184 lines
19 KiB
XML
184 lines
19 KiB
XML
<document id="C770EAF5CF37414CF6A25925C9F2D20C" ID-CLB-Dataset="28501" ID-DOI="http://dx.doi.org/10.3897/zookeys.826.26488" ID-GBIF-Dataset="9d62af83-e4f5-423b-8e12-7dd567f69921" ID-PMC="PMC6405737" ID-Pensoft-Pub="1313-2970-826-1" ID-PubMed="30858752" ID-ZooBank="B1550A3AE54744509A44BC4366D5E110" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-826-1" ModsDocOrigin="ZooKeys 826" ModsDocTitle="Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records" checkinTime="1551446249571" checkinUser="pensoft" docAuthor="Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z." docDate="2019" docId="EFD1115C0F557520C3F834CCE8C085AB" docLanguage="en" docName="ZooKeys 826: 1-343" docOrigin="ZooKeys 826" docSource="http://dx.doi.org/10.3897/zookeys.826.26488" docTitle="Canthidium (Neocanthidium) coerulescens Balthasar 1939" docType="treatment" docVersion="7" lastPageNumber="33" masterDocId="FFECFFBAFFFC2D25D00B9843FFD67749" masterDocTitle="Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records" masterLastPageNumber="343" masterPageNumber="1" pageNumber="33" updateTime="1701379963039" updateUser="ExternalLinkService">
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<mods:title id="A4DFC09F706CBAE58AF76FF5CA6FD7EE">Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records</mods:title>
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<mods:namePart id="D88BB193175AC80495905EA071283526">Chamorro, William</mods:namePart>
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<mods:namePart id="3C3B1AA039C7D036717E1BA13942AE48">Marin-Armijos, Diego</mods:namePart>
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<mods:namePart id="CBF60089A5338823A0DA237756AE4763">senjo, Angelico</mods:namePart>
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<mods:name id="C40B80711613670FACFC02A93222F706" type="personal">
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<mods:namePart id="8DBB8A5C950C1D896D30CA9CD41C049D">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="EFCCF90620EA9106531E2DFEB62E7089">text</mods:typeOfResource>
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<mods:date id="31FEBD51F6077B96E495E43016B10BD0">2019</mods:date>
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<mods:number id="2182DC278F53B4256F25FB31DA431150">826</mods:number>
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<mods:classification id="92131AE3FBF1081713AB58DBDB4F15F3">journal article</mods:classification>
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<mods:identifier id="660E89AE6DB16955665EEF890A3A9800" type="DOI">http://dx.doi.org/10.3897/zookeys.826.26488</mods:identifier>
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</mods:mods>
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<treatment id="EFD1115C0F557520C3F834CCE8C085AB" ID-GBIF-Taxon="154127594" LSID="urn:lsid:plazi:treatment:EFD1115C0F557520C3F834CCE8C085AB" httpUri="http://treatment.plazi.org/id/EFD1115C0F557520C3F834CCE8C085AB" lastPageNumber="33" pageId="32" pageNumber="33">
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<subSubSection id="9830761A4D0B3C35E45A25B228C8E097" pageId="32" pageNumber="33" type="nomenclature">
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<paragraph id="99E2424D0E283D10EF1D2DF34764DE99" pageId="32" pageNumber="33">
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<taxonomicName id="39BA9E2B725BCDF5066E331A81FE9D32" ID-CoL="5WXGD" authority="Balthasar, 1939" authorityName="Balthasar" authorityYear="1939" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium (Neocanthidium) coerulescens" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="coerulescens" subGenus="Neocanthidium">Canthidium (Neocanthidium) coerulescens Balthasar, 1939</taxonomicName>
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Plates 10B, 55D
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</paragraph>
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</subSubSection>
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<subSubSection id="4D401C38FED1D6BF602BF03AE90D8EFC" pageId="32" pageNumber="33" type="reference_group">
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<paragraph id="BB28BA687B6BF76D93F65D6DE8EE133D" pageId="32" pageNumber="33">
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<taxonomicName id="4AF7EC86FD4DECA74BF318ED160F877D" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium coerulescens" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="coerulescens">Canthidium coerulescens</taxonomicName>
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Balthasar, 1939c: 117 (original description. Type locality: Ecuador, Ostcordillieren, Teremotillo, Jivaria, Santa
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<normalizedToken id="B01BED8D56F00B3EC790B4AB75C713DC" originalValue="Inéz">Inez</normalizedToken>
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[= Santa
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<normalizedToken id="688865FBC04FCE23AC43CEB909B3CBE9" originalValue="Inés">Ines</normalizedToken>
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]).
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</paragraph>
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<paragraph id="636B05A0B11AEC1D5D597469591D27FF" pageId="32" pageNumber="33">
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<taxonomicName id="FC31EABDD5A92F0E2F68CFA89CAA9361" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium coerulescens" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="coerulescens">Canthidium coerulescens</taxonomicName>
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:
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<bibRefCitation id="3D2EA68DC1C271A094F4F8D498DE978D" pageId="32" pageNumber="33">Krajcik 2012</bibRefCitation>
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: 62 (complete list of species);
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<bibRefCitation id="7975E4B19CE6F9CDAF499D4FA9779F75" author="Ratcliffe, BC" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="276" pageNumber="277" pagination="186 - 207" title="Beetles (Coleoptera) of Peru: A Survey of the Families." url="https://doi.org/10.2317/kent-88-02-186-207.1" volume="88" year="2015">Ratcliffe et al. 2015</bibRefCitation>
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: 196 (cited for Peru).
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</paragraph>
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<paragraph id="EAFB91B3A66E4A2E9775835BB7677232" pageId="32" pageNumber="33">
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<taxonomicName id="7276AEF2186804ABDAA41D5462FAF316" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium (Canthidium) coerulescens" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="coerulescens" subGenus="Canthidium">Canthidium (Canthidium) coerulescens</taxonomicName>
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:
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<bibRefCitation id="C7EABF0768046B14886966FB0E444281" author="Martinez, A" journalOrPublisher="Acta Zoologica Mexicana" pageId="273" pageNumber="274" pagination="19 - 40" title="Situacion del genero Canthidium." volume="18" year="1986">
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<normalizedToken id="F5D82D957B89B226BEF6C979ED83D6F3" originalValue="Martínez">Martinez</normalizedToken>
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and Halffter 1986
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</bibRefCitation>
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: 26 (transferred to the subgenus
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<taxonomicName id="B314386DE5C9AF23A2A9F6F3FFE81634" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium (Canthidium)" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Canthidium">Canthidium (Canthidium)</taxonomicName>
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s. str., cited for Ecuador);
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<bibRefCitation id="ECD96D5F3A84C421CE5ADF260EE939DB" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
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: 318-319 (cited for Ecuador);
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<bibRefCitation id="97109086B28ABDB399DE80B64A9A3795" author="Bezdek, A" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="262" pageNumber="263" pagination="297 - 334" title="Catalogue of type specimens of beetles deposited in the National Museum, Prague, Czech Republic. Scarabaeidae: Scarabaeinae: Coprini, Eurysternini, Gymnopleurini and Oniticellini." volume="52" year="2012">Bezdek and Hajek 2012</bibRefCitation>
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: 300 (catalog of types NMPC).
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</paragraph>
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<paragraph id="01B9210E95516EDF41B1DF40D9914645" pageId="32" pageNumber="33">
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<taxonomicName id="A2F92013CF7722255008B03A788AEF89" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium (Neocanthidium) coerulescens" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="coerulescens" subGenus="Neocanthidium">Canthidium (Neocanthidium) coerulescens</taxonomicName>
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:
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<bibRefCitation id="0DC259A785F1B4F50ED9E8AF67BA39D8" author="Vulcano, MA" journalOrPublisher="Atas do Simposio sobre a Biota Amazonica" pageId="279" pageNumber="280" pagination="533 - 603" title="Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera)." volume="5" year="1967">Vulcano and Pereira 1967</bibRefCitation>
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: 592 (characters in key. Transferred to the genus
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<taxonomicName id="6AFAE110238BAC6E6F896B054B1EA701" class="Insecta" family="Scarabaeidae" genus="Neocanthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neocanthidium" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Neocanthidium</taxonomicName>
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<normalizedToken id="85676366E9EC21E4BBB16654098E8A15" originalValue="Martínez">Martinez</normalizedToken>
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, Halffter & Pereira, 1964); Cupello, 2018: 468 (distribution, cited for Ecuador);
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<bibRefCitation id="3CFF75B06B9BA2FBE47AF2A40128E848" author="Chamorro, W" journalOrPublisher="Revista Colombiana de Entomologi ́ a" pageId="264" pageNumber="265" pagination="72 - 100" title="Listado de especies y clave de ge ́ neros y subge ́ neros de escarabajos estercoleros (Coleoptera: Scarabaei- dae: Scarabaeinae) presentes y presuntos para Ecuador." url="https://doi.org/10.25100/socolen.v44i1.6545" volume="44" year="2018">Chamorro et al. 2018</bibRefCitation>
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: 89 (figure 12C), 92 (cited for Ecuador).
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</paragraph>
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</subSubSection>
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<subSubSection id="BC46931457E2687215CB863F03995B5B" pageId="32" pageNumber="33" type="type specimens">
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<paragraph id="3158994EAF16CFFB94B0E06224F3346D" pageId="32" pageNumber="33">Type specimens.</paragraph>
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<paragraph id="EB2C3AF1ED66579FB77C4A07F0B3FC02" pageId="32" pageNumber="33">
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<taxonomicName id="4867DA630B42F1BC732C579765993632" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium coerulescens" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="coerulescens">Canthidium coerulescens</taxonomicName>
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Balthasar, 1939. The lectotype (♂) (here designated) and one paralectotype are deposited at the NMPC (ex coll. V Balthasar). Locality: Jivaria, examined.
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</paragraph>
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<paragraph id="B64C91E9F4CFEC4A07B88B9D5B707BB3" pageId="32" pageNumber="33">
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Lectotype (here designated) (♂): "O. Ecuador / Jivaria / 17 12. 05 F. Ohs.S. [p]", "Typus [p, red label, black margin]", "LECTOTYPE ♂ /
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<taxonomicName id="CF33B862F148392AC2E4D9A0CB4C2BAB" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Canthidium</taxonomicName>
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/
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<taxonomicName id="DE88634342E46943F19CEE4326708A15" lsidName="coerulescens" pageId="32" pageNumber="33" rank="species" species="coerulescens">coerulescens</taxonomicName>
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/ Balth. / des. F.Z. Vaz-de-Mello, 2014 [hw and p, red label, black margin]".
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</paragraph>
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<paragraph id="35F5BAE0D2287789C249A9CE5A92C69A" pageId="32" pageNumber="33">
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Paralectotype (♂): "O. Ecuador / Jivaria / 17 12. 05 F. Ohs.S. [p]", "Typus [p, red label, black margin]", "PARALECTOTYPE /
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<taxonomicName id="FE036B15534695AD9C550ADE1D7FD0D8" class="Insecta" family="Scarabaeidae" genus="Canthidium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Canthidium" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Canthidium</taxonomicName>
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♂ /
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<taxonomicName id="DCB81E08B40E4460E399859A78176074" lsidName="coerulescens" pageId="32" pageNumber="33" rank="species" species="coerulescens">coerulescens</taxonomicName>
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Balth / des. F.Z. Vaz-de-Mello, 2013 [hw and p, yellow label, black margin]".
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</paragraph>
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</subSubSection>
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<subSubSection id="CCFD92CFA2B6D8969EF4BC2E7E2DB739" pageId="32" pageNumber="33" type="distribution">
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<paragraph id="9B165385A0809B21AE385DA2A1DDE4CA" pageId="32" pageNumber="33">Distribution.</paragraph>
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<paragraph id="2F90B921AE908C84F4B9490E26649521" pageId="32" pageNumber="33">Ecuador, Peru, and Venezuela.</paragraph>
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</subSubSection>
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<subSubSection id="E809049820E44B5F4BAE690CD0BB37E9" pageId="32" pageNumber="33" type="records examined">
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<paragraph id="CD8D86790DA9D8E970E6C2AA01D50F83" pageId="32" pageNumber="33">Records examined.</paragraph>
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<paragraph id="1C1F795E145DACBA4A56D3EEB075B08D" pageId="32" pageNumber="33">
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<normalizedToken id="5A7322EBB150B95BA7266A1B63A93694" originalValue="SUCUMBÍOS">SUCUMBIOS</normalizedToken>
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:
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<normalizedToken id="BF00AC59E88142350B6BC4A30BB1B11D" originalValue="Río">Rio</normalizedToken>
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Azuela Reventador, 1720 m (1 specimen CEMT); La
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<normalizedToken id="DAF3897CEE692F415AA90C9E2F14FE39" originalValue="Sofía">Sofia</normalizedToken>
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, 1800 m (2 specimens MUTPL). TUNGURAHUA:
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<normalizedToken id="11CD90006378A00178EAFFCBB5C24E3E" originalValue="Baños">Banos</normalizedToken>
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El Topo, 1590 m (6 specimens CEMT; 8 specimens MUTPL). ZAMORA CHINCHIPE: Chito
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<normalizedToken id="8EB64C3012861D937E5CB14A5ADE53CA" originalValue="Río">Rio</normalizedToken>
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San Francisco, 1800 m (2 specimens MUTPL); RVS El Zarza
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<normalizedToken id="83F3D6707B2CC6DA3D59B1AB1C3A1223" originalValue="conseción">consecion</normalizedToken>
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El Zarza, campamento las
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<normalizedToken id="7EB34730E1AB0E52933DB006CDF63FCB" originalValue="Peñas">Penas</normalizedToken>
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, Cordillera del
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<normalizedToken id="8C384E567173DE9DE117B8F3FF21B242" originalValue="Cóndor">Condor</normalizedToken>
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, 1710 m (2 specimens MUTPL); Tundayme campamento Mirador
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<normalizedToken id="2D47B31A3F303AC5492478869C7B8805" originalValue="Jardín">Jardin</normalizedToken>
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<normalizedToken id="54F8C14F3776AF308E272432FE4605B8" originalValue="Botánico">Botanico</normalizedToken>
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, 925 m (1 specimen CEMT); Tundayme campamento Mirador Escombrera Norte, 1225 m (2 specimens).
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</paragraph>
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</subSubSection>
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<subSubSection id="F98C8DC575869D536F81DE52B205B8A3" pageId="32" pageNumber="33" type="literature records">
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<paragraph id="D8A46C6AC7FF325415AD02323F94F152" pageId="32" pageNumber="33">Literature records.</paragraph>
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<paragraph id="45B5B40BD8D9356ABBF35028E3DD543B" pageId="32" pageNumber="33">
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UNDETERMINED PROVINCE: Teremotillo (
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<bibRefCitation id="196637A482CC96EE341D35663F64A570" author="Balthasar, V" journalOrPublisher="Entomologische Nachrichten" pageId="261" pageNumber="262" pagination="111 - 140" title="Neue Canthidium arten." volume="13" year="1939 c">Balthasar 1939c</bibRefCitation>
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: 118); Jivaria (
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<bibRefCitation id="4EDF821B8A41FB3BE55EE8014FAB9901" author="Balthasar, V" journalOrPublisher="Entomologische Nachrichten" pageId="261" pageNumber="262" pagination="111 - 140" title="Neue Canthidium arten." volume="13" year="1939 c">Balthasar 1939c</bibRefCitation>
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: 118;
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<bibRefCitation id="FA4AE46ACAEE85B465D9B8272671B8C0" author="Bezdek, A" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="262" pageNumber="263" pagination="297 - 334" title="Catalogue of type specimens of beetles deposited in the National Museum, Prague, Czech Republic. Scarabaeidae: Scarabaeinae: Coprini, Eurysternini, Gymnopleurini and Oniticellini." volume="52" year="2012">Bezdek and Hajek 2012</bibRefCitation>
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: 300); Santa
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<normalizedToken id="EFD2E92F46FC2847B195F1AF4E2E5B8B" originalValue="Jnéz">Jnez</normalizedToken>
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[= Santa
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<normalizedToken id="B03E168D496A03D3AD8A3FC1723DC7D1" originalValue="Inés">Ines</normalizedToken>
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] (
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<bibRefCitation id="1CA693314402B681C7FD6B573E147F36" author="Balthasar, V" journalOrPublisher="Entomologische Nachrichten" pageId="261" pageNumber="262" pagination="111 - 140" title="Neue Canthidium arten." volume="13" year="1939 c">Balthasar 1939c</bibRefCitation>
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: 118;
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<bibRefCitation id="51EB98C2CFEF700338B70684D7FB5CCD" author="Bezdek, A" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="262" pageNumber="263" pagination="297 - 334" title="Catalogue of type specimens of beetles deposited in the National Museum, Prague, Czech Republic. Scarabaeidae: Scarabaeinae: Coprini, Eurysternini, Gymnopleurini and Oniticellini." volume="52" year="2012">Bezdek and Hajek 2012</bibRefCitation>
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: 300).
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</paragraph>
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</subSubSection>
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<subSubSection id="9689A0A294BC78F1CB77AC2F12169C2E" pageId="32" pageNumber="33" type="temporal data">
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<paragraph id="A132E17C79F5103791087372A0775AD5" pageId="32" pageNumber="33">Temporal data.</paragraph>
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<paragraph id="8E02AA8831286780563F5D0457FA0AD8" pageId="32" pageNumber="33">Collected in January, February, May, August, September, and December.</paragraph>
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</subSubSection>
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<subSubSection id="D7F059659DFC1975A777CB2DF70432DD" pageId="32" pageNumber="33" type="remarks">
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<paragraph id="DD9EBD6AAAD3F0B1F5EB21AFD954614D" pageId="32" pageNumber="33">Remarks.</paragraph>
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<paragraph id="6E762D68D70D946FFBDC72A683E887B3" pageId="32" pageNumber="33">Inhabits lower montane forests and in the montane cloud forests of the Andean regions from 1710-1800 m a.s.l. It was also registered in the evergreen foothill forests and evergreen lower montane forests in the Amazon region from 925-1700 m a.s.l. Collected with pitfall traps baited with human feces.</paragraph>
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<paragraph id="1D1CD4E40BD4EFDEBC8E5D5535FA3354" pageId="32" pageNumber="33">The lectotype is here designated and illustrated (♂ deposited at the NMPC, Plate 56B) and belongs to the locality Ecuador, Jivaria [= eastern slopes of the Andes mountains].</paragraph>
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</subSubSection>
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</treatment>
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</document> |