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<document ID-CLB-Dataset="9127" ID-DOI="http://dx.doi.org/10.3897/zookeys.1008.61040" ID-GBIF-Dataset="6883cbae-4fcb-4a05-ab30-f0673672c13d" ID-PMC="PMC7790813" ID-Pensoft-Pub="1313-2970-1008-61" ID-Pensoft-UUID="F43798A25D4A53C8B58BACF7281D0CBD" ID-PubMed="33505188" ID-ZooBank="E17BB2E712A94F4295956082848B8FC9" ModsDocID="1313-2970-1008-61" checkinTime="1609463508641" checkinUser="pensoft" docAuthor="Tian, Mingyi &amp; He, Li" docDate="2020" docId="EE95CFE4E6B55F048E3EF4D17ED7ECE9" docLanguage="en" docName="ZooKeys 1008: 61-91" docOrigin="ZooKeys 1008" docPubDate="2020-12-31" docSource="http://dx.doi.org/10.3897/zookeys.1008.61040" docTitle="Uenoaphaenops Tian &amp; He 2020, gen. nov." docType="treatment" docUuid="20CF5C96-DFC5-419F-97D8-924D18433C3F" docUuidSource="ZooBank" docVersion="6" id="F43798A25D4A53C8B58BACF7281D0CBD" lastPageNumber="61" masterDocId="F43798A25D4A53C8B58BACF7281D0CBD" masterDocTitle="A contribution to the knowledge of cavernicolous ground beetles from Sichuan Province, southwestern China (Coleoptera, Carabidae, Trechini, Platynini)" masterLastPageNumber="91" masterPageNumber="61" pageNumber="61" updateTime="1732750414890" updateUser="ExternalLinkService">
<mods:mods id="C85BE0BF65B23D7AEC60880CD75DA09C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E32A523C8F7C45D87ADFA7462B6BB4D8">
<mods:title id="E83547F6033DEC60911A47BAF651AAF0">A contribution to the knowledge of cavernicolous ground beetles from Sichuan Province, southwestern China (Coleoptera, Carabidae, Trechini, Platynini)</mods:title>
</mods:titleInfo>
<mods:name id="6D354F1ADDF53118EEB2810F0B1E5B39" type="personal">
<mods:role id="8C6D2F5324A05DC992A82D1C772CB323">
<mods:roleTerm id="255F63505D6E7520B73060AE876AD513">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3E0802C02ACF22F943653CE644E3A642">Tian, Mingyi</mods:namePart>
<mods:nameIdentifier id="757A20E02FA8D0E680FC02DFA1DE455E" type="ORCID">https://orcid.org/0000-0003-2823-7619</mods:nameIdentifier>
<mods:affiliation id="E51739D36A210B48D4451E859B34C1E7">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, China</mods:affiliation>
<mods:nameIdentifier id="81DB885FA0E6D4A1ED06D0C2A3C9B18A" type="email">mytian168@aliyun.com</mods:nameIdentifier>
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<mods:name id="0B6404C974BEDDEA670F3C73CA45EDDD" type="personal">
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</mods:role>
<mods:namePart id="69489DCB251D87C7A3422C8935112CBB">He, Li</mods:namePart>
<mods:nameIdentifier id="CB551BDD1D4948518E528E65B7FAE2E3" type="ORCID">https://orcid.org/0000-0002-4597-3442</mods:nameIdentifier>
<mods:affiliation id="A28BFADD287E11EDB80774A2E11AF7D6">Sichuan Cave Exploration Team, No. 66, 5 th Shuangcheng Road, Chenghua District, Chengdu, 610051, China</mods:affiliation>
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<mods:title id="3B79B8C5A8E5D3C9FF2C17BE6C426F07">ZooKeys</mods:title>
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<mods:part id="D53377AF0AF7572460F0A0B89EF82D21">
<mods:date id="3DFE1D1E32B88701AED6C0985ECF1527">2020</mods:date>
<mods:detail id="E0E817D60350E5CE623D3A3AE0FF7213" type="pubDate">
<mods:number id="88B63852201C0C79920062C660D5C790">2020-12-31</mods:number>
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<mods:detail id="D96FBE93EABBC4D5D6DE37CF3ED22DED" type="volume">
<mods:number id="A31602FB61A095CA73D25171FA12A9A9">1008</mods:number>
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<mods:start id="EF93892C6783CC988230E6AEB1030784">61</mods:start>
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<mods:url id="C485F27FE5E283FAE5D55F91DA9E5179">http://dx.doi.org/10.3897/zookeys.1008.61040</mods:url>
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<mods:classification id="E59CA03800481BE64F608A9C99E10065">journal article</mods:classification>
<mods:identifier id="36B984FBB6CC70C3E5DADCF4D7CBBD66" type="DOI">http://dx.doi.org/10.3897/zookeys.1008.61040</mods:identifier>
<mods:identifier id="426862AA5F28F209F169E82E19F9FF08" type="Pensoft-Pub">1313-2970-1008-61</mods:identifier>
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<treatment id="EE95CFE4E6B55F048E3EF4D17ED7ECE9" ID-GBIF-Taxon="176352580" LSID="urn:lsid:zoobank.org:act:20CF5C96-DFC5-419F-97D8-924D18433C3F" httpUri="http://treatment.plazi.org/id/EE95CFE4E6B55F048E3EF4D17ED7ECE9" lastPageNumber="61" pageId="0" pageNumber="61" scope_family="Carabidae" scope_order="Coleoptera" scope_tribe="Trechini">
<subSubSection id="F62A996A2599D98A2ED212F59444468B" pageId="0" pageNumber="61" type="nomenclature">
<paragraph id="B9EBABFC5BC80E5D0C455A22620419F0" pageId="0" pageNumber="61">
<taxonomicName id="6353B16F4C485AA696F8EE91BB4288B0" ID-CoL="8LYP2" LSID="http://zoobank.org/20CF5C96-DFC5-419F-97D8-924D18433C3F" authority="Tian &amp; He, 2020" authorityName="Tian &amp; He" authorityYear="2020" class="Insecta" family="Carabidae" genus="Uenoaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uenoaphaenops" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus" status="gen. nov.">Uenoaphaenops</taxonomicName>
<taxonomicNameLabel id="2DB6762BD1C63169D86A4928259DF784" pageId="0" pageNumber="61">gen. nov.</taxonomicNameLabel>
<figureCitation id="F0D92945E2413A332E868976F024F46E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Map of Sichuan Province showing the locations of the related caves." figureDoi="10.3897/zookeys.1008.61040.figure2" httpUri="https://binary.pensoft.net/fig/492963" pageId="0" pageNumber="61">
Chinese name:
<normalizedToken id="6A6D643C63CCFA5AC48EA332AE6A2590" originalValue="上野穴步甲属">上野穴步甲属</normalizedToken>
Figs 2
</figureCitation>
<figureCitation id="000BCC04F80D467B72D942FE5F3AC92F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus of Uenoaphaenops fani (Ueno, 2003), comb. nov., female." figureDoi="10.3897/zookeys.1008.61040.figure3" httpUri="https://binary.pensoft.net/fig/492964" pageId="0" pageNumber="61">, 3</figureCitation>
<figureCitation id="1BDB41650634270E702A9590D139CC8A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cave trechine beetles A ventral head of Uenoaphaenops fani (Ueno, 2003), comb. nov., female B ventral head of Chu pheggomisetoides gen. nov. &amp; sp. nov., female C-E elytral chaetotaxy of Uenoaphaenops fani (Ueno, 2003), comb. nov., female, Chu pheggomisetoides gen. nov. &amp; sp. nov., female paratype, and Boreaphaenops liyuani sp. nov., female holotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.61040.figure4" httpUri="https://binary.pensoft.net/fig/492965" pageId="0" pageNumber="61">, 4A, C</figureCitation>
<figureCitation id="018BE8583BD4A3F31BECF5B644DABE2F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia of cave trechine species, lateral and dorsal views A, B Uenoaphaenops fani (Ueno, 2003), comb. nov. C, D Chu pheggomisetoides gen. nov. &amp; sp. nov." figureDoi="10.3897/zookeys.1008.61040.figure5" httpUri="https://binary.pensoft.net/fig/492966" pageId="0" pageNumber="61">, 5A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E0394D9186BA7C2981D6A65A6024D72A" pageId="0" pageNumber="61" type="type species">
<paragraph id="16E5D94C5CD72DA914A9699E2C0AF73E" pageId="0" pageNumber="61">Type species.</paragraph>
<paragraph id="391D37671BAAC3352D27640663DE4A34" pageId="0" pageNumber="61">
<taxonomicName id="C0BDCE373ACEF3BB6ADD00C102C27005" authorityName="Ueno" authorityYear="2003" class="Insecta" family="Carabidae" genus="Qianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qianotrechus fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis id="8069DB8A4F055AA6EC62C4361E2E4987" italics="true" pageId="0" pageNumber="61">Qianotrechus fani</emphasis>
</taxonomicName>
<normalizedToken id="3E2352ACDB3F110E73C89E47C1A34565" originalValue="Uéno">Ueno</normalizedToken>
, 2003 from the cave
<normalizedToken id="8DBED793061AC3CE0A19BDB60774B53E" originalValue="Hua’er">Hua'er</normalizedToken>
Dong, Gulin, southeastern Sichuan).
</paragraph>
<caption id="D1DFE75C224FC2A698B0CDE083D3833F" doi="10.3897/zookeys.1008.61040.figure3" httpUri="https://binary.pensoft.net/fig/492964" pageId="0" pageNumber="61" start="Figure 3" startId="F3">
<paragraph id="659DFB68C903627B15BFE5127C8DAA3C" pageId="0" pageNumber="61">
<emphasis id="89C5662548D9E317D91A2FEBB25B4BB7" bold="true" pageId="0" pageNumber="61">Figure 3.</emphasis>
Habitus of
<taxonomicName id="2C5299955A811CDFF354281812659946" authorityName="Tian &amp; He" authorityYear="2020" baseAuthorityName="Ueno" baseAuthorityYear="2003" class="Insecta" family="Carabidae" genus="Uenoaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uenoaphaenops fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis id="23A622A1657838EBABCDAF5AD59C7BB3" italics="true" pageId="0" pageNumber="61">Uenoaphaenops fani</emphasis>
</taxonomicName>
(
<normalizedToken id="EE6393A99D17B7689D8D0433EB67F2F5" originalValue="Uéno">Ueno</normalizedToken>
, 2003), comb. nov., female.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="49C0BBABE4A24BB4D8D2EE6A409297B6" pageId="0" pageNumber="61" type="description">
<paragraph id="33E8D097C0DCB5A9474023FE4B6D1387" pageId="0" pageNumber="61">Generic characteristics.</paragraph>
<paragraph id="5C4D29D92048A310246E871F7B220DD3" pageId="0" pageNumber="61">
Medium-sized, aphaenopsian and depigmented; body moderately elongate, wholly pubescent. Head strongly elongate, much longer than wide, nearly parallel-sided; neck weakly-marked, ring-shaped; two pairs of supraorbital setiferous pores present; frontal furrows incomplete, parallel-sided in most part though briefly divergent posteriorly; frons and vertex convex; right mandibular tooth bidentate; mentum and submentum completely fused; mentum bisetose, and covered with short setae on basal area of mental tooth, and along the site of labial suture; base largely concave, uni-foveate, tooth short and simple at tip; submentum with a row of 12 setae; labial palpomere 2 much longer than 3 (1.3 times), bisetose on inner margin, without additional setae; maxillary palpomere 3 much longer than 4 (1.4 times) (Fig.
<figureCitation id="5A51C5CBFACD203B1D232E7DEF08D8BB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cave trechine beetles A ventral head of Uenoaphaenops fani (Ueno, 2003), comb. nov., female B ventral head of Chu pheggomisetoides gen. nov. &amp; sp. nov., female C-E elytral chaetotaxy of Uenoaphaenops fani (Ueno, 2003), comb. nov., female, Chu pheggomisetoides gen. nov. &amp; sp. nov., female paratype, and Boreaphaenops liyuani sp. nov., female holotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1008.61040.figure4" httpUri="https://binary.pensoft.net/fig/492965" pageId="0" pageNumber="61">4A</figureCitation>
); antennae long and thin, extending to about 1/4 of elytra from apex; one pair of suborbital pores present. Prothorax distinctly expanded, propleura visible from above; pronotum elongate, as long as head excluding mandibles, wider than head, widest at about 1/3 from front, lateral margins sinuate before hind angles which is nearly rectangular, base slightly narrower than front; two pairs of latero-marginal setae present, disc moderately convex. Elytra ovate, shoulders obtuse, almost rounded, distinctly serrate at prehumeral part, while ciliate on other parts; disc extraordinarily convex though depressed near base; striae reduced though traceable; presence of two dorsal pores along the 3rd striae and the preapical; prehumeral set of the marginal umbilicate pores not aggregated, the 5th pore much closer to 4th than to 6th. Protarsi not modified in male; tibiae without longitudinal sulci.
<taxonomicName id="A4E7EFF07F6FC76C1626913122F67615" genus="Ventrites" lsidName="Ventrites" pageId="0" pageNumber="61" rank="genus">Ventrites</taxonomicName>
IV-VI each with pair of paramedian setae; ventrite VII quadrisetose in both sexes.
</paragraph>
<caption id="80FFA62713C05F6139558CCEE775425F" doi="10.3897/zookeys.1008.61040.figure4" httpUri="https://binary.pensoft.net/fig/492965" pageId="0" pageNumber="61" start="Figure 4" startId="F4">
<paragraph id="B3A60114E071444C01F2690AB48C853B" pageId="0" pageNumber="61">
<emphasis id="F0095EF1B5395684302F3FB073AB2708" bold="true" pageId="0" pageNumber="61">Figure 4.</emphasis>
Cave trechine beetles
<emphasis id="4919DC22A9CF0CCE504ACC209DA8E948" bold="true" pageId="0" pageNumber="61">A</emphasis>
ventral head of
<taxonomicName id="FA96E19912FBA6FD90133357A310321B" authorityName="Tian &amp; He" authorityYear="2020" baseAuthorityName="Ueno" baseAuthorityYear="2003" class="Insecta" family="Carabidae" genus="Uenoaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uenoaphaenops fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis id="C3DB0FE84A2B41D30DD74DD75DC7BFD3" italics="true" pageId="0" pageNumber="61">Uenoaphaenops fani</emphasis>
</taxonomicName>
(
<normalizedToken id="A5616C28DCCEDE1319BCE4E51BABFCAF" originalValue="Uéno">Ueno</normalizedToken>
, 2003), comb. nov., female
<emphasis id="8FE9920F4BA0BA02D7CEE3DAAF36D385" bold="true" pageId="0" pageNumber="61">B</emphasis>
ventral head of
<taxonomicName id="61ADE504C6EB2D91534B928BAA96834E" authorityName="Tian &amp; He" authorityYear="2020" class="Insecta" family="Carabidae" genus="Chu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chu pheggomisetoides" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="pheggomisetoides">
<emphasis id="3B998C6712014FA55B5FBB14DEBA85FF" italics="true" pageId="0" pageNumber="61">Chu pheggomisetoides</emphasis>
</taxonomicName>
gen. nov. &amp; sp. nov., female
<emphasis id="7A90BBB75BB8552185EA1EBA27E1E019" bold="true" pageId="0" pageNumber="61">C-E</emphasis>
elytral chaetotaxy of
<taxonomicName id="69079FE1744B3CFF9CA103F5F949125F" authorityName="Tian &amp; He" authorityYear="2020" baseAuthorityName="Ueno" baseAuthorityYear="2003" class="Insecta" family="Carabidae" genus="Uenoaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uenoaphaenops fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis id="36FDF4EDEC66CA87D31E3B79AD1880E3" italics="true" pageId="0" pageNumber="61">Uenoaphaenops fani</emphasis>
</taxonomicName>
(
<normalizedToken id="0BE8D47A1DD669F425DE571133D8E5BF" originalValue="Uéno">Ueno</normalizedToken>
, 2003), comb. nov., female,
<taxonomicName id="4FF8C5DBDFEAE9CBA65377847B01AA6E" authorityName="Tian &amp; He" authorityYear="2020" class="Insecta" family="Carabidae" genus="Chu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chu pheggomisetoides" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="pheggomisetoides">
<emphasis id="9D051F9431C92D57EE86F8D720879146" italics="true" pageId="0" pageNumber="61">Chu pheggomisetoides</emphasis>
</taxonomicName>
gen. nov. &amp; sp. nov., female paratype, and
<taxonomicName id="E5C2B6045BEB9BD6772827F40AABED89" authorityName="Tian &amp; He" authorityYear="2020" class="Insecta" family="Carabidae" genus="Boreaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Boreaphaenops liyuani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="liyuani">
<emphasis id="5111894B369A19EDC40925F18857C99F" italics="true" pageId="0" pageNumber="61">Boreaphaenops liyuani</emphasis>
</taxonomicName>
sp. nov., female holotype. Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="27812F256958477BA69925378907EFEB" pageId="0" pageNumber="61">
<emphasis id="BEDC52E75DD4C189CBBB8ABC9551B167" bold="true" pageId="0" pageNumber="61">Male genitalia</emphasis>
(Fig.
<figureCitation id="7B82DE5FB415B8EBB4B11AC7DCD70330" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia of cave trechine species, lateral and dorsal views A, B Uenoaphaenops fani (Ueno, 2003), comb. nov. C, D Chu pheggomisetoides gen. nov. &amp; sp. nov." figureDoi="10.3897/zookeys.1008.61040.figure5" httpUri="https://binary.pensoft.net/fig/492966" pageId="0" pageNumber="61">5A, B</figureCitation>
). Aedeagus very short and small, but thick, weakly sclerotized. The median lobe slightly arcuate at median portion, but strongly sinuate before apex which is obtuse, with a large round opening; base quite large, without a sagittal aileron; inner sac provided with a thick and long copulatory piece, which is about 2/5 as long as aedeagus; in dorsal view, apical lobe gradually contracted towards apex which is broadly rounded; parameres well-developed, but much shorter than the median lobe, truncate at apical margin, each armed with four long setae at apex.
</paragraph>
<caption id="78DD83A077E0463409C2DD7EBC9FC25C" doi="10.3897/zookeys.1008.61040.figure5" httpUri="https://binary.pensoft.net/fig/492966" pageId="0" pageNumber="61" start="Figure 5" startId="F5">
<paragraph id="ECCC9EF2F3DDE57857586CE3953E14B3" pageId="0" pageNumber="61">
<emphasis id="17D2BB2E93861A82A774091AB613F912" bold="true" pageId="0" pageNumber="61">Figure 5.</emphasis>
Male genitalia of cave trechine species, lateral and dorsal views
<emphasis id="E756B524C00400A84EDF35781DB59683" bold="true" pageId="0" pageNumber="61">A, B</emphasis>
<taxonomicName id="7FE8A84E98CECA056EDCA455F037F3AD" authorityName="Tian &amp; He" authorityYear="2020" baseAuthorityName="Ueno" baseAuthorityYear="2003" class="Insecta" family="Carabidae" genus="Uenoaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Uenoaphaenops fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis id="B13D81C16AB4CAFAA8AB7D90EA37F116" italics="true" pageId="0" pageNumber="61">Uenoaphaenops fani</emphasis>
</taxonomicName>
(
<normalizedToken id="BBF32FDD06494268D84F376549CDB442" originalValue="Uéno">Ueno</normalizedToken>
, 2003), comb. nov.
<emphasis id="B0F92F97405E87C732B726A9E50C3F49" bold="true" pageId="0" pageNumber="61">C, D</emphasis>
<taxonomicName id="EACB6F9CDA696FA8CDBA46999C516920" authorityName="Tian &amp; He" authorityYear="2020" class="Insecta" family="Carabidae" genus="Chu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chu pheggomisetoides" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="pheggomisetoides">
<emphasis id="28AEC4822DDFDA4FD34315464A735C82" italics="true" pageId="0" pageNumber="61">Chu pheggomisetoides</emphasis>
</taxonomicName>
gen. nov. &amp; sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="D09BF8F83F5466A21F825625DB0BE1BA" pageId="0" pageNumber="61" type="remarks">
<paragraph id="3C2B82E52EE401EDD32C51E6B540FD8F" pageId="0" pageNumber="61">Remarks.</paragraph>
<paragraph id="3563FD2B01F0901B8AB5E86475EF7530" pageId="0" pageNumber="61">
This peculiar species was put into the genus
<taxonomicName id="7789707F5051C07617884857134B8FD1" authorityName="Ueno" authorityYear="2000" class="Insecta" family="Carabidae" genus="Qianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qianotrechus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="4D33881560209031AD0D977984AA2CF4" italics="true" pageId="0" pageNumber="61">Qianotrechus</emphasis>
</taxonomicName>
<normalizedToken id="F1F3442C278FD2468A3E18B33B8F474E" originalValue="Uéno">Ueno</normalizedToken>
, 2000 due to the lack of a male at that time. However,
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<normalizedToken id="AF1CF2339592D2A702725529BB8866D1" originalValue="Uéno">Ueno</normalizedToken>
(2003)
</bibRefCitation>
pointed out the following characteristics of this species which are not present in the congeners of
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<emphasis id="48636B815179DD4882EFFFFB8167A76D" italics="true" pageId="0" pageNumber="61">Qianotrechus</emphasis>
</taxonomicName>
: body wholly pubescent; humeral margins of elytra strongly serrated; and the 5th pore of marginal umbilicate series forwardly and inwardly shifted, widely distant from the 6th pore. Hence, he mentioned that the above peculiarities may suggest a generic separation of this species from the Guizhou genus. Our examination of male individuals provided further evidence to support his opinion. First, protarsi are not modified in the male of
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<emphasis id="8524484F351FC735B3A0DF6C2A6DA0BE" italics="true" pageId="0" pageNumber="61">Qianotrechus fani</emphasis>
</taxonomicName>
, while the 1st and 2nd protarsomeres are spurred inwards at the apices in all other
<taxonomicName id="F78C879D5EF2FA630FBE4A70B1241BFE" authorityName="Ueno" authorityYear="2000" class="Insecta" family="Carabidae" genus="Qianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qianotrechus" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="E01FB6D2358645C8778CA83E94449360" italics="true" pageId="0" pageNumber="61">Qianotrechus</emphasis>
</taxonomicName>
. Second, ventrite VII is quadrisetose in both sexes in
<taxonomicName id="CECF2F05B512B682F57E64C3536CE73A" authorityName="Ueno" authorityYear="2003" class="Insecta" family="Carabidae" genus="Qianotrechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qianotrechus fani" order="Coleoptera" pageId="0" pageNumber="61" phylum="Arthropoda" rank="species" species="fani">
<emphasis id="4688283C4C7DCED090B76E4EEEBAB5BF" italics="true" pageId="0" pageNumber="61">Qianotrechus fani</emphasis>
</taxonomicName>
, vs. bisetose in males of other
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<emphasis id="8EFAD5002CE3541DFCFADC01D0007525" italics="true" pageId="0" pageNumber="61">Qianotrechus</emphasis>
</taxonomicName>
. Third, the aedeagus of
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<emphasis id="E9704D47B15038A4D1EE2E979CD5D467" italics="true" pageId="0" pageNumber="61">Qianotrechus fani</emphasis>
</taxonomicName>
is very small and stout, not the same type as in other
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<emphasis id="728ECE0687F2D2B515AF5C8E125AC9B0" italics="true" pageId="0" pageNumber="61">Qianotrechus</emphasis>
</taxonomicName>
species, which are always large and elongate (
<bibRefCitation id="C1727CF5F837A2CCBC94ABC13DC476EC" author="Ueno, SI" journalOrPublisher="Journal of the Speleological Society of Japan" pageId="0" pageNumber="61" pagination="1 - 38" refId="B13" refString="Ueno, SI, 2000. New cave trechines (Coleoptera, Trechinae) from northeastern Guizhou, South China. Journal of the Speleological Society of Japan 25: 1 - 38" title="New cave trechines (Coleoptera, Trechinae) from northeastern Guizhou, South China." volume="25" year="2000">
<normalizedToken id="1057CB36C6B9EACE5A638848BE8B6776" originalValue="Uéno">Ueno</normalizedToken>
2000
</bibRefCitation>
,
<bibRefCitation id="89C0195E175BF4FF02147FEEA1590787" author="Ueno, SI" journalOrPublisher="Journal of the Speleological Society of Japan" pageId="0" pageNumber="61" pagination="1 - 11" refId="B15" refString="Ueno, SI, 2003. Two new cave trechines of the genus Qianotrechus (Coleoptera, Trechinae). Journal of the Speleological Society of Japan 28: 1 - 11" title="Two new cave trechines of the genus Qianotrechus (Coleoptera, Trechinae)." volume="28" year="2003">2003</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D35A3525164A7962A52BC0B10AC918F9" pageId="0" pageNumber="61" type="etymology">
<paragraph id="CD6F24B603A43E8E7E2340E35679A465" pageId="0" pageNumber="61">Etymology.</paragraph>
<paragraph id="BC4EC5F223D2DDB4641DE7B91A5BBD86" pageId="0" pageNumber="61">
&quot;
<emphasis id="20E39AA04FA2766775325D18F1FF87DD" italics="true" pageId="0" pageNumber="61">Ueno</emphasis>
&quot;+ &quot;-
<emphasis id="9FA0BA9DAB07E4A980A7323FE0C165FF" italics="true" pageId="0" pageNumber="61">aphaenops</emphasis>
&quot;. Dedicated to the late Dr Shun-Ichi
<normalizedToken id="661FDBE1831C32055D88EE9797CB7577" originalValue="Uéno">Ueno</normalizedToken>
who made a great contribution to the knowledge of Chinese subterranean ground beetles. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="553982BB6789393794AF904CD5E1B08B" pageId="0" pageNumber="61" type="generic range">
<paragraph id="FFBED883585CC148AD725D6554779BB3" pageId="0" pageNumber="61">Generic range.</paragraph>
<paragraph id="E2EAF62938359841C745589BBF9F3365" pageId="0" pageNumber="61">
China (Sichuan) (Fig.
<figureCitation id="96ACF586A337D30D2244E3BAD13F7F74" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Map of Sichuan Province showing the locations of the related caves." figureDoi="10.3897/zookeys.1008.61040.figure2" httpUri="https://binary.pensoft.net/fig/492963" pageId="0" pageNumber="61">2</figureCitation>
). A monospecific genus only recorded from the cave
<normalizedToken id="2957710EDE0DF7BE25C45862E021BDFF" originalValue="Hua’er">Hua'er</normalizedToken>
Dong, Gulin County, southeastern Sichuan.
</paragraph>
</subSubSection>
</treatment>
</document>
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