160,173,2018,2042]" pageId="2" pageNumber="351">8</emphasis>
) epipharynx (scale bar = 0.3 mm; arrow: torma).
</paragraph>
</caption>
<caption id="2CFF6602C22E114EFF4CF889FC7138FC" ID-DOI="http://doi.org/10.5281/zenodo.3934212" ID-Zenodo-Dep="3934212" httpUri="https://zenodo.org/record/3934212/files/figure.png" pageId="3" pageNumber="352" startId="3.[151,264,1918,1942]" targetBox="[178,1402,212,1826]" targetPageId="3">
<paragraph id="783F368AC22E114EFF4CF889FC7138FC" blockId="3.[151,1437,1918,2014]" pageId="3" pageNumber="352">
<emphasis id="4AF4EA98C22E114EFF4CF889FEBA38B7" bold="true" box="[151,327,1918,1942]" pageId="3" pageNumber="352">FIGURES 916.</emphasis>
<taxonomicName id="BF804D09C22E114EFE97F888FD7738B7" authority="Crotch, 1876" authorityName="Crotch" authorityYear="1876" box="[332,650,1918,1942]" class="Insecta" family="Erotylidae" genus="Bacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="4AF4EA98C22E114EFE97F888FE0138B7" box="[332,508,1919,1942]" italics="true" pageId="3" pageNumber="352">Bacis nigropictus</emphasis>
<bibRefCitation id="1C114B7BC22E114EFDD8F889FD7738B7" author="Crotch, G. R." box="[515,650,1918,1942]" pageId="3" pageNumber="352" pagination="377 - 572" refId="ref5170" refString="Crotch, G. R. (1876) A revision of the coleopterous family Erotylidae. Cistula Entomologica, 1, 377 - 572." type="journal article" year="1876">Crotch, 1876</bibRefCitation>
</taxonomicName>
. (
<emphasis id="4AF4EA98C22E114EFD40F889FD3338B7" bold="true" box="[667,718,1918,1942]" pageId="3" pageNumber="352">910</emphasis>
) Mandibles, dorsal view (arrow: proximal incisor); (
<emphasis id="4AF4EA98C22E114EFB39F889FADD38B7" bold="true" box="[1250,1312,1918,1942]" pageId="3" pageNumber="352">1112</emphasis>
) mandibles, ventral view (arrow: proximal incisor); (
<emphasis id="4AF4EA98C22E114EFDE0F855FDA9389B" bold="true" box="[571,596,1954,1978]" pageId="3" pageNumber="352">13</emphasis>
) maxilla, dorsal view, (
<emphasis id="4AF4EA98C22E114EFC96F855FC9B389B" bold="true" box="[845,870,1954,1978]" pageId="3" pageNumber="352">14</emphasis>
) maxilla, ventral view, (
<emphasis id="4AF4EA98C22E114EFBBDF855FB82389B" bold="true" box="[1126,1151,1954,1978]" pageId="3" pageNumber="352">15</emphasis>
) hypopharynx, dorsal view (arrow: hypopharynx arm); (
<emphasis id="4AF4EA98C22E114EFE61F831FE2E38FF" bold="true" box="[442,467,1990,2014]" pageId="3" pageNumber="352">16</emphasis>
) labium, dorsal view. Scale bars = 0.3 mm.
</paragraph>
</caption>
<subSubSection id="309A6501C2291149FF4CFF6EFD3C3972" pageId="4" pageNumber="353" type="description">
<paragraph id="783F368AC2291149FF4CFF6EFE913E62" blockId="4.[151,1437,153,2016]" pageId="4" pageNumber="353">
<emphasis id="4AF4EA98C2291149FF4CFF6EFEB83F92" bold="true" box="[151,325,153,179]" pageId="4" pageNumber="353">Redescription.</emphasis>
Length:
<quantity id="BF789B6FC2291149FE72FF6EFE163F95" box="[425,491,153,180]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="353" unit="mm" value="9.0">9 mm</quantity>
, thorax/abdomen R=1.6. Body moderately oval, subcordiform, sides slightly curved, moderately convex. Head orange with irregular vertical black band, antennae black, clypeus black apically, orange at base, pronotum black with two orange lateral maculae, mesoventrite, mesanepisterna, mesepimera and metaventrite orange, coxae and femora dark yellow, elytra dark orange with eight black maculae, abdominal ventrites dark orange (
<figureCitation id="E0BB2A0FC2291149FF28FEDEFEA23E62" box="[243,351,297,323]" captionStart="FIGURES 18" captionStartId="2.[151,264,1946,1970]" captionTargetBox="[164,1427,184,1906]" captionTargetId="figure@2.[153,1434,179,1917]" captionTargetPageId="2" captionText="FIGURES 18. Bacis nigropictus Crotch, 1876, specimen from Brazil. Habitus: (12) dorsal, (3) lateral, and (4) ventral view (scale bars for 14 = 3 mm); (5) head (scale bar = 0.3 mm); (6) left antenna (scale bar = 1 mm); (7) labrum (scale bar = 0.3 mm); (8) epipharynx (scale bar = 0.3 mm; arrow: torma)." figureDoi="http://doi.org/10.5281/zenodo.3934210" httpUri="https://zenodo.org/record/3934210/files/figure.png" pageId="4" pageNumber="353">Figs. 14</figureCitation>
).
</paragraph>
<paragraph id="783F368AC2291149FF1CFEBAFEBA3C41" blockId="4.[151,1437,153,2016]" pageId="4" pageNumber="353">
<emphasis id="4AF4EA98C2291149FF1CFEBAFEFA3E46" bold="true" box="[199,263,333,359]" pageId="4" pageNumber="353">Head</emphasis>
(R=0.8): ocular striae restricted to eye margin, eyes faceted, interocular distance ~2/3 of head width (
<figureCitation id="E0BB2A0FC2291149FAB4FEBAFF5A3EAD" captionStart="FIGURES 18" captionStartId="2.[151,264,1946,1970]" captionTargetBox="[164,1427,184,1906]" captionTargetId="figure@2.[153,1434,179,1917]" captionTargetPageId="2" captionText="FIGURES 18. Bacis nigropictus Crotch, 1876, specimen from Brazil. Habitus: (12) dorsal, (3) lateral, and (4) ventral view (scale bars for 14 = 3 mm); (5) head (scale bar = 0.3 mm); (6) left antenna (scale bar = 1 mm); (7) labrum (scale bar = 0.3 mm); (8) epipharynx (scale bar = 0.3 mm; arrow: torma)." figureDoi="http://doi.org/10.5281/zenodo.3934210" httpUri="https://zenodo.org/record/3934210/files/figure.png" pageId="4" pageNumber="353">Fig. 5</figureCitation>
). Stridulatory organs absent at base of head in both sexes. Antennae (R=11.6): antennomeres VIII and XI elongate (
<figureCitation id="E0BB2A0FC2291149FF7BFE62FF193E91" box="[160,228,405,432]" captionStart="FIGURES 18" captionStartId="2.[151,264,1946,1970]" captionTargetBox="[164,1427,184,1906]" captionTargetId="figure@2.[153,1434,179,1917]" captionTargetPageId="2" captionText="FIGURES 18. Bacis nigropictus Crotch, 1876, specimen from Brazil. Habitus: (12) dorsal, (3) lateral, and (4) ventral view (scale bars for 14 = 3 mm); (5) head (scale bar = 0.3 mm); (6) left antenna (scale bar = 1 mm); (7) labrum (scale bar = 0.3 mm); (8) epipharynx (scale bar = 0.3 mm; arrow: torma)." figureDoi="http://doi.org/10.5281/zenodo.3934210" httpUri="https://zenodo.org/record/3934210/files/figure.png" pageId="4" pageNumber="353">Fig. 6</figureCitation>
). Clypeus (R=0.4) sub-rectangular, apex truncate (
<figureCitation id="E0BB2A0FC2291149FCC4FE62FC993E91" box="[799,868,405,432]" captionStart="FIGURES 18" captionStartId="2.[151,264,1946,1970]" captionTargetBox="[164,1427,184,1906]" captionTargetId="figure@2.[153,1434,179,1917]" captionTargetPageId="2" captionText="FIGURES 18. Bacis nigropictus Crotch, 1876, specimen from Brazil. Habitus: (12) dorsal, (3) lateral, and (4) ventral view (scale bars for 14 = 3 mm); (5) head (scale bar = 0.3 mm); (6) left antenna (scale bar = 1 mm); (7) labrum (scale bar = 0.3 mm); (8) epipharynx (scale bar = 0.3 mm; arrow: torma)." figureDoi="http://doi.org/10.5281/zenodo.3934210" httpUri="https://zenodo.org/record/3934210/files/figure.png" pageId="4" pageNumber="353">Fig. 5</figureCitation>
). Labrum (R=0.7): sub-rectangular, without apical elevation, absence of membranous cover at proximal half (
<figureCitation id="E0BB2A0FC2291149FCEBFE4EFC8B3EF5" box="[816,886,441,468]" captionStart="FIGURES 18" captionStartId="2.[151,264,1946,1970]" captionTargetBox="[164,1427,184,1906]" captionTargetId="figure@2.[153,1434,179,1917]" captionTargetPageId="2" captionText="FIGURES 18. Bacis nigropictus Crotch, 1876, specimen from Brazil. Habitus: (12) dorsal, (3) lateral, and (4) ventral view (scale bars for 14 = 3 mm); (5) head (scale bar = 0.3 mm); (6) left antenna (scale bar = 1 mm); (7) labrum (scale bar = 0.3 mm); (8) epipharynx (scale bar = 0.3 mm; arrow: torma)." figureDoi="http://doi.org/10.5281/zenodo.3934210" httpUri="https://zenodo.org/record/3934210/files/figure.png" pageId="4" pageNumber="353">Fig. 7</figureCitation>
). Epipharynx with setae in median region and at anterior margin, median region covered with microtrichiae, tormae with anterior projection reaching the basal third of labrum, posterior projection of tormae almost as long as the anterior projection (
<figureCitation id="E0BB2A0FC2291149FBFCFDF6FB943D3A" box="[1063,1129,513,539]" captionStart="FIGURES 18" captionStartId="2.[151,264,1946,1970]" captionTargetBox="[164,1427,184,1906]" captionTargetId="figure@2.[153,1434,179,1917]" captionTargetPageId="2" captionText="FIGURES 18. Bacis nigropictus Crotch, 1876, specimen from Brazil. Habitus: (12) dorsal, (3) lateral, and (4) ventral view (scale bars for 14 = 3 mm); (5) head (scale bar = 0.3 mm); (6) left antenna (scale bar = 1 mm); (7) labrum (scale bar = 0.3 mm); (8) epipharynx (scale bar = 0.3 mm; arrow: torma)." figureDoi="http://doi.org/10.5281/zenodo.3934210" httpUri="https://zenodo.org/record/3934210/files/figure.png" pageId="4" pageNumber="353">Fig. 8</figureCitation>
, arrow). Mandibles slightly asymmetrical, with three incisors, membranous lobe covered with microtrichiae on molar region, with two carinae on dorsal region, ventral cavity with internal incisor margin sinuate, left mandible with proximal incisor smaller and attached to middle incisor as a lobe (
<figureCitation id="E0BB2A0FC2291149FDE3FD9AFD483DA6" box="[568,693,621,647]" captionStart="FIGURES 916" captionStartId="3.[151,264,1918,1942]" captionTargetBox="[178,1402,212,1826]" captionTargetId="figure@3.[166,1422,178,1892]" captionTargetPageId="3" captionText="FIGURES 916. Bacis nigropictus Crotch, 1876. (910) Mandibles, dorsal view (arrow: proximal incisor); (1112) mandibles, ventral view (arrow: proximal incisor); (13) maxilla, dorsal view, (14) maxilla, ventral view, (15) hypopharynx, dorsal view (arrow: hypopharynx arm); (16) labium, dorsal view. Scale bars = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3934212" httpUri="https://zenodo.org/record/3934212/files/figure.png" pageId="4" pageNumber="353">Figs. 912</figureCitation>
). Maxillae (R=2.7): lacinia with an apical curved hook, with long setae denser distally, palpomere I elongate, palpomeres IIIII as wide as long, palpomere IV wide, galea (R=1.9) subfalciform, with moderately long setae denser distally (
<figureCitation id="E0BB2A0FC2291149FCC9FD42FC673DEE" box="[786,922,693,719]" captionStart="FIGURES 916" captionStartId="3.[151,264,1918,1942]" captionTargetBox="[178,1402,212,1826]" captionTargetId="figure@3.[166,1422,178,1892]" captionTargetPageId="3" captionText="FIGURES 916. Bacis nigropictus Crotch, 1876. (910) Mandibles, dorsal view (arrow: proximal incisor); (1112) mandibles, ventral view (arrow: proximal incisor); (13) maxilla, dorsal view, (14) maxilla, ventral view, (15) hypopharynx, dorsal view (arrow: hypopharynx arm); (16) labium, dorsal view. Scale bars = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3934212" httpUri="https://zenodo.org/record/3934212/files/figure.png" pageId="4" pageNumber="353">Figs. 1314</figureCitation>
). Hypopharynx with two basal arms extending to posterior region of mentum (
<figureCitation id="E0BB2A0FC2291149FE21FD2EFDB03DD5" box="[506,589,729,756]" captionStart="FIGURES 916" captionStartId="3.[151,264,1918,1942]" captionTargetBox="[178,1402,212,1826]" captionTargetId="figure@3.[166,1422,178,1892]" captionTargetPageId="3" captionText="FIGURES 916. Bacis nigropictus Crotch, 1876. (910) Mandibles, dorsal view (arrow: proximal incisor); (1112) mandibles, ventral view (arrow: proximal incisor); (13) maxilla, dorsal view, (14) maxilla, ventral view, (15) hypopharynx, dorsal view (arrow: hypopharynx arm); (16) labium, dorsal view. Scale bars = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3934212" httpUri="https://zenodo.org/record/3934212/files/figure.png" pageId="4" pageNumber="353">Fig. 15</figureCitation>
, arrow). Labium (R=1.5): ligula long (~3/4 width of the mentum), fused at middle, with a basal sclerite extending through almost the entire ligula, outer angles rounded, basal palpomere with moderately curved base, inner angle of distal margin slightly narrow, mentum with medial region without definite shape (
<figureCitation id="E0BB2A0FC2291149FF3DFCB2FEC73C41" box="[230,314,837,864]" captionStart="FIGURES 916" captionStartId="3.[151,264,1918,1942]" captionTargetBox="[178,1402,212,1826]" captionTargetId="figure@3.[166,1422,178,1892]" captionTargetPageId="3" captionText="FIGURES 916. Bacis nigropictus Crotch, 1876. (910) Mandibles, dorsal view (arrow: proximal incisor); (1112) mandibles, ventral view (arrow: proximal incisor); (13) maxilla, dorsal view, (14) maxilla, ventral view, (15) hypopharynx, dorsal view (arrow: hypopharynx arm); (16) labium, dorsal view. Scale bars = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3934212" httpUri="https://zenodo.org/record/3934212/files/figure.png" pageId="4" pageNumber="353">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="783F368AC2291149FF1CFC9EFB113B16" blockId="4.[151,1437,153,2016]" pageId="4" pageNumber="353">
<emphasis id="4AF4EA98C2291149FF1CFC9EFED43CA2" bold="true" box="[199,297,873,899]" pageId="4" pageNumber="353">Thorax:</emphasis>
pronotum (R=0.4) glossy, lateral region of posterior margin with a depression, surface of depression punctate, prosternum with few short setae, procoxal lines absent. Scutellar shield (R=0.7) semiovoid. Metaventrite (R=0.4): mesocoxal lines conspicuous. Legs: metathoracic legs with femora as wide as the prothoracic legs. Elytra (R=2.4): surface with fine punctures (φ~
<quantity id="BF789B6FC2291149FDB7FC22FD243CD1" box="[620,729,981,1008]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="353" unit="mm" value="0.05">0.05 mm</quantity>
), with elytral striae moderately conspicuous, geminate; interstriae with punctures moderately conspicuous. Metathoracic wings (R=2.3): with two axillary veins, veins AA
<subScript id="E40434CFC2291149FA58FBFDFA603B39" attach="right" box="[1411,1437,1034,1048]" fontSize="6" pageId="4" pageNumber="353">3+4</subScript>
reaching CuA
<subScript id="E40434CFC2291149FEE8FBD9FEB03B1D" attach="left" box="[307,333,1070,1084]" fontSize="6" pageId="4" pageNumber="353">3+4</subScript>
, veins MP
<subScript id="E40434CFC2291149FE1DFBD9FE223B1D" attach="left" box="[454,479,1070,1084]" fontSize="6" pageId="4" pageNumber="353">3+4</subScript>
reaching MP
<subScript id="E40434CFC2291149FDADFBD9FD723B1D" attach="left" box="[630,655,1070,1084]" fontSize="6" pageId="4" pageNumber="353">1+2</subScript>
, vein cua1-mp4 complete, vein r4 complete (
<figureCitation id="E0BB2A0FC2291149FB57FBEAFB223B16" box="[1164,1247,1053,1079]" captionStart="FIGURES 1725" captionStartId="5.[151,264,1912,1936]" captionTargetBox="[262,1300,195,1849]" captionTargetId="figure@5.[201,1387,181,1886]" captionTargetPageId="5" captionText="FIGURES 1725. Bacis nigropictus Crotch, 1876. (17) Right metathoracic wing (scale bar = 3 mm); (18) eighth abdominal tergite; (19) eighth abdominal sternite; (20) tenth abdominal tergite; (21) ninth abdominal sternite (scale bars for 1821 = 0.3 mm); (2224) aedeagus, dorsal, ventral, and lateral view, respectively (scale bar = 0.5 mm); (25) head of flagellum (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3934214" httpUri="https://zenodo.org/record/3934214/files/figure.png" pageId="4" pageNumber="353">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="783F368AC2291149FF1CFBB6FDF5392A" blockId="4.[151,1437,153,2016]" pageId="4" pageNumber="353">
<emphasis id="4AF4EA98C2291149FF1CFBB6FEB83B7A" bold="true" box="[199,325,1089,1115]" pageId="4" pageNumber="353">Abdomen:</emphasis>
surface with few short setae. Male genital segments and genitalia: tergite VIII (R=0.6) subtriangular, uniformly sclerotized, base curved, setae denser distally (
<figureCitation id="E0BB2A0FC2291149FCE1FB92FC703B5E" box="[826,909,1125,1151]" captionStart="FIGURES 1725" captionStartId="5.[151,264,1912,1936]" captionTargetBox="[262,1300,195,1849]" captionTargetId="figure@5.[201,1387,181,1886]" captionTargetPageId="5" captionText="FIGURES 1725. Bacis nigropictus Crotch, 1876. (17) Right metathoracic wing (scale bar = 3 mm); (18) eighth abdominal tergite; (19) eighth abdominal sternite; (20) tenth abdominal tergite; (21) ninth abdominal sternite (scale bars for 1821 = 0.3 mm); (2224) aedeagus, dorsal, ventral, and lateral view, respectively (scale bar = 0.5 mm); (25) head of flagellum (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3934214" httpUri="https://zenodo.org/record/3934214/files/figure.png" pageId="4" pageNumber="353">Fig. 18</figureCitation>
); sternite VIII (R=0.3) transverse, distal margin emarginate, setae denser distally (
<figureCitation id="E0BB2A0FC2291149FDCBFB7EFD9F3B82" box="[528,610,1161,1187]" captionStart="FIGURES 1725" captionStartId="5.[151,264,1912,1936]" captionTargetBox="[262,1300,195,1849]" captionTargetId="figure@5.[201,1387,181,1886]" captionTargetPageId="5" captionText="FIGURES 1725. Bacis nigropictus Crotch, 1876. (17) Right metathoracic wing (scale bar = 3 mm); (18) eighth abdominal tergite; (19) eighth abdominal sternite; (20) tenth abdominal tergite; (21) ninth abdominal sternite (scale bars for 1821 = 0.3 mm); (2224) aedeagus, dorsal, ventral, and lateral view, respectively (scale bar = 0.5 mm); (25) head of flagellum (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3934214" httpUri="https://zenodo.org/record/3934214/files/figure.png" pageId="4" pageNumber="353">Fig. 19</figureCitation>
); tergite X (R=0.8) U-shaped; lateral shafts slightly curved, apex truncate, setae denser distally (
<figureCitation id="E0BB2A0FC2291149FE5CFB5AFE263BE6" box="[391,475,1197,1223]" captionStart="FIGURES 1725" captionStartId="5.[151,264,1912,1936]" captionTargetBox="[262,1300,195,1849]" captionTargetId="figure@5.[201,1387,181,1886]" captionTargetPageId="5" captionText="FIGURES 1725. Bacis nigropictus Crotch, 1876. (17) Right metathoracic wing (scale bar = 3 mm); (18) eighth abdominal tergite; (19) eighth abdominal sternite; (20) tenth abdominal tergite; (21) ninth abdominal sternite (scale bars for 1821 = 0.3 mm); (2224) aedeagus, dorsal, ventral, and lateral view, respectively (scale bar = 0.5 mm); (25) head of flagellum (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3934214" httpUri="https://zenodo.org/record/3934214/files/figure.png" pageId="4" pageNumber="353">Fig. 20</figureCitation>
); laterotergite IX (R=1.4) elongate, asymmetrical, lateral lobes sub-triangular; sternite IX setae denser distally (
<figureCitation id="E0BB2A0FC2291149FE72FB26FE063BCA" box="[425,507,1233,1259]" captionStart="FIGURES 1725" captionStartId="5.[151,264,1912,1936]" captionTargetBox="[262,1300,195,1849]" captionTargetId="figure@5.[201,1387,181,1886]" captionTargetPageId="5" captionText="FIGURES 1725. Bacis nigropictus Crotch, 1876. (17) Right metathoracic wing (scale bar = 3 mm); (18) eighth abdominal tergite; (19) eighth abdominal sternite; (20) tenth abdominal tergite; (21) ninth abdominal sternite (scale bars for 1821 = 0.3 mm); (2224) aedeagus, dorsal, ventral, and lateral view, respectively (scale bar = 0.5 mm); (25) head of flagellum (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3934214" httpUri="https://zenodo.org/record/3934214/files/figure.png" pageId="4" pageNumber="353">Fig. 21</figureCitation>
). Aedeagus: tegmen elongate at middle, dorsal longitudinal line sclerotized, tegmen arm long, with two sclerotized lines, lateral lobes width ~1/8 of tegmen length, setae denser distally, lateral region of median lobe narrow, internal sac short (
<figureCitation id="E0BB2A0FC2291149FDBCFAEEFD0D3A12" box="[615,752,1305,1331]" captionStart="FIGURES 1725" captionStartId="5.[151,264,1912,1936]" captionTargetBox="[262,1300,195,1849]" captionTargetId="figure@5.[201,1387,181,1886]" captionTargetPageId="5" captionText="FIGURES 1725. Bacis nigropictus Crotch, 1876. (17) Right metathoracic wing (scale bar = 3 mm); (18) eighth abdominal tergite; (19) eighth abdominal sternite; (20) tenth abdominal tergite; (21) ninth abdominal sternite (scale bars for 1821 = 0.3 mm); (2224) aedeagus, dorsal, ventral, and lateral view, respectively (scale bar = 0.5 mm); (25) head of flagellum (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3934214" httpUri="https://zenodo.org/record/3934214/files/figure.png" pageId="4" pageNumber="353">Figs. 2224</figureCitation>
); head of flagellum subtriangular (
<figureCitation id="E0BB2A0FC2291149FBB6FAEEFB433A15" box="[1133,1214,1305,1332]" captionStart="FIGURES 1725" captionStartId="5.[151,264,1912,1936]" captionTargetBox="[262,1300,195,1849]" captionTargetId="figure@5.[201,1387,181,1886]" captionTargetPageId="5" captionText="FIGURES 1725. Bacis nigropictus Crotch, 1876. (17) Right metathoracic wing (scale bar = 3 mm); (18) eighth abdominal tergite; (19) eighth abdominal sternite; (20) tenth abdominal tergite; (21) ninth abdominal sternite (scale bars for 1821 = 0.3 mm); (2224) aedeagus, dorsal, ventral, and lateral view, respectively (scale bar = 0.5 mm); (25) head of flagellum (scale bar = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.3934214" httpUri="https://zenodo.org/record/3934214/files/figure.png" pageId="4" pageNumber="353">Fig. 25</figureCitation>
, MAAEIS). Female genital segments and genitalia: tergite VIII (R=0.5) transverse, setae denser distally (
<figureCitation id="E0BB2A0FC2291149FB97FACAFB5D3A79" box="[1100,1184,1341,1368]" captionStart="FIGURES 2629" captionStartId="6.[151,264,1515,1539]" captionTargetBox="[216,1296,329,1451]" captionTargetId="figure@6.[151,1436,306,1492]" captionTargetPageId="6" captionText="FIGURES 2629. Bacis nigropictus Crotch, 1876. (26) Eighth abdominal tergite; (27) eighth abdominal sternite; (28) ovipositor, dorsal view (arrow: proctiger lobe); (29) ovipositor, ventral view. Scale bars = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3934216" httpUri="https://zenodo.org/record/3934216/files/figure.png" pageId="4" pageNumber="353">Fig. 26</figureCitation>
); sternite VIII (R=0.6) transverse, setae denser distally, median strut approximately three times longer than base (
<figureCitation id="E0BB2A0FC2291149FB4EFA96FB173A5A" box="[1173,1258,1377,1403]" captionStart="FIGURES 2629" captionStartId="6.[151,264,1515,1539]" captionTargetBox="[216,1296,329,1451]" captionTargetId="figure@6.[151,1436,306,1492]" captionTargetPageId="6" captionText="FIGURES 2629. Bacis nigropictus Crotch, 1876. (26) Eighth abdominal tergite; (27) eighth abdominal sternite; (28) ovipositor, dorsal view (arrow: proctiger lobe); (29) ovipositor, ventral view. Scale bars = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3934216" httpUri="https://zenodo.org/record/3934216/files/figure.png" pageId="4" pageNumber="353">Fig. 27</figureCitation>
); tergite IX and sternite IX indistinct, membranous, surface with microtrichiae uniformly distributed. Ovipositor: proctiger lobe long (
<figureCitation id="E0BB2A0FC2291149FF02FA5EFE9F3AE2" box="[217,354,1449,1475]" captionStart="FIGURES 2629" captionStartId="6.[151,264,1515,1539]" captionTargetBox="[216,1296,329,1451]" captionTargetId="figure@6.[151,1436,306,1492]" captionTargetPageId="6" captionText="FIGURES 2629. Bacis nigropictus Crotch, 1876. (26) Eighth abdominal tergite; (27) eighth abdominal sternite; (28) ovipositor, dorsal view (arrow: proctiger lobe); (29) ovipositor, ventral view. Scale bars = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3934216" httpUri="https://zenodo.org/record/3934216/files/figure.png" pageId="4" pageNumber="353">Figs. 2829</figureCitation>
, arrow), covering the vulval lobe, subvulval lobe as wide as the basal segments of the gonocoxites, gonocoxites ~1/2 of ovipositor length with narrowed and rounded apex, gonostyli setae moderately long (
<figureCitation id="E0BB2A0FC2291149FABFFA3AFF1F392A" captionStart="FIGURES 2629" captionStartId="6.[151,264,1515,1539]" captionTargetBox="[216,1296,329,1451]" captionTargetId="figure@6.[151,1436,306,1492]" captionTargetPageId="6" captionText="FIGURES 2629. Bacis nigropictus Crotch, 1876. (26) Eighth abdominal tergite; (27) eighth abdominal sternite; (28) ovipositor, dorsal view (arrow: proctiger lobe); (29) ovipositor, ventral view. Scale bars = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3934216" httpUri="https://zenodo.org/record/3934216/files/figure.png" pageId="4" pageNumber="353">Figs. 2829</figureCitation>
). Spermatheca ellipsoidal.
</paragraph>
<paragraph id="783F368AC2291149FF1CF9E2FD3C3972" blockId="4.[151,1437,153,2016]" pageId="4" pageNumber="353">
<emphasis id="4AF4EA98C2291149FF1CF9E2FE25390E" bold="true" box="[199,472,1557,1583]" pageId="4" pageNumber="353">Intraspecific variation.</emphasis>
Integument color from light-orange to light-brown, with bands, legs and sternites dark brown; inner spots of elytra can be fused in pairs.
</paragraph>
</subSubSection>
<subSubSection id="309A6501C2291149FF1CF9AAFACD39BD" pageId="4" pageNumber="353" type="diagnosis">
<paragraph id="783F368AC2291149FF1CF9AAFACD39BD" blockId="4.[151,1437,153,2016]" pageId="4" pageNumber="353">
<emphasis id="4AF4EA98C2291149FF1CF9AAFEBF3956" bold="true" box="[199,322,1629,1655]" pageId="4" pageNumber="353">Diagnosis.</emphasis>
<taxonomicName id="BF804D09C2291149FE90F9A9FDEF3956" authorityName="Crotch" authorityYear="1876" box="[331,530,1630,1655]" class="Insecta" family="Erotylidae" genus="Bacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="4AF4EA98C2291149FE90F9A9FDEF3956" box="[331,530,1630,1655]" italics="true" pageId="4" pageNumber="353">Bacis nigropictus</emphasis>
</taxonomicName>
can be recognized among other species of
<taxonomicName id="BF804D09C2291149FBD2F9A9FBBA3956" authorityName="Dejean" authorityYear="1836" box="[1033,1095,1630,1655]" class="Insecta" family="Erotylidae" genus="Bacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="4AF4EA98C2291149FBD2F9A9FBBA3956" box="[1033,1095,1630,1655]" italics="true" pageId="4" pageNumber="353">Bacis</emphasis>
</taxonomicName>
by its dorsal coloration, with eight black elytral spots and pronotum orange with black longitudinal medial band, and clypeus truncate.
</paragraph>
</subSubSection>
<subSubSection id="309A6501C2291149FF1CF952FAF239C2" pageId="4" pageNumber="353" type="etymology">
<paragraph id="783F368AC2291149FF1CF952FAF239C2" blockId="4.[151,1437,153,2016]" pageId="4" pageNumber="353">
<emphasis id="4AF4EA98C2291149FF1CF952FEB3399E" bold="true" box="[199,334,1701,1727]" pageId="4" pageNumber="353">Etymology.</emphasis>
The genus name is derived from the Greek
<emphasis id="4AF4EA98C2291149FCE8F952FC93399E" box="[819,878,1701,1727]" italics="true" pageId="4" pageNumber="353">bacis</emphasis>
, “soothsayer”; the specific epithet is derived from the Latin
<emphasis id="4AF4EA98C2291149FED9F93DFEC339C2" box="[258,318,1738,1763]" italics="true" pageId="4" pageNumber="353">nigro</emphasis>
, “black” and
<emphasis id="4AF4EA98C2291149FE03F93DFDE639C2" box="[472,539,1738,1763]" italics="true" pageId="4" pageNumber="353">pictus</emphasis>
, “painted, colored”, referring to the elytral and pronotal coloration.
</paragraph>
</subSubSection>
<subSubSection id="309A6501C2291149FF1CF91AFC42386E" pageId="4" pageNumber="353" type="materials_examined">
<paragraph id="783F368AC2291149FF1CF91AFDA8380A" blockId="4.[151,1437,153,2016]" pageId="4" pageNumber="353">
<emphasis id="4AF4EA98C2291149FF1CF91AFE523826" bold="true" box="[199,431,1773,1799]" pageId="4" pageNumber="353">Material examined.</emphasis>
<materialsCitation id="C8E83CD7C2291149FE6FF91AFB113826" ID-GBIF-Occurrence="2805034302" box="[436,1260,1773,1800]" collectionCode="SMTD" collectorName="Venezuela-ex-Coll. C. Felsche" pageId="4" pageNumber="353" specimenCount="2">
<collectorName id="D575535CC2291149FE6FF91AFCE63826" box="[436,795,1773,1799]" pageId="4" pageNumber="353">Venezuela—ex-Coll. C. Felsche</collectorName>
, donation 1907,
<specimenCount id="6E86FD03C2291149FC09F91AFBF73829" box="[978,1034,1773,1800]" pageId="4" pageNumber="353" type="generic">2 ex.</specimenCount>
: †846, 845 (
<collectionCode id="1E91AE4FC2291149FB4AF91AFB183826" box="[1169,1253,1773,1799]" pageId="4" pageNumber="353">SMTD</collectionCode>
)
</materialsCitation>
; †843 (
<collectionCode id="1E91AE4FC2291149FAE7F91AFA6D3826" box="[1340,1424,1773,1799]" pageId="4" pageNumber="353">SMTD</collectionCode>
);
<materialsCitation id="C8E83CD7C2291149FF4CF8E6FDAC380A" ID-GBIF-Occurrence="2805034301" box="[151,593,1809,1836]" collectionCode="SMTD" pageId="4" pageNumber="353" specimenCount="1" specimenCount-male="1">
Brazil—
<specimenCount id="6E86FD03C2291149FF26F8E6FEED380A" box="[253,272,1809,1835]" pageId="4" pageNumber="353" type="male"></specimenCount>
, Mus antiqu., †844 (
<collectionCode id="1E91AE4FC2291149FE2EF8E6FDB4380A" box="[501,585,1809,1835]" pageId="4" pageNumber="353">SMTD</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="783F368AC2291149FF1CF8C2FC42386E" blockId="4.[151,1437,153,2016]" box="[199,959,1845,1872]" pageId="4" pageNumber="353">
<emphasis id="4AF4EA98C2291149FF1CF8C2FDF8386E" bold="true" box="[199,517,1845,1871]" pageId="4" pageNumber="353">Geographical distribution.</emphasis>
<collectingCountry id="0097761AC2291149FDD7F8C2FD80386E" box="[524,637,1845,1871]" name="Colombia" pageId="4" pageNumber="353">Colombia</collectingCountry>
,
<collectingCountry id="0097761AC2291149FD5CF8C2FD07386E" box="[647,762,1845,1871]" name="Venezuela" pageId="4" pageNumber="353">Venezuela</collectingCountry>
(
<collectingRegion id="BA44F868C2291149FCD2F8C2FC9D3871" box="[777,864,1845,1872]" country="Venezuela" name="Merida" pageId="4" pageNumber="353">Mérida</collectingRegion>
),
<collectingCountry id="0097761AC2291149FCA8F8C2FC46386E" box="[883,955,1845,1871]" name="Brazil" pageId="4" pageNumber="353">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="309A6501C229114BFF1CF8AEFECB3E01" lastPageId="6" lastPageNumber="355" pageId="4" pageNumber="353" type="discussion">
<paragraph id="783F368AC229114BFF1CF8AEFECB3E01" blockId="4.[151,1437,153,2016]" lastBlockId="6.[151,1437,153,288]" lastPageId="6" lastPageNumber="355" pageId="4" pageNumber="353">
<emphasis id="4AF4EA98C2291149FF1CF8AEFEC63852" bold="true" box="[199,315,1881,1907]" pageId="4" pageNumber="353">Remarks.</emphasis>
Although the type series of
<taxonomicName id="BF804D09C2291149FDA8F8ADFCEC3852" authorityName="Crotch" authorityYear="1876" box="[627,785,1882,1907]" class="Insecta" family="Erotylidae" genus="Bacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="4AF4EA98C2291149FDA8F8ADFCEC3852" box="[627,785,1882,1907]" italics="true" pageId="4" pageNumber="353">B. nigropictus</emphasis>
</taxonomicName>
has not been examined, Crotchs original description indicates that
<taxonomicName id="BF804D09C2291149FEDEF889FE5F38B6" authorityName="Crotch" authorityYear="1876" box="[261,418,1918,1943]" class="Insecta" family="Erotylidae" genus="Bacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="4AF4EA98C2291149FEDEF889FE5F38B6" box="[261,418,1918,1943]" italics="true" pageId="4" pageNumber="353">B. nigropictus</emphasis>
</taxonomicName>
and
<taxonomicName id="BF804D09C2291149FE0DF88AFD2938B6" box="[470,724,1917,1943]" class="Insecta" family="Erotylidae" genus="Oligocorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="4AF4EA98C2291149FE0DF88AFD2938B6" box="[470,724,1917,1943]" italics="true" pageId="4" pageNumber="353">Oligocorynus limbatus</emphasis>
</taxonomicName>
are the same species. By comparing specimens of
<taxonomicName id="BF804D09C2291149FADFF88AFA6138B6" authorityName="Dejean" authorityYear="1836" box="[1284,1436,1917,1943]" class="Insecta" family="Erotylidae" genus="Oligocorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="4AF4EA98C2291149FADFF88AFA6138B6" box="[1284,1436,1917,1943]" italics="true" pageId="4" pageNumber="353">Oligocorynus</emphasis>
</taxonomicName>
to specimens of
<taxonomicName id="BF804D09C2291149FE91F855FE1A389A" authorityName="Crotch" authorityYear="1876" box="[330,487,1954,1979]" class="Insecta" family="Erotylidae" genus="Bacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="4AF4EA98C2291149FE91F855FE1A389A" box="[330,487,1954,1979]" italics="true" pageId="4" pageNumber="353">B. nigropictus</emphasis>
</taxonomicName>
, it is evident that the latter have an increased and more gradual curvature of the elytral sides (“
<emphasis id="4AF4EA98C2291149FF35F832FE6538FE" box="[238,408,1989,2015]" italics="true" pageId="4" pageNumber="353">breviter ovatus</emphasis>
”, as described by
<bibRefCitation id="1C114B7BC2291149FDB4F832FCEF38FE" author="Crotch, G. R." box="[623,786,1988,2015]" pageId="4" pageNumber="353" pagination="377 - 572" refId="ref5170" refString="Crotch, G. R. (1876) A revision of the coleopterous family Erotylidae. Cistula Entomologica, 1, 377 - 572." type="journal article" year="1876">Crotch [1876]</bibRefCitation>
), a trait typical of the genus
<taxonomicName id="BF804D09C2291149FBBAF831FB6238FE" authorityName="Dejean" authorityYear="1836" box="[1121,1183,1990,2015]" class="Insecta" family="Erotylidae" genus="Bacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="4AF4EA98C2291149FBBAF831FB6238FE" box="[1121,1183,1990,2015]" italics="true" pageId="4" pageNumber="353">Bacis</emphasis>
</taxonomicName>
, not of
<taxonomicName id="BF804D09C2291149FB26F832FA6838FE" authorityName="Dejean" authorityYear="1836" box="[1277,1429,1989,2015]" class="Insecta" family="Erotylidae" genus="Oligocorynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="4AF4EA98C2291149FB26F832FA6838FE" box="[1277,1429,1989,2015]" italics="true" pageId="4" pageNumber="353">Oligocorynus</emphasis>
</taxonomicName>
. In any case, the diagnosis presented above places this species among those currently included in
<taxonomicName id="BF804D09C22B114BFB3CFF6DFAD83F92" authorityName="Dejean" authorityYear="1836" box="[1255,1317,154,179]" class="Insecta" family="Erotylidae" genus="Bacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="355" phylum="Arthropoda" rank="genus">
<emphasis id="4AF4EA98C22B114BFB3CFF6DFAD83F92" box="[1255,1317,154,179]" italics="true" pageId="6" pageNumber="355">Bacis</emphasis>
</taxonomicName>
. The Venezuelan
<typeStatus id="A73B8828C22B114BFF2FFF49FEB63FF9" box="[244,331,190,216]" pageId="6" pageNumber="355" type="syntype">syntype</typeStatus>
of
<taxonomicName id="BF804D09C22B114BFEA9FF49FDC23FF6" authorityName="Kuhnt" authorityYear="1910" box="[370,575,189,215]" class="Insecta" family="Erotylidae" genus="Zonarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="355" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="4AF4EA98C22B114BFEA9FF49FDC23FF6" box="[370,575,189,215]" italics="true" pageId="6" pageNumber="355">Zonarius limbatus</emphasis>
</taxonomicName>
is here chosen as the
<typeStatus id="A73B8828C22B114BFCE3FF4AFC5F3FF6" box="[824,930,189,215]" pageId="6" pageNumber="355" type="lectotype">lectotype</typeStatus>
, because it came from the same type locality as
<taxonomicName id="BF804D09C22B114BFF6CFF15FE803FDA" authorityName="Crotch" authorityYear="1876" box="[183,381,226,251]" class="Insecta" family="Erotylidae" genus="Bacis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="355" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="4AF4EA98C22B114BFF6CFF15FE803FDA" box="[183,381,226,251]" italics="true" pageId="6" pageNumber="355">Bacis nigropictus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="1C114B7BC22B114BFE67FF16FDB03FDD" author="Crotch, G. R." box="[444,589,225,252]" pageId="6" pageNumber="355" pagination="377 - 572" refId="ref5170" refString="Crotch, G. R. (1876) A revision of the coleopterous family Erotylidae. Cistula Entomologica, 1, 377 - 572." type="journal article" year="1876">Crotch 1876</bibRefCitation>
). The Colombian
<typeStatus id="A73B8828C22B114BFCC0FF15FC803FDD" box="[795,893,226,252]" pageId="6" pageNumber="355" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="BF804D09C22B114BFC7DFF15FBD83FDA" authorityName="Kuhnt" authorityYear="1910" box="[934,1061,225,252]" class="Insecta" family="Erotylidae" genus="Zonarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="355" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="4AF4EA98C22B114BFC7DFF15FC4B3FDA" box="[934,950,226,251]" italics="true" pageId="6" pageNumber="355">Z</emphasis>
.
<emphasis id="4AF4EA98C22B114BFC1EFF16FBD83FDA" box="[965,1061,225,251]" italics="true" pageId="6" pageNumber="355">limbatus</emphasis>
</taxonomicName>
are designated as
<typeStatus id="A73B8828C22B114BFB20FF16FA603FDA" box="[1275,1437,225,251]" pageId="6" pageNumber="355" type="paralectotype">paralectotypes</typeStatus>
(
<figureCitation id="E0BB2A0FC22B114BFF44FEF2FED73E01" box="[159,298,261,288]" captionStart="FIGURES 5760" captionStartId="12.[151,264,1673,1697]" captionTargetBox="[158,1431,185,1636]" captionTargetId="figure@12.[151,1436,181,1647]" captionTargetPageId="12" captionText="FIGURES 5760. Type material of Zonarius limbatus Kuhnt, 1911. (57) Lectotype; (5860), paralectotypes. Photographs provided by Bernd Jaeger." figureDoi="http://doi.org/10.5281/zenodo.3934226" httpUri="https://zenodo.org/record/3934226/files/figure.png" pageId="6" pageNumber="355">Figs. 5760</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>