412 lines
52 KiB
XML
412 lines
52 KiB
XML
<document id="4329C08B249A618E7E513F3131D4F6D4" ID-DOI="10.11646/zootaxa.3736.3.2" ID-GBIF-Dataset="6e974289-b881-47e6-ab54-e7b21a9d8a38" ID-ISSN="1175-5326" ID-Zenodo-Dep="215704" ID-ZooBank="36CBB3F6-59C7-47DC-BF47-E7DB6BFCB391" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460697698505" checkinUser="plazi" docAuthor="Suvarnaraksha, Apinun" docDate="2013" docId="F026EF11FFAA2176FF59C50D7BA2F855" docLanguage="en" docName="zt03736p248.pdf" docOrigin="Zootaxa 3736 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Physoschistura chulabhornae Suvarnaraksha, 2013, new species" docType="treatment" docVersion="8" lastPageNumber="244" masterDocId="0C1F9769FFAB217EFFCEC74A7B41FFA0" masterDocTitle="A new species of Physoschistura (Pisces: Nemacheilidae) from northern Thailand" masterLastPageNumber="248" masterPageNumber="236" pageNumber="237" updateTime="1698353975721" updateUser="plazi">
|
||
<mods:mods id="3F6A317D0AB45C04B330DAE60FBF1950" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="99D589C0CAC8C24F223CDC7AA42E9039">
|
||
<mods:title id="62CF614E6FCEFF98D191301549CBA937">A new species of Physoschistura (Pisces: Nemacheilidae) from northern Thailand</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="15ED4286991DB2E62144CF9431733F15" type="personal">
|
||
<mods:role id="2C9E07909E4C686CA33FFFE1F9E2F31E">
|
||
<mods:roleTerm id="10A2EB4D22B5AE615DAA6EEB2D2FE7F0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5A0DC80D0BCA97E02521DD9642007676">Suvarnaraksha, Apinun</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="41CC527B2CCDCE964D7A5931D1A007EC">text</mods:typeOfResource>
|
||
<mods:relatedItem id="96ECBB7001BFEA4068C92844EF60FA08" type="host">
|
||
<mods:titleInfo id="521707A38280E3332DE376CAF0EE34A7">
|
||
<mods:title id="180098626E731B82E89C7E278289E2AF">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="98C4D4CA7035A86C076C5D6E954991F8">
|
||
<mods:date id="BF6B9F3020233C2C734E16CA90CBD868">2013</mods:date>
|
||
<mods:detail id="120746A83BC74E555FF52F17350A7E9C" type="volume">
|
||
<mods:number id="7836F0EA5F68E7592BAE3CFFC4341ACB">3736</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="DAEE0CF02E5AC1D9B7914D345FBAF8BC" type="issue">
|
||
<mods:number id="B5E03B151DC34AE759CB618CFEF625ED">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="BFE13B9A04EB685A2D34791B09187B60" unit="page">
|
||
<mods:start id="450DE0CE2D65D2C90A6B7FA067458899">236</mods:start>
|
||
<mods:end id="AAB8B636DC8B3EF2E8333AC3D70C55C0">248</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="DA400A85372B8877B9D2CC1360A30CFE">journal article</mods:classification>
|
||
<mods:identifier id="FD614E3BEF5A8E592399A6BF178FBD7D" type="DOI">10.11646/zootaxa.3736.3.2</mods:identifier>
|
||
<mods:identifier id="0BAC452D63CC778886E7E862B7A594B0" type="GBIF-Dataset">6e974289-b881-47e6-ab54-e7b21a9d8a38</mods:identifier>
|
||
<mods:identifier id="261F5D63B728A471F84FDC80438B1CB8" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="A32199F9ED37A86FABA31270852A491A" type="Zenodo-Dep">215704</mods:identifier>
|
||
<mods:identifier id="FA54E5044AD94472431AD5D933AC5D16" type="ZooBank">36CBB3F6-59C7-47DC-BF47-E7DB6BFCB391</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="F026EF11FFAA2176FF59C50D7BA2F855" ID-DOI="http://doi.org/10.5281/zenodo.6148130" ID-GBIF-Taxon="119582184" ID-Zenodo-Dep="6148130" LSID="urn:lsid:plazi:treatment:F026EF11FFAA2176FF59C50D7BA2F855" httpUri="http://treatment.plazi.org/id/F026EF11FFAA2176FF59C50D7BA2F855" lastPageId="8" lastPageNumber="244" pageId="1" pageNumber="237">
|
||
<subSubSection id="30950D8CFFAA217FFF59C50D7A53FD23" pageId="1" pageNumber="237" type="nomenclature">
|
||
<paragraph id="78305E07FFAA217FFF59C50D79DFFDC2" blockId="1.[151,670,583,643]" box="[151,670,583,610]" pageId="1" pageNumber="237">
|
||
<heading id="2378E96BFFAA217FFF59C50D79DFFDC2" bold="true" box="[151,670,583,610]" fontSize="11" level="1" pageId="1" pageNumber="237" reason="1">
|
||
<emphasis id="4AFB8215FFAA217FFF59C50D79DFFDC2" bold="true" box="[151,670,583,610]" pageId="1" pageNumber="237">
|
||
<taxonomicName id="BF8F2584FFAA217FFF59C50D7ABEFDC1" box="[151,511,583,609]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="237" phylum="Chordata" rank="species" species="chulabhornae" status="sp. nov.">
|
||
<emphasis id="4AFB8215FFAA217FFF59C50D7ABEFDC1" bold="true" box="[151,511,583,609]" italics="true" pageId="1" pageNumber="237">Physoschistura chulabhornae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="51C83F6EFFAA217FFDC0C50279DFFDC2" box="[526,670,584,610]" pageId="1" pageNumber="237" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="78305E07FFAA217FFF59C5207A53FD23" blockId="1.[151,670,583,643]" box="[151,274,618,643]" pageId="1" pageNumber="237">
|
||
(
|
||
<figureCitation id="E0B44282FFAA217FFF51C5207BADFD23" box="[159,236,618,643]" captionStart="FIGURE 1" captionStartId="1.[151,250,1465,1487]" captionTargetBox="[151,1436,1022,1440]" captionTargetId="figure@1.[151,1436,1022,1441]" captionTargetPageId="1" captionText="FIGURE 1. Lateral view of holotype of Physoschistura chulabhornae sp. nov." httpUri="https://zenodo.org/record/215705/files/figure.png" pageId="1" pageNumber="237">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="E0B44282FFAA217FFF34C5217A48FD23" box="[250,265,619,643]" captionStart="FIGURE 2" captionStartId="1.[151,250,1963,1985]" captionTargetBox="[151,1436,1537,1939]" captionTargetId="figure@1.[151,1436,1537,1940]" captionTargetPageId="1" captionText="FIGURE 2. Live individual of Physoschistura chulabhornae (uncataloged)." httpUri="https://zenodo.org/record/215706/files/figure.png" pageId="1" pageNumber="237">2</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="30950D8CFFAA217FFF59C5F87E10FCFB" pageId="1" pageNumber="237" type="materials_examined">
|
||
<paragraph id="78305E07FFAA217FFF59C5F87E28FCB3" blockId="1.[151,1436,690,967]" pageId="1" pageNumber="237">
|
||
<emphasis id="4AFB8215FFAA217FFF59C5F87A4AFD6B" bold="true" box="[151,267,690,715]" pageId="1" pageNumber="237">
|
||
<typeStatus id="A734E0A5FFAA217FFF59C5F87A47FD6B" box="[151,262,690,715]" pageId="1" pageNumber="237" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
MARNM 3998,
|
||
<date id="0C3178C7FFAA217FFE2CC5F8793AFD6B" box="[482,635,690,715]" pageId="1" pageNumber="237" value="2008-05-06">6 May 2008</date>
|
||
, Huay Maetalanoi, Ban Maetala, T. Maedaed, Maechaem District, Chiangmai, near a village,
|
||
<geoCoordinate id="1DBB38C0FFAA217FFE00C59C79C1FD4F" box="[462,640,726,751]" direction="north" orientation="latitude" pageId="1" pageNumber="237" precision="1" value="18.926508">18°55' 35.43''N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1DBB38C0FFAA217FFD40C59C7803FD4F" box="[654,834,726,751]" direction="east" orientation="longitude" pageId="1" pageNumber="237" precision="1" value="98.37934">098°22'45.63''E</geoCoordinate>
|
||
; tributary of Ping River, northern Chaophraya River basin, Maechaem District, Chiangmai Province, coll. A. Suvarnaraksha, R. Wattanasiriserekul and colleagues.
|
||
</paragraph>
|
||
<paragraph id="78305E07FFAA217FFF09C4547E10FCFB" blockId="1.[151,1436,690,967]" pageId="1" pageNumber="237">
|
||
<emphasis id="4AFB8215FFAA217FFF09C4547A07FC97" bold="true" box="[199,326,798,823]" pageId="1" pageNumber="237">
|
||
<typeStatus id="A734E0A5FFAA217FFF09C4547A00FC97" box="[199,321,798,823]" pageId="1" pageNumber="237" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
MARNM 3999, 36 ex.,
|
||
<quantity id="BF77F3E2FFAA217FFD90C4547843FC96" box="[606,770,798,823]" metricMagnitude="-2" metricUnit="m" metricValue="3.155" metricValueMax="3.65" metricValueMin="2.66" pageId="1" pageNumber="237" unit="mm" value="31.55" valueMax="36.5" valueMin="26.6">26.6–36.5 mm</quantity>
|
||
SL,
|
||
<quantity id="BF77F3E2FFAA217FFCF4C454789FFC96" box="[826,990,798,823]" metricMagnitude="-2" metricUnit="m" metricValue="4.12" metricValueMax="4.76" metricValueMin="3.4799999999999995" pageId="1" pageNumber="237" unit="mm" value="41.2" valueMax="47.6" valueMin="34.8">34.8–47.6 mm</quantity>
|
||
TL, NIFI: 0 4836, 3 ex., 31.0–
|
||
<quantity id="BF77F3E2FFAA217FFAF6C4557EDDFC96" box="[1336,1436,799,823]" metricMagnitude="-2" metricUnit="m" metricValue="3.38" pageId="1" pageNumber="237" unit="mm" value="33.8">33.8 mm</quantity>
|
||
SL,
|
||
<quantity id="BF77F3E2FFAA217FFF0BC4087A28FCFA" box="[197,361,834,859]" metricMagnitude="-2" metricUnit="m" metricValue="4.045" metricValueMax="4.24" metricValueMin="3.85" pageId="1" pageNumber="237" unit="mm" value="40.45" valueMax="42.4" valueMin="38.5">38.5–42.4 mm</quantity>
|
||
TL, and UF 185354, 3 ex.,
|
||
<quantity id="BF77F3E2FFAA217FFD6EC4097802FCFA" box="[672,835,835,859]" metricMagnitude="-2" metricUnit="m" metricValue="3.08" metricValueMax="3.32" metricValueMin="2.84" pageId="1" pageNumber="237" unit="mm" value="30.8" valueMax="33.2" valueMin="28.4">28.4–33.2 mm</quantity>
|
||
SL, 38.0–
|
||
<quantity id="BF77F3E2FFAA217FFC79C4097F5AFCFA" box="[951,1051,835,859]" metricMagnitude="-2" metricUnit="m" metricValue="4.34" pageId="1" pageNumber="237" unit="mm" value="43.4">43.4 mm</quantity>
|
||
TL; same data as
|
||
<typeStatus id="A734E0A5FFAA217FFB29C4087E0CFCFB" box="[1255,1357,834,859]" pageId="1" pageNumber="237" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="30950D8CFFAA217FFF09C42C7FFBFC67" pageId="1" pageNumber="237" type="diagnosis">
|
||
<paragraph id="78305E07FFAA217FFF09C42C7FFBFC67" blockId="1.[151,1436,690,967]" pageId="1" pageNumber="237">
|
||
<emphasis id="4AFB8215FFAA217FFF09C42C7A00FCDF" bold="true" box="[199,321,870,895]" pageId="1" pageNumber="237">Diagnosis.</emphasis>
|
||
This new species can be distinguished from other members of
|
||
<taxonomicName id="BF8F2584FFAA217FFBE9C42C7F93FCDE" box="[1063,1234,870,894]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="237" phylum="Chordata" rank="genus">
|
||
<emphasis id="4AFB8215FFAA217FFBE9C42C7F93FCDE" box="[1063,1234,870,894]" italics="true" pageId="1" pageNumber="237">Physoschistura</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters: incomplete lateral line with 62–83 pores, reaching between anal-fin origin and anal-fin tip; no axillary lobe on pelvic-fin base; male with hammer head-shaped suborbital flap (
|
||
<figureCitation id="E0B44282FFAA217FFBBEC4E47FECFC67" box="[1136,1197,942,967]" captionStart="FIGURE 1" captionStartId="1.[151,250,1465,1487]" captionTargetBox="[151,1436,1022,1440]" captionTargetId="figure@1.[151,1436,1022,1441]" captionTargetPageId="1" captionText="FIGURE 1. Lateral view of holotype of Physoschistura chulabhornae sp. nov." httpUri="https://zenodo.org/record/215705/files/figure.png" pageId="1" pageNumber="237">Fig.1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="2CF00E8FFFAA217FFF59C2F378F6FA6F" box="[151,951,1465,1487]" httpUri="https://zenodo.org/record/215705/files/figure.png" pageId="1" pageNumber="237" targetBox="[151,1436,1022,1440]" targetPageId="1">
|
||
<paragraph id="78305E07FFAA217FFF59C2F378F6FA6F" blockId="1.[151,951,1465,1487]" box="[151,951,1465,1487]" pageId="1" pageNumber="237">
|
||
<emphasis id="4AFB8215FFAA217FFF59C2F37A52FA6F" bold="true" box="[151,275,1465,1487]" pageId="1" pageNumber="237">FIGURE 1.</emphasis>
|
||
Lateral view of holotype of
|
||
<taxonomicName id="BF8F2584FFAA217FFDFAC2F37820FA6F" box="[564,865,1465,1487]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="237" phylum="Chordata" rank="species" species="chulabhornae" status="sp. nov.">
|
||
<emphasis id="4AFB8215FFAA217FFDFAC2F37820FA6F" box="[564,865,1465,1487]" italics="true" pageId="1" pageNumber="237">Physoschistura chulabhornae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="51C83F6EFFAA217FFCA6C2F078F6FA6F" box="[872,951,1466,1487]" pageId="1" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2CF00E8FFFAA217FFF59C0E178DDF861" box="[151,924,1963,1985]" httpUri="https://zenodo.org/record/215706/files/figure.png" pageId="1" pageNumber="237" targetBox="[151,1436,1537,1939]" targetPageId="1">
|
||
<paragraph id="78305E07FFAA217FFF59C0E178DDF861" blockId="1.[151,924,1963,1985]" box="[151,924,1963,1985]" pageId="1" pageNumber="237">
|
||
<emphasis id="4AFB8215FFAA217FFF59C0E17A53F861" bold="true" box="[151,274,1963,1985]" pageId="1" pageNumber="237">FIGURE 2.</emphasis>
|
||
Live individual of
|
||
<taxonomicName id="BF8F2584FFAA217FFE18C0E17842F861" box="[470,771,1963,1985]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="237" phylum="Chordata" rank="species" species="chulabhornae">
|
||
<emphasis id="4AFB8215FFAA217FFE18C0E17842F861" box="[470,771,1963,1985]" italics="true" pageId="1" pageNumber="237">Physoschistura chulabhornae</emphasis>
|
||
</taxonomicName>
|
||
(uncataloged).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="30950D8CFFA92176FF09C7DD7F6EF8C4" lastPageId="8" lastPageNumber="244" pageId="2" pageNumber="238" type="description">
|
||
<paragraph id="78305E07FFA9217CFF09C7DD793CFD9C" blockId="2.[151,1436,151,573]" pageId="2" pageNumber="238">
|
||
<emphasis id="4AFB8215FFA9217CFF09C7DD7A18FF10" bold="true" box="[199,345,151,176]" pageId="2" pageNumber="238">Description.</emphasis>
|
||
Morphometric measurements, proportions and meristic counts of the
|
||
<typeStatus id="A734E0A5FFA9217CFBBCC7DD7F95FF10" box="[1138,1236,151,176]" pageId="2" pageNumber="238" type="holotype">holotype</typeStatus>
|
||
and 42
|
||
<typeStatus id="A734E0A5FFA9217CFAFFC7D27EDDFF10" box="[1329,1436,152,176]" pageId="2" pageNumber="238" type="paratype">paratypes</typeStatus>
|
||
are presented in
|
||
<tableCitation id="350D6BBCFFA9217CFEA0C7F67A92FF74" box="[366,467,188,213]" captionStart="TABLE 1" captionStartId="2.[151,239,620,643]" captionText="TABLE 1. Morphometric data on the holotype and 42 paratypes of Physoschistura chulabhornae, sp. nov." pageId="2" pageNumber="238">Tables 1</tableCitation>
|
||
–3. A relatively small to medium-sized species compared to other species of
|
||
<taxonomicName id="BF8F2584FFA9217CFF59C7AA7A03FF58" box="[151,322,224,248]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="238" phylum="Chordata" rank="genus">
|
||
<emphasis id="4AFB8215FFA9217CFF59C7AA7A03FF58" box="[151,322,224,248]" italics="true" pageId="2" pageNumber="238">Physoschistura</emphasis>
|
||
</taxonomicName>
|
||
with a moderately elongated body; body sub-cylindrical anteriorly in cross section, compressed posteriorly; dorsal profile slightly arched, ventral profile straight throughout; snout obtuse to pointed; strongly arched mouth, 1.5–1.7±0.08 times wider than long with medially interrupted lower lip forming two lateral broadly triangular pads with deep furrows, pads 1.5±0.07 times wider than long (
|
||
<figureCitation id="E0B44282FFA9217CFC2BC6067F6AFEC4" box="[997,1067,332,357]" captionStart="FIGURE 3. A" captionStartId="6.[151,250,1969,1991]" captionTargetBox="[216,1367,195,1940]" captionTargetId="figure@6.[216,1367,195,1940]" captionTargetPageId="6" captionText="FIGURE 3. A) Ventral view of head of Physoschistura chulabhornae, MARNM 3999 / 7, 32.37 mm SL; B) lateral view of head of Physoschistura chulabhornae. showing suborbital flap of male, MARNM 3999 / 20, 30.03 mm SL." httpUri="https://zenodo.org/record/215707/files/figure.png" pageId="2" pageNumber="238">Fig. 3</figureCitation>
|
||
); lips thin, upper lip with small incision in middle, lower lip with deep median interruption; anterior nostril in front of flap-like tube with pointed tip; eye small, 4.0–4.8 times in HL (mean 4.3±0.2), eye diameter 3.0–3.8 times in interorbital space. Inner rostral barbel longer than outer rostral barbel and maxillary barbel, extending to middle of eye, longer than snout (0.7– 0.96±0.06); outer rostral barbel not reaching to anterior margin of eye; maxillary barbel reaching to posterior margin of eye. Processus dentiformis present, not prominent. Free posterior chamber of air bladder well formed, not encapsulated, and slightly oval (
|
||
<figureCitation id="E0B44282FFA9217CFDE4C56E792EFD9C" box="[554,623,548,573]" captionStart="FIGURES 4 – 5. 4" captionStartId="7.[151,264,1297,1319]" captionTargetBox="[193,1388,203,1246]" captionTargetId="figure@7.[151,1436,193,1276]" captionTargetPageId="7" captionText="FIGURES 4 – 5. 4. Free posterior chamber of air bladder of Physoschistura chulabhornae. Scale bar 1 mm. 5. Stomach and loop of intestine of Physoschistura chulabhornae. Scale bar 1 mm." httpUri="https://zenodo.org/record/215708/files/figure.png" pageId="2" pageNumber="238">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="2CF00E8FFFA9217CFF59C5267E48FD23" box="[151,1289,619,643]" pageId="2" pageNumber="238">
|
||
<paragraph id="78305E07FFA9217CFF59C5267E48FD23" blockId="2.[151,1289,619,643]" box="[151,1289,619,643]" pageId="2" pageNumber="238">
|
||
<emphasis id="4AFB8215FFA9217CFF59C5267A48FD23" bold="true" box="[151,265,620,643]" pageId="2" pageNumber="238">TABLE 1.</emphasis>
|
||
Morphometric data on the holotype and 42 paratypes of
|
||
<taxonomicName id="BF8F2584FFA9217CFCA7C5217FE7FD23" box="[873,1190,619,643]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="238" phylum="Chordata" rank="species" species="chulabhornae" status="sp. nov.">
|
||
<emphasis id="4AFB8215FFA9217CFCA7C5217FE7FD23" box="[873,1190,619,643]" italics="true" pageId="2" pageNumber="238">Physoschistura chulabhornae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="51C83F6EFFA9217CFB7AC5267E48FD23" box="[1204,1289,620,643]" pageId="2" pageNumber="238" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="78305E07FFA9217CFF51C5D47F61FD14" blockId="2.[159,1424,670,2015]" box="[159,1056,670,692]" pageId="2" pageNumber="238">
|
||
Morphometric characters
|
||
<typeStatus id="A734E0A5FFA9217CFD70C5D4785DFD14" box="[702,796,670,692]" pageId="2" pageNumber="238" type="holotype">Holotype</typeStatus>
|
||
<typeStatus id="A734E0A5FFA9217CFCA8C5D57889FD14" box="[870,968,671,692]" pageId="2" pageNumber="238" type="paratype">Paratypes</typeStatus>
|
||
(n = 42)
|
||
</paragraph>
|
||
<paragraph id="78305E07FFA9217CFCA8C5837E4AFD7E" blockId="2.[159,1424,670,2015]" box="[870,1291,712,734]" pageId="2" pageNumber="238">Range mean SD</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C5BB7BBDFC90" blockId="2.[159,1424,670,2015]" pageId="2" pageNumber="238">
|
||
Standard length (mm) 30.49 25.82–39.98 31.16 2.2
|
||
<emphasis id="4AFB8215FFA9217CFF70C4507BBDFC90" bold="true" box="[190,252,794,816]" pageId="2" pageNumber="238">% SL</emphasis>
|
||
</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C40E7E4BFCFA" blockId="2.[159,1424,670,2015]" box="[159,1290,836,858]" pageId="2" pageNumber="238">Total length 126.2 123.3–133.7 128.6 2.3</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C4277E4BFC23" blockId="2.[159,1424,670,2015]" box="[159,1290,877,899]" pageId="2" pageNumber="238">Fork length 121.9 117.5–126.9 122.4 2.2</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C4DC7E4BFC0C" blockId="2.[159,1424,670,2015]" box="[159,1290,918,940]" pageId="2" pageNumber="238">Prepectoral-fin length 25.0 21.8–28.7 24.2 1.5</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C48A7E4BFC76" blockId="2.[159,1424,670,2015]" box="[159,1290,960,982]" pageId="2" pageNumber="238">Pectoral-fin length 21.7 20.8–25.3 23.2 1.1</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C4A37E4BFC5F" blockId="2.[159,1424,670,2015]" box="[159,1290,1001,1023]" pageId="2" pageNumber="238">Predorsal-fin length 50.2 46.8–52.7 49.4 1.4</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C3587E4BFB88" blockId="2.[159,1424,670,2015]" box="[159,1290,1042,1064]" pageId="2" pageNumber="238">Preanal-fin length 76.2 71.5–79.8 75.9 1.7</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C3767E4BFBF2" blockId="2.[159,1424,670,2015]" box="[159,1290,1084,1106]" pageId="2" pageNumber="238">Prepelvic-fin length 48.5 47.0–53.9 49.9 1.6</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C32F7E4BFBDB" blockId="2.[159,1424,670,2015]" box="[159,1290,1125,1147]" pageId="2" pageNumber="238">Pelvic-fin length 21.3 18.4–23.5 20.3 1.1</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C3C47E4BFB04" blockId="2.[159,1424,670,2015]" box="[159,1290,1166,1188]" pageId="2" pageNumber="238">Pelvic to vent distance 21.3 20.5–27.1 23.0 1.5</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C3F27E4BFB6E" blockId="2.[159,1424,670,2015]" box="[159,1290,1208,1230]" pageId="2" pageNumber="238">Dorsal-fin base length 17.7 15.3–22.9 17.9 1.5</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C3AB7E4BFB57" blockId="2.[159,1424,670,2015]" box="[159,1290,1249,1271]" pageId="2" pageNumber="238">Dorsal-fin length 23.1 19.5–26.7 23.6 1.8</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C2407E4BFA80" blockId="2.[159,1424,670,2015]" box="[159,1290,1290,1312]" pageId="2" pageNumber="238">Anal-fin base length 9.4 7.1–13.9 11.1 1.6</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C27E7E4BFAEA" blockId="2.[159,1424,670,2015]" box="[159,1290,1332,1354]" pageId="2" pageNumber="238">Anal-fin length 19.9 16.8–24.7 20.5 1.6</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C2177E4BFAD3" blockId="2.[159,1424,670,2015]" box="[159,1290,1373,1395]" pageId="2" pageNumber="238">Body depth at dorsal-fin origin 17.8 15.6–23.2 19.6 1.8</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C2CC7E4BFA3C" blockId="2.[159,1424,670,2015]" box="[159,1290,1414,1436]" pageId="2" pageNumber="238">Body width at dorsal-fin origin 10.1 8.6–18.8 12.8 2.2</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C2FA7E4BFA66" blockId="2.[159,1424,670,2015]" box="[159,1290,1456,1478]" pageId="2" pageNumber="238">Body depth at anal-fin origin 13.6 13.0–16.3 14.1 0.8</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C2937E4BFA4F" blockId="2.[159,1424,670,2015]" box="[159,1290,1497,1519]" pageId="2" pageNumber="238">Body width at anal-fin origin 6.2 5.3–8.5 7.0 0.8</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C1487E4BF9B8" blockId="2.[159,1424,670,2015]" box="[159,1290,1538,1560]" pageId="2" pageNumber="238">Caudal-peduncle length 12.4 10.7–15.0 12.4 1.0</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C1667E4BF9E2" blockId="2.[159,1424,670,2015]" box="[159,1290,1580,1602]" pageId="2" pageNumber="238">Caudal-peduncle depth 10.7 8.2–11.9 10.7 0.6</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C11F7E4BF9CB" blockId="2.[159,1424,670,2015]" box="[159,1290,1621,1643]" pageId="2" pageNumber="238">Caudal-fin length 26.2 23.3–33.7 29.2 2.3</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C1347E4BF934" blockId="2.[159,1424,670,2015]" box="[159,1290,1662,1684]" pageId="2" pageNumber="238">Head length 22.3 22.3–26.6 24.1 1.2</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C1E27E4BF91E" blockId="2.[159,1424,670,2015]" box="[159,1290,1704,1726]" pageId="2" pageNumber="238">Dorsal head length 18.3 18.2–23.4 20.6 1.2</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C19B7E4BF947" blockId="2.[159,1424,670,2015]" box="[159,1290,1745,1767]" pageId="2" pageNumber="238">Head width 10.7 10.0–12.7 11.3 0.6</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C1B07A43F89A" blockId="2.[159,1424,670,2015]" pageId="2" pageNumber="238">
|
||
Head depth at occiput 13.2 12.7–15.2 13.8 0.6
|
||
<emphasis id="4AFB8215FFA9217CFF70C06E7A43F89A" bold="true" box="[190,258,1828,1850]" pageId="2" pageNumber="238">% HL</emphasis>
|
||
</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C0077E4BF8C3" blockId="2.[159,1424,670,2015]" box="[159,1290,1869,1891]" pageId="2" pageNumber="238">Dorsal head length 82.0 81.2–86.1 84.8 1.4</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C03C7E4BF82C" blockId="2.[159,1424,670,2015]" box="[159,1290,1910,1932]" pageId="2" pageNumber="238">Head width 48.0 42.6–48.9 44.9 1.7</paragraph>
|
||
<paragraph id="78305E07FFA9217CFF51C0EA7E4AF816" blockId="2.[159,1424,670,2015]" box="[159,1291,1952,1974]" pageId="2" pageNumber="238">Head depth at occiput 56.4 52.3–56.4 55.1 1.0</paragraph>
|
||
<paragraph id="78305E07FFA9217DFB94C0837F61FF7F" blockId="2.[159,1424,670,2015]" lastBlockId="3.[159,1303,160,1422]" lastPageId="3" lastPageNumber="239" pageId="2" pageNumber="238">
|
||
<emphasis id="4AFB8215FFA9217CFB94C0837ED1F87F" box="[1114,1424,1993,2015]" italics="true" pageId="2" pageNumber="238">......continued on the next page</emphasis>
|
||
Morphometric characters
|
||
<typeStatus id="A734E0A5FFA8217DFD70C783785DFF7F" box="[702,796,201,223]" pageId="3" pageNumber="239" type="holotype">Holotype</typeStatus>
|
||
<typeStatus id="A734E0A5FFA8217DFCA8C7807889FF7F" box="[870,968,202,223]" pageId="3" pageNumber="239" type="paratype">Paratypes</typeStatus>
|
||
(n = 42)
|
||
</paragraph>
|
||
<caption id="2CF00E8FFFA8217DFF51C7EA7ACDFF16" box="[159,396,160,182]" pageId="3" pageNumber="239">
|
||
<paragraph id="78305E07FFA8217DFF51C7EA7ACDFF16" blockId="3.[159,1303,160,1422]" box="[159,396,160,182]" pageId="3" pageNumber="239">
|
||
<emphasis id="4AFB8215FFA8217DFF51C7EA7A4DFF16" bold="true" box="[159,268,160,182]" pageId="3" pageNumber="239">TABLE 1.</emphasis>
|
||
(Continued)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="78305E07FFA8217DFCA8C7B97E4AFEA8" blockId="3.[159,1303,160,1422]" box="[870,1291,242,264]" pageId="3" pageNumber="239">Range mean SD</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C6577E4BFE92" blockId="3.[159,1303,160,1422]" box="[159,1290,284,306]" pageId="3" pageNumber="239">Eye diameter 25.8 22.5–26.0 24.4 1.0</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C60F7E4BFEFB" blockId="3.[159,1303,160,1422]" box="[159,1290,325,347]" pageId="3" pageNumber="239">Interorbital length 29.9 28.0–31.9 29.9 0.7</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C6247E4BFE24" blockId="3.[159,1303,160,1422]" box="[159,1290,366,388]" pageId="3" pageNumber="239">Inter-nostril distance 20.9 19.0–21.7 20.4 0.6</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C6D27E4BFE0E" blockId="3.[159,1303,160,1422]" box="[159,1290,408,430]" pageId="3" pageNumber="239">Snout length 35.1 33.2–35.9 35.0 0.7</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C68B7E4BFE77" blockId="3.[159,1303,160,1422]" box="[159,1290,449,471]" pageId="3" pageNumber="239">Mouth width 28.7 25.9–29.9 28.4 0.9</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C6A07E4BFDA0" blockId="3.[159,1303,160,1422]" box="[159,1290,490,512]" pageId="3" pageNumber="239">Mouth length 18.6 16.3–19.1 17.6 0.7</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C55E7E4BFD8A" blockId="3.[159,1303,160,1422]" box="[159,1290,532,554]" pageId="3" pageNumber="239">Rostral barbel I length 29.5 27.3–29.9 29.2 0.7</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C5777E4BFDF3" blockId="3.[159,1303,160,1422]" box="[159,1290,573,595]" pageId="3" pageNumber="239">Rostral barbel II length 42.0 39.1–43.0 41.4 1.0</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C52C7A4BFD06" blockId="3.[159,1303,160,1422]" pageId="3" pageNumber="239">
|
||
Maxillary-barbel length 35.2 34.2–36.9 35.1 0.7
|
||
<emphasis id="4AFB8215FFA8217DFF0BC5DA7A4BFD06" bold="true" box="[197,266,656,678]" pageId="3" pageNumber="239">Ratios</emphasis>
|
||
</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C5F37E56FD6F" blockId="3.[159,1303,160,1422]" box="[159,1303,697,719]" pageId="3" pageNumber="239">Mouth width/mouth length 1.5 1.5–1.7 1.6 0.08</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C5A87E56FD58" blockId="3.[159,1303,160,1422]" box="[159,1303,738,760]" pageId="3" pageNumber="239">Snout/maxillary-barbel length 1.0 0.9–1.0 1.0 0.03</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C4467E4BFC82" blockId="3.[159,1303,160,1422]" box="[159,1290,780,802]" pageId="3" pageNumber="239">Head length/eye diameter 3.9 3.9–4.4 4.1 0.2</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C47C7E56FCEB" blockId="3.[159,1303,160,1422]" box="[159,1303,821,843]" pageId="3" pageNumber="239">Eye diameter/Interorbital space 0.9 0.7–0.9 0.8 0.04</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C4147E56FCD4" blockId="3.[159,1303,160,1422]" box="[159,1303,862,884]" pageId="3" pageNumber="239">Lateral head length/pectoral-fin length 1.0 0.9–1.2 1.0 0.06</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C4C27E56FC3E" blockId="3.[159,1303,160,1422]" box="[159,1303,904,926]" pageId="3" pageNumber="239">Caudal-peduncle depth/caudal-peduncle length 1.2 1.0–1.4 1.2 0.12</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C4FB7E4BFBBA" blockId="3.[159,1303,160,1422]" pageId="3" pageNumber="239">Standard length/pelvic-fin length 4.7 3.9–4.8 4.3 0.06 males 4.7 4.3–5.0 4.7 0.2 females - 4.6–5.4 5.0 0.2</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C3677E4BFB36" blockId="3.[159,1303,160,1422]" pageId="3" pageNumber="239">Body depth/body width at dorsal-fin origin 1.8 0.9–2.1 1.5 0.2 males 1.8 1.4–2.1 1.8 0.2 females - 0.9–1.6 1.5 0.1</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C3E37E56FAB2" blockId="3.[159,1303,160,1422]" pageId="3" pageNumber="239">Pelvic-fin length/vent to anus distance 1.0 0.8–1.1 0.9 0.1 males 1.0 0.78–1.0 0.95 0.07 females - 0.78–0.96 0.87 0.05</paragraph>
|
||
<paragraph id="78305E07FFA8217DFF51C26F7AAEF840" blockId="3.[159,1303,160,1422]" lastBlockId="3.[151,1436,1485,2017]" pageId="3" pageNumber="239">
|
||
Body depth at dorsal fin/body depth at anal fin 1.3 1.2–1.6 1.4 0.1 males 1.3 1.2–1.4 1.3 0.1 females - 1.3–1.6 1.4 0.1 Body covered with embedded minute scales, except belly naked between pectoral fin bases. Incomplete lateral line reaching between anal-fin origin and tip of anal fin with 62–83 pores. Cephalic lateral-line system with 7 supraorbital, 4+8–10 infraorbital, 7 preoperculo-mandibular, and 3 supratemporal pores. Unculi present on lips, barbels, and pectoral-fin rays. Dorsal fin half-way between tip of snout and caudal-fin base; dorsal fin with 4 simple and 8½ branched rays, distal margin slightly convex. Pectoral fin rounded, the length 1.04±0.06 times lateral head length, origin near surface of breast, with 1 simple and 11 branched rays, reaching two-thirds distance to pelvic-fin origin in female, but close to pelvic-fin origin in male. Pelvic-fin origin slightly behind dorsal-fin origin, at vertical from 1st or 2nd branched dorsal-fin rays, with 1 simple and 7 branched rays. Distal margin of pelvic fin not reaching vent in female, nearly reaching vent in male when adpressed. Pelvic-fin length 20.3±1.1% SL; fin not extending beyond anus, except reaching anus in some males (0.96±0.04% HL). Anal fin not reaching base of caudal fin, with 3 simple rays and 5½ branched rays; no adipose crest on caudal peduncle. Anus is much nearer to anal fin than to caudal-fin base. Intestine curved toward lower part of stomach (
|
||
<figureCitation id="E0B44282FFA8217DFBBAC0167FF9F8D5" box="[1140,1208,1884,1909]" captionStart="FIGURES 4 – 5. 4" captionStartId="7.[151,264,1297,1319]" captionTargetBox="[193,1388,203,1246]" captionTargetId="figure@7.[151,1436,193,1276]" captionTargetPageId="7" captionText="FIGURES 4 – 5. 4. Free posterior chamber of air bladder of Physoschistura chulabhornae. Scale bar 1 mm. 5. Stomach and loop of intestine of Physoschistura chulabhornae. Scale bar 1 mm." httpUri="https://zenodo.org/record/215708/files/figure.png" pageId="3" pageNumber="239">Fig. 5</figureCitation>
|
||
). Caudal fin with 8– 9 upper and 8 lower branched rays, forked, upper lobe slightly shorter than lower lobe, both lobes with rounded tips. Caudal peduncle 1.2±0.1 times longer than deep. Largest recorded size:
|
||
<quantity id="BF77F3E2FFA8217DFBC2C0EE7F33F81C" box="[1036,1138,1956,1981]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" pageId="3" pageNumber="239" unit="mm" value="36.5">36.5 mm</quantity>
|
||
SL (
|
||
<quantity id="BF77F3E2FFA8217DFB65C0EE7E52F81C" box="[1195,1299,1956,1981]" metricMagnitude="-2" metricUnit="m" metricValue="4.76" pageId="3" pageNumber="239" unit="mm" value="47.6">47.6 mm</quantity>
|
||
TL) female (
|
||
<typeStatus id="A734E0A5FFA8217DFF50C0827A42F840" box="[158,259,1992,2016]" pageId="3" pageNumber="239" type="paratype">paratype</typeStatus>
|
||
, MARNM 3999/14).
|
||
</paragraph>
|
||
<caption id="2CF00E8FFFAD2178FF59C0FB7FD4F846" httpUri="https://zenodo.org/record/215707/files/figure.png" pageId="6" pageNumber="242" targetBox="[216,1367,195,1940]" targetPageId="6">
|
||
<paragraph id="78305E07FFAD2178FF59C0FB7FD4F846" blockId="6.[151,1436,1969,2022]" pageId="6" pageNumber="242">
|
||
<emphasis id="4AFB8215FFAD2178FF59C0FB7A53F867" bold="true" box="[151,274,1969,1991]" pageId="6" pageNumber="242">FIGURE 3.</emphasis>
|
||
A) Ventral view of head of
|
||
<taxonomicName id="BF8F2584FFAD2178FDE5C0FB7816F867" box="[555,855,1969,1991]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="242" phylum="Chordata" rank="species" species="chulabhornae">
|
||
<emphasis id="4AFB8215FFAD2178FDE5C0FB7816F867" box="[555,855,1969,1991]" italics="true" pageId="6" pageNumber="242">Physoschistura chulabhornae</emphasis>
|
||
</taxonomicName>
|
||
, MARNM 3999/7, 32.37 mm SL; B) lateral view of head of
|
||
<taxonomicName id="BF8F2584FFAD2178FF7CC09A7A9EF846" box="[178,479,2000,2022]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="242" phylum="Chordata" rank="species" species="chulabhornae">
|
||
<emphasis id="4AFB8215FFAD2178FF7CC09A7A9EF846" box="[178,479,2000,2022]" italics="true" pageId="6" pageNumber="242">Physoschistura chulabhornae</emphasis>
|
||
</taxonomicName>
|
||
. showing suborbital flap of male, MARNM 3999/20, 30.03 mm SL.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2CF00E8FFFAC2179FF59C25B787CFAE9" httpUri="https://zenodo.org/record/215708/files/figure.png" pageId="7" pageNumber="243" targetBox="[193,1388,203,1246]" targetPageId="7">
|
||
<paragraph id="78305E07FFAC2179FF59C25B787CFAE9" blockId="7.[151,1435,1297,1353]" pageId="7" pageNumber="243">
|
||
<emphasis id="4AFB8215FFAC2179FF59C25B7A16FA87" bold="true" box="[151,343,1297,1319]" pageId="7" pageNumber="243">FIGURES 4–5. 4.</emphasis>
|
||
Free posterior chamber of air bladder of
|
||
<taxonomicName id="BF8F2584FFAC2179FCC4C25B7F78FA87" box="[778,1081,1297,1319]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="243" phylum="Chordata" rank="species" species="chulabhornae">
|
||
<emphasis id="4AFB8215FFAC2179FCC4C25B7F78FA87" box="[778,1081,1297,1319]" italics="true" pageId="7" pageNumber="243">Physoschistura chulabhornae</emphasis>
|
||
</taxonomicName>
|
||
. Scale bar 1 mm.
|
||
<emphasis id="4AFB8215FFAC2179FB35C25B7E4FFA87" bold="true" box="[1275,1294,1297,1319]" pageId="7" pageNumber="243">5.</emphasis>
|
||
Stomach and loop of intestine of
|
||
<taxonomicName id="BF8F2584FFAC2179FE93C27979CBFAE9" box="[349,650,1331,1353]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="243" phylum="Chordata" rank="species" species="chulabhornae">
|
||
<emphasis id="4AFB8215FFAC2179FE93C27979CBFAE9" box="[349,650,1331,1353]" italics="true" pageId="7" pageNumber="243">Physoschistura chulabhornae</emphasis>
|
||
</taxonomicName>
|
||
. Scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="78305E07FFAC2179FF09C23278E7F9C9" blockId="7.[151,1436,1400,2001]" pageId="7" pageNumber="243">
|
||
<emphasis id="4AFB8215FFAC2179FF09C2327A2AFA31" bold="true" box="[199,363,1400,1425]" pageId="7" pageNumber="243">Colour in life.</emphasis>
|
||
Body translucent and light yellowish-brown. Eight-10 dark olivaceous blotches on side of body, 8–10 dark saddles on back not extending to belly; large males have fewer blotches on side of body and faint dorsal saddles; iridescent stripes on belly and lower half of body from pectoral fin to anus (
|
||
<figureCitation id="E0B44282FFAC2179FBA0C28A7FF7FA78" box="[1134,1206,1472,1497]" captionStart="FIGURE 2" captionStartId="1.[151,250,1963,1985]" captionTargetBox="[151,1436,1537,1939]" captionTargetId="figure@1.[151,1436,1537,1940]" captionTargetPageId="1" captionText="FIGURE 2. Live individual of Physoschistura chulabhornae (uncataloged)." httpUri="https://zenodo.org/record/215706/files/figure.png" pageId="7" pageNumber="243">Fig. 2</figureCitation>
|
||
). Lower lip without black mark on each side of median interruption (
|
||
<figureCitation id="E0B44282FFAC2179FD11C2AE7877FA5D" box="[735,822,1508,1533]" captionStart="FIGURE 3. A" captionStartId="6.[151,250,1969,1991]" captionTargetBox="[216,1367,195,1940]" captionTargetId="figure@6.[216,1367,195,1940]" captionTargetPageId="6" captionText="FIGURE 3. A) Ventral view of head of Physoschistura chulabhornae, MARNM 3999 / 7, 32.37 mm SL; B) lateral view of head of Physoschistura chulabhornae. showing suborbital flap of male, MARNM 3999 / 20, 30.03 mm SL." httpUri="https://zenodo.org/record/215707/files/figure.png" pageId="7" pageNumber="243">Fig. 3a</figureCitation>
|
||
). Dorsal and dorso-lateral portion of head with few irregular dark spots. Basicaudal bar fragmented into small blotch on upper caudal-fin base, vertical olivaceous blotch on lower caudal-fin base; thin, horizontally oriented, faint Vs on median part of caudal fin. Black spot at base of simple ray of dorsal fin. Pectoral, pelvic and anal fins hyaline.
|
||
</paragraph>
|
||
<paragraph id="78305E07FFAC2179FF09C13E7B91F975" blockId="7.[151,1436,1400,2001]" pageId="7" pageNumber="243">
|
||
<emphasis id="4AFB8215FFAC2179FF09C13E7A9AF92D" bold="true" box="[199,475,1652,1677]" pageId="7" pageNumber="243">Colour in preservative.</emphasis>
|
||
Body creamy white; head and snout with few, scattered dark brown spots; basicaudal bar dissociated into two blotches, one blotch at upper caudal-fin base, larger vertical blotch on lower caudal fin base.
|
||
</paragraph>
|
||
<paragraph id="78305E07FFAC2179FF09C1AA7F1AF871" blockId="7.[151,1436,1400,2001]" pageId="7" pageNumber="243">
|
||
<emphasis id="4AFB8215FFAC2179FF09C1AA7AFCF959" bold="true" box="[199,445,1760,1785]" pageId="7" pageNumber="243">Sexual dimorphism.</emphasis>
|
||
Male with hammer head-shaped suborbital flap (
|
||
<figureCitation id="E0B44282FFAC2179FBDAC1AA7F1FF959" box="[1044,1118,1760,1785]" captionStart="FIGURE 3. A" captionStartId="6.[151,250,1969,1991]" captionTargetBox="[216,1367,195,1940]" captionTargetId="figure@6.[216,1367,195,1940]" captionTargetPageId="6" captionText="FIGURE 3. A) Ventral view of head of Physoschistura chulabhornae, MARNM 3999 / 7, 32.37 mm SL; B) lateral view of head of Physoschistura chulabhornae. showing suborbital flap of male, MARNM 3999 / 20, 30.03 mm SL." httpUri="https://zenodo.org/record/215707/files/figure.png" pageId="7" pageNumber="243">Fig. 3</figureCitation>
|
||
b), and stiff first branched pectoral-fin ray with numerous rows of unculi on upper surface. Pectoral-fin length slightly longer or same length as head (vs. shorter in female); tip of pectoral fin length reaches pelvic-fin origin (vs. not reaching in female). Some females have a small groove in same position as suborbital flab in male. Fewer lateral blotches in large male, changing to longitudinal stripe along lateral line. Male has more slender body; body depth/body width at dorsal-fin origin
|
||
<quantity id="BF77F3E2FFAC2179FF2CC0DF7A12F80D" box="[226,339,1940,1965]" metricMagnitude="-2" metricUnit="m" metricValue="4.445" metricValueMax="5.334" metricValueMin="3.5559999999999996" pageId="7" pageNumber="243" unit="in" value="1.75" valueMax="2.1" valueMin="1.4">1.4–2.1 in</quantity>
|
||
male vs.
|
||
<quantity id="BF77F3E2FFAC2179FE0FC0DE7972F80D" box="[449,563,1940,1965]" metricMagnitude="-2" metricUnit="m" metricValue="3.175" metricValueMax="4.064" metricValueMin="2.286" pageId="7" pageNumber="243" unit="in" value="1.25" valueMax="1.6" valueMin="0.9">0.9–1.6 in</quantity>
|
||
female (
|
||
<tableCitation id="350D6BBCFFAC2179FD58C0DE79AAF80D" box="[662,747,1940,1965]" captionStart="TABLE 1" captionStartId="2.[151,239,620,643]" captionText="TABLE 1. Morphometric data on the holotype and 42 paratypes of Physoschistura chulabhornae, sp. nov." pageId="7" pageNumber="243">Table 1</tableCitation>
|
||
). Pelvic-fin/SL 4.3–5.0 in males vs.
|
||
<quantity id="BF77F3E2FFAC2179FB40C0DE7E41F80D" box="[1166,1280,1940,1965]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.3716" metricValueMin="1.1683999999999999" pageId="7" pageNumber="243" unit="in" value="5.0" valueMax="5.4" valueMin="4.6">4.6–5.4 in</quantity>
|
||
females. One female (MARNM 3999/14 30.0 mm SL) contained 260 large eggs (
|
||
<quantity id="BF77F3E2FFAC2179FC45C0F278A3F870" box="[907,994,1976,2001]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="243" unit="mm" value="0.6">0.6 mm</quantity>
|
||
diameter
|
||
<emphasis id="4AFB8215FFAC2179FB85C0F27F1AF871" bold="true" box="[1099,1115,1976,2001]" pageId="7" pageNumber="243">).</emphasis>
|
||
</paragraph>
|
||
<caption id="2CF00E8FFFA32176FF59C2807FD6FA40" box="[151,1175,1482,1504]" httpUri="https://zenodo.org/record/215709/files/figure.png" pageId="8" pageNumber="244" targetBox="[339,1246,187,1470]" targetPageId="8">
|
||
<paragraph id="78305E07FFA32176FF59C2807FD6FA40" blockId="8.[151,1175,1482,1504]" box="[151,1175,1482,1504]" pageId="8" pageNumber="244">
|
||
<emphasis id="4AFB8215FFA32176FF59C2807A53FA40" bold="true" box="[151,274,1482,1504]" pageId="8" pageNumber="244">FIGURE 6.</emphasis>
|
||
Map of Maechaem River basin indicating type locality of
|
||
<taxonomicName id="BF8F2584FFA32176FCAAC2807FD0FA40" box="[868,1169,1482,1504]" class="Actinopterygii" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="244" phylum="Chordata" rank="species" species="chulabhornae">
|
||
<emphasis id="4AFB8215FFA32176FCAAC2807FD0FA40" box="[868,1169,1482,1504]" italics="true" pageId="8" pageNumber="244">Physoschistura chulabhornae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="78305E07FFA32176FF09C14D7F6EF8C4" blockId="8.[151,1437,1543,2037]" pageId="8" pageNumber="244">
|
||
<emphasis id="4AFB8215FFA32176FF09C14D7A6BF980" bold="true" box="[199,298,1543,1568]" pageId="8" pageNumber="244">Habitat.</emphasis>
|
||
Specimens were collected from the Huay Maetalanoi, Ban Maetala, T. Maedaed, Maechaem District, Chiangmai (
|
||
<figureCitation id="E0B44282FFA32176FEEDC1667A2BF9E5" box="[291,362,1580,1605]" captionStart="FIGURE 6" captionStartId="8.[151,250,1482,1504]" captionTargetBox="[339,1246,187,1470]" captionTargetId="figure@8.[339,1247,185,1470]" captionTargetPageId="8" captionText="FIGURE 6. Map of Maechaem River basin indicating type locality of Physoschistura chulabhornae." httpUri="https://zenodo.org/record/215709/files/figure.png" pageId="8" pageNumber="244">Fig. 6</figureCitation>
|
||
). The stream has a mixed sand, gravel, pebble, silt, mud and rock bottom, with clear running water (
|
||
<figureCitation id="E0B44282FFA32176FF2BC1057A6AF9C8" box="[229,299,1615,1640]" captionStart="FIGURE 7" captionStartId="9.[151,250,1221,1243]" captionTargetBox="[151,1436,223,1187]" captionTargetId="figure@9.[151,1436,223,1187]" captionTargetPageId="9" captionText="FIGURE 7. Huay Maetalanoi, Ban Maetala, T. Maedaed, Maechaem District, Chiangmai, Province, Thailand. Type locality of Physoschistura chulabhornae." httpUri="https://zenodo.org/record/215710/files/figure.png" pageId="8" pageNumber="244">Fig. 7</figureCitation>
|
||
). Elevation at the
|
||
<typeStatus id="A734E0A5FFA32176FE34C11A796BF9C8" box="[506,554,1616,1640]" pageId="8" pageNumber="244">type</typeStatus>
|
||
locality is
|
||
<quantity id="BF77F3E2FFA32176FD67C10579B0F9C7" box="[681,753,1615,1640]" metricMagnitude="2" metricUnit="m" metricValue="7.64" pageId="8" pageNumber="244" unit="m" value="764.0">764 m</quantity>
|
||
above mean sea level, and annual rainfall is over
|
||
<quantity id="BF77F3E2FFA32176FAE7C1057ED9F9C7" box="[1321,1432,1615,1640]" metricMagnitude="0" metricUnit="m" metricValue="1.45" pageId="8" pageNumber="244" unit="mm" value="1450.0">1450 mm</quantity>
|
||
. The surrounding landscape has semi-evergreen forest, plantations, and agricultural crops (mainly corn, cabbage and tomato). Water temperature was 17°C, air temperature 20.5°C, pH 7.7, DO
|
||
<quantity id="BF77F3E2FFA32176FBEEC1DD7F30F910" box="[1056,1137,1687,1712]" metricMagnitude="-6" metricUnit="kg" metricValue="6.5" pageId="8" pageNumber="244" unit="mg" value="6.5">6.5 mg</quantity>
|
||
/l, and conductivity 20 ΜS. Stream width averaged 4 meters, and water depth
|
||
<quantity id="BF77F3E2FFA32176FD0AC1F67877F974" box="[708,822,1724,1749]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="8" pageNumber="244" unit="cm" value="50.0" valueMax="60.0" valueMin="40.0">40–60 cm</quantity>
|
||
. All specimens were collected during the daylight. The
|
||
<typeStatus id="A734E0A5FFA32176FF59C1957BB8F958" box="[151,249,1759,1784]" pageId="8" pageNumber="244" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="A734E0A5FFA32176FE8EC1AA7AEAF958" box="[320,427,1760,1784]" pageId="8" pageNumber="244" type="paratype">paratypes</typeStatus>
|
||
were caught by electro-fishing. Other species collected at the
|
||
<typeStatus id="A734E0A5FFA32176FB75C1AA7FAAF958" box="[1211,1259,1760,1784]" pageId="8" pageNumber="244">type</typeStatus>
|
||
locality were:
|
||
<taxonomicName id="BF8F2584FFA32176FF59C04E7A02F8BD" box="[151,323,1796,1821]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="244" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
:
|
||
<taxonomicName id="BF8F2584FFA32176FE97C04E797EF8BC" box="[345,575,1796,1820]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="244" phylum="Chordata" rank="species" species="breviceps">
|
||
<emphasis id="4AFB8215FFA32176FE97C04E797EF8BC" box="[345,575,1796,1820]" italics="true" pageId="8" pageNumber="244">Schistura breviceps</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="BF8F2584FFA32176FD96C04E7827F8BC" box="[600,870,1796,1820]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="244" phylum="Chordata" rank="species" species="maejotigrina">
|
||
<emphasis id="4AFB8215FFA32176FD96C04E7827F8BC" box="[600,870,1796,1820]" italics="true" pageId="8" pageNumber="244">Schistura maejotigrina</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="BF8F2584FFA32176FCB1C04E78ADF8BD" box="[895,1004,1796,1821]" class="Actinopterygii" family="Sisoridae" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="244" phylum="Chordata" rank="family">Sisoridae</taxonomicName>
|
||
:
|
||
<taxonomicName id="BF8F2584FFA32176FBCFC04E7FBCF8BC" box="[1025,1277,1796,1820]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="244" phylum="Chordata" rank="species" species="siamensis">
|
||
<emphasis id="4AFB8215FFA32176FBCFC04E7FBCF8BC" box="[1025,1277,1796,1820]" italics="true" pageId="8" pageNumber="244">Oreoglanis siamensis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="BF8F2584FFA32176FAD6C04E7ED8F8BD" box="[1304,1433,1796,1821]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="244" phylum="Chordata" rank="family">Cyprinidae</taxonomicName>
|
||
:
|
||
<taxonomicName id="BF8F2584FFA32176FF59C0627BB2F89F" box="[151,243,1832,1855]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="244" phylum="Chordata" rank="genus">
|
||
<emphasis id="4AFB8215FFA32176FF59C0627BB2F89F" box="[151,243,1832,1855]" italics="true" pageId="8" pageNumber="244">Devario</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName id="BF8F2584FFA32176FEE2C06279CDF8E0" box="[300,652,1832,1856]" class="Actinopterygii" family="Cyprinidae" genus="Scaphiodonichthys" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="244" phylum="Chordata" rank="species" species="burmanicus">
|
||
<emphasis id="4AFB8215FFA32176FEE2C06279CDF8E0" box="[300,652,1832,1856]" italics="true" pageId="8" pageNumber="244">Scaphiodonichthys burmanicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="BF8F2584FFA32176FD52C062783EF8E0" box="[668,895,1832,1856]" class="Actinopterygii" family="Cyprinidae" genus="Opsarius" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="244" phylum="Chordata" rank="species" species="pulchellus">
|
||
<emphasis id="4AFB8215FFA32176FD52C062783EF8E0" box="[668,895,1832,1856]" italics="true" pageId="8" pageNumber="244">Opsarius pulchellus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="BF8F2584FFA32176FC41C0627FC5F8E0" box="[911,1156,1832,1856]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="244" phylum="Chordata" rank="species" species="cambodgiensis">
|
||
<emphasis id="4AFB8215FFA32176FC41C0627FC5F8E0" box="[911,1156,1832,1856]" italics="true" pageId="8" pageNumber="244">Garra cambodgiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="BF8F2584FFA32176FB5AC0627E76F8E0" box="[1172,1335,1832,1856]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="244" phylum="Chordata" rank="species" species="brevis">
|
||
<emphasis id="4AFB8215FFA32176FB5AC0627E76F8E0" box="[1172,1335,1832,1856]" italics="true" pageId="8" pageNumber="244">Puntius brevis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="BF8F2584FFA32176FA88C0627A61F8C4" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="244" phylum="Chordata" rank="species" species="stoliczkanus">
|
||
<emphasis id="4AFB8215FFA32176FA88C0627A61F8C4" italics="true" pageId="8" pageNumber="244">Puntius stoliczkanus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="BF8F2584FFA32176FEE0C0067A98F8C5" box="[302,473,1868,1893]" class="Actinopterygii" family="Synbranchidae" kingdom="Animalia" order="Synbranchiformes" pageId="8" pageNumber="244" phylum="Chordata" rank="family">Synbranchidae</taxonomicName>
|
||
:
|
||
<emphasis id="4AFB8215FFA32176FE2AC00779F9F8C4" box="[484,696,1868,1892]" italics="true" pageId="8" pageNumber="244">
|
||
<taxonomicName id="BF8F2584FFA32176FE2AC00779F0F8C4" box="[484,689,1868,1892]" class="Actinopterygii" family="Synbranchidae" genus="Monopterus" kingdom="Animalia" order="Synbranchiformes" pageId="8" pageNumber="244" phylum="Chordata" rank="species" species="albus">Monopterus albus</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="BF8F2584FFA32176FD20C006782BF8C5" box="[750,874,1868,1893]" class="Actinopterygii" family="Channidae" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="244" phylum="Chordata" rank="family">Channidae</taxonomicName>
|
||
:
|
||
<emphasis id="4AFB8215FFA32176FCBBC0067F6EF8C4" box="[885,1071,1868,1892]" italics="true" pageId="8" pageNumber="244">
|
||
<taxonomicName id="BF8F2584FFA32176FCBBC0067F6AF8C4" box="[885,1067,1868,1892]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="244" phylum="Chordata" rank="species" species="gachua">Channa gachua</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="30950D8CFFA32176FF09C0257E2DF80D" pageId="8" pageNumber="244" type="distribution">
|
||
<paragraph id="78305E07FFA32176FF09C0257E2DF80D" blockId="8.[151,1437,1543,2037]" pageId="8" pageNumber="244">
|
||
<emphasis id="4AFB8215FFA32176FF09C0257A21F828" bold="true" box="[199,352,1903,1928]" pageId="8" pageNumber="244">Distribution.</emphasis>
|
||
This species has only been collected at the
|
||
<typeStatus id="A734E0A5FFA32176FC85C03A783AF828" box="[843,891,1904,1928]" pageId="8" pageNumber="244">type</typeStatus>
|
||
locality in the Maechaem River basin (
|
||
<figureCitation id="E0B44282FFA32176FAF8C0257E3BF828" box="[1334,1402,1903,1928]" captionStart="FIGURE 7" captionStartId="9.[151,250,1221,1243]" captionTargetBox="[151,1436,223,1187]" captionTargetId="figure@9.[151,1436,223,1187]" captionTargetPageId="9" captionText="FIGURE 7. Huay Maetalanoi, Ban Maetala, T. Maedaed, Maechaem District, Chiangmai, Province, Thailand. Type locality of Physoschistura chulabhornae." httpUri="https://zenodo.org/record/215710/files/figure.png" pageId="8" pageNumber="244">Fig. 7</figureCitation>
|
||
), a tributary of Ping River, northern Chaophraya River Basin, Maechaem District, Chiangmai Province,
|
||
<collectingCountry id="00981E97FFA32176FACEC0DE7E26F80D" box="[1280,1383,1940,1965]" name="Thailand" pageId="8" pageNumber="244">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="30950D8CFFA32176FF09C0FD7BA2F855" pageId="8" pageNumber="244" type="etymology">
|
||
<paragraph id="78305E07FFA32176FF09C0FD7BA2F855" blockId="8.[151,1437,1543,2037]" pageId="8" pageNumber="244">
|
||
<emphasis id="4AFB8215FFA32176FF09C0FD7A0FF870" bold="true" box="[199,334,1975,2000]" pageId="8" pageNumber="244">Etymology.</emphasis>
|
||
The species epithet honours Her Royal Highness Princess Chulabhorn for her valuable scientific works.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |