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<document id="6074A931C8634734158C5339D92B4AE6" ID-CLB-Dataset="264111" ID-DOI="10.5852/ejt.2023.863.2081" ID-GBIF-Dataset="59ed2548-4c91-4485-ae44-08609e1028bb" ID-ISSN="2118-9773" ID-Zenodo-Dep="7767958" ID-ZooBank="10981377-CCE7-4487-A415-4E409E55A507" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1679676818438" checkinUser="felipe" docAuthor="Gibbs, David" docDate="2023" docId="EE3F8791FFCE4C63FE263C11D76FE5B0" docLanguage="en" docName="ejt-863_1-162.pdf" docOrigin="European Journal of Taxonomy 863 (1)" docSource="http://zoobank.org/10981377-cce7-4487-a415-4e409e55a507" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Protypusia zimini Gibbs 2023, gen. et comb. nov." docType="treatment" docVersion="7" lastPageNumber="121" masterDocId="1206FFE9FFB84C1BFFEA3D18D57BE31F" masterDocTitle="A world review of the bee fly tribe Usiini (Diptera, Bombyliidae) - Part 3: Parageron Paramonov s. lat." masterLastPageNumber="162" masterPageNumber="1" pageNumber="119" updateTime="1699477371262" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0E3A055CD5CFE3BD37162BEE4246BD6D">A world review of the bee fly tribe Usiini (Diptera, Bombyliidae) - Part 3: Parageron Paramonov s. lat.</mods:title>
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<mods:namePart id="3B9279C1EC2F5A6FB8C0F0019314E35C">Gibbs, David</mods:namePart>
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<mods:title id="7475218961AAE9261D45883F676EDC69">European Journal of Taxonomy</mods:title>
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<mods:date id="15FC8776A219FFBEEC811E26F1820A98">2023</mods:date>
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<mods:number id="214ED4D23029E9C3AA017A32DF3EBDB6">2023-03-24</mods:number>
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<treatment id="EE3F8791FFCE4C63FE263C11D76FE5B0" ID-DOI="http://doi.org/10.5281/zenodo.7768085" ID-GBIF-Taxon="212472458" ID-Zenodo-Dep="7768085" LSID="urn:lsid:plazi:treatment:EE3F8791FFCE4C63FE263C11D76FE5B0" httpUri="http://treatment.plazi.org/id/EE3F8791FFCE4C63FE263C11D76FE5B0" lastPageId="120" lastPageNumber="121" pageId="118" pageNumber="119">
<subSubSection id="2E8C650CFFCE4C6DFE263C11D113E23C" box="[460,1128,265,291]" pageId="118" pageNumber="119" type="nomenclature">
<paragraph id="66293687FFCE4C6DFE263C11D113E23C" blockId="118.[460,1128,265,327]" box="[460,1128,265,291]" pageId="118" pageNumber="119">
<heading id="3D6181EBFFCE4C6DFE263C11D113E23C" box="[460,1128,265,291]" centered="true" fontSize="11" level="2" pageId="118" pageNumber="119" reason="2">
<taxonomicName id="A1964D04FFCE4C6DFE263C11D6F2E23C" ID-CoL="BDP2L" authority="(Paramonov, 1947)" authorityName="Gibbs" authorityYear="2023" baseAuthorityName="Paramonov" baseAuthorityYear="1947" box="[460,905,265,291]" class="Insecta" family="Bombyliidae" genus="Protypusia" kingdom="Animalia" order="Diptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="zimini" status="gen. et comb. nov.">
<emphasis id="54E2EA95FFCE4C6DFE263C11D7DAE23C" bold="true" box="[460,673,265,291]" italics="true" pageId="118" pageNumber="119">Protypusia zimini</emphasis>
(
<bibRefCitation id="02074B76FFCE4C6DFD5A3C11D6FAE23C" author="Paramonov S. J." box="[688,897,265,291]" pageId="118" pageNumber="119" pagination="207 - 220" refId="ref74939" refString="Paramonov S. J. 1947. Uebersicht der mit der Gattung Usia Latr. (Bombyliidae, Diptera) naechstverwandten Gattungen. Eos 23: 207 - 220. Available from http: // hdl. handle. net / 10261 / 149924 [accessed 25 Jan. 2023]." type="journal article" year="1947">Paramonov, 1947</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFCE4C6DFC7A3C11D113E23C" box="[912,1128,265,291]" pageId="118" pageNumber="119" rank="species">gen. et comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2E8C650CFFCE4C6DFD1A3C34D638E258" box="[752,835,300,327]" pageId="118" pageNumber="119" type="description">
<paragraph id="66293687FFCE4C6DFD1A3C34D638E258" blockId="118.[460,1128,265,327]" box="[752,835,300,327]" pageId="118" pageNumber="119">
<figureCitation id="FEAD2A02FFCE4C6DFD1A3C34D638E258" box="[752,835,300,327]" captionStart="Fig" captionStartId="119.[189,232,1928,1954]" captionTargetBox="[256,1331,275,1877]" captionTargetId="figure-12@119.[241,1347,265,1886]" captionTargetPageId="119" captionText="Fig. 59. Protypusia zimini (Paramonov, 1947) gen. et comb. nov. a. Epiphallic complex ventral. b. Epiphallic complex lateral. c. Gonocoxite ventral. d. Gonocoxite lateral. e. Epandrium dorsal. f. Female genitalia ventral." figureDoi="http://doi.org/10.5281/zenodo.7768061" httpUri="https://zenodo.org/record/7768061/files/figure.png" pageId="118" pageNumber="119">Fig. 59</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2E8C650CFFCE4C6DFF573C76D7D9E298" box="[189,674,365,392]" pageId="118" pageNumber="119" type="reference_group">
<paragraph id="66293687FFCE4C6DFF573C76D7D9E298" blockId="118.[189,674,365,392]" box="[189,674,365,392]" pageId="118" pageNumber="119">
<treatmentCitationGroup id="468611A9FFCE4C6DFF573C76D7D9E298" box="[189,674,365,392]" pageId="118" pageNumber="119">
<treatmentCitation id="E7371096FFCE4C6DFF573C76D7E5E298" author="Paramonov S. J." box="[189,670,365,392]" page="216" pageId="118" pageNumber="119" year="1947">
<taxonomicName id="A1964D04FFCE4C6DFF573C76D7E5E298" ID-CoL="4CVP2" authority="Paramonov, 1947: 216" authorityName="Paramonov" authorityPageNumber="216" authorityYear="1947" box="[189,670,365,392]" class="Insecta" family="Bombyliidae" genus="Parageron" kingdom="Animalia" order="Diptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="zimini">
<emphasis id="54E2EA95FFCE4C6DFF573C76D4F1E298" box="[189,394,366,391]" italics="true" pageId="118" pageNumber="119">Parageron zimini</emphasis>
<bibRefCitation id="02074B76FFCE4C6DFE7B3C76D7E5E298" author="Paramonov S. J." box="[401,670,365,392]" pageId="118" pageNumber="119" pagination="207 - 220" refId="ref74939" refString="Paramonov S. J. 1947. Uebersicht der mit der Gattung Usia Latr. (Bombyliidae, Diptera) naechstverwandten Gattungen. Eos 23: 207 - 220. Available from http: // hdl. handle. net / 10261 / 149924 [accessed 25 Jan. 2023]." type="journal article" year="1947">Paramonov, 1947: 216</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="2E8C650CFFCE4C6DFF573CB7D1CEE2E7" box="[189,1205,431,504]" pageId="118" pageNumber="119" type="etymology">
<paragraph id="66293687FFCE4C6DFF573CB7D43FE2D6" blockId="118.[189,324,431,457]" box="[189,324,431,457]" pageId="118" pageNumber="119">
<heading id="3D6181EBFFCE4C6DFF573CB7D43FE2D6" bold="true" box="[189,324,431,457]" fontSize="11" level="3" pageId="118" pageNumber="119" reason="3">
<emphasis id="54E2EA95FFCE4C6DFF573CB7D43FE2D6" bold="true" box="[189,324,431,457]" pageId="118" pageNumber="119">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="66293687FFCE4C6DFF573CC5D1CEE2E7" blockId="118.[189,1205,477,504]" box="[189,1205,477,504]" pageId="118" pageNumber="119">
Named in memory of Leonid Sergeevich Zimin (1902–1970), who collected the
<typeStatus id="B92D8825FFCE4C6DFB853CC6D1CAE2E7" box="[1135,1201,478,504]" pageId="118" pageNumber="119">types</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="2E8C650CFFCE4C6DFF573F07D736E0F3" pageId="118" pageNumber="119" type="materials_examined">
<paragraph id="66293687FFCE4C6DFF573F07D417E126" blockId="118.[189,364,543,569]" box="[189,364,543,569]" pageId="118" pageNumber="119">
<heading id="3D6181EBFFCE4C6DFF573F07D417E126" bold="true" box="[189,364,543,569]" fontSize="11" level="3" pageId="118" pageNumber="119" reason="3">
<emphasis id="54E2EA95FFCE4C6DFF573F07D417E126" bold="true" box="[189,364,543,569]" pageId="118" pageNumber="119">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="66293687FFCE4C6DFF003F56D743E177" blockId="118.[189,1398,589,686]" box="[234,568,589,616]" pageId="118" pageNumber="119">
<emphasis id="54E2EA95FFCE4C6DFF003F56D41DE177" bold="true" box="[234,358,590,616]" pageId="118" pageNumber="119">
<typeStatus id="B92D8825FFCE4C6DFF003F56D41DE177" box="[234,358,590,616]" pageId="118" pageNumber="119" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated)
</paragraph>
<paragraph id="66293687FFCE4C6DFF573F69D152E1B1" blockId="118.[189,1398,589,686]" pageId="118" pageNumber="119">
<materialsCitation id="D6FE3CDAFFCE4C6DFF573F69D152E1B1" ID-GBIF-Occurrence="4401245574" collectingDate="1930-06-13" collectionCode="ZMHB" collectorName="L. S. Zimin" country="Uzbekistan" determinerName="S. Paramonov &amp; Syntypus" location="Tschangyr" municipality="Bukhara" pageId="118" pageNumber="119" specimenCount="3" specimenCount-male="3" typeStatus="syntype">
<collectingCountry id="1E817617FFCE4C6DFF573F69D408E194" box="[189,371,625,651]" name="Uzbekistan" pageId="118" pageNumber="119">UZBEKISTAN</collectingCountry>
•
<specimenCount id="7090FD0EFFCE4C6DFE7B3F69D4D1E194" box="[401,426,625,651]" pageId="118" pageNumber="119" type="male">♂</specimenCount>
; “
<location id="6349605CFFCE4C6DFE293F69D739E194" LSID="urn:lsid:plazi:treatment:EE3F8791FFCE4C63FE263C11D76FE5B0:6349605CFFCE4C6DFE293F69D739E194" box="[451,578,625,651]" country="Uzbekistan" municipality="Bukhara" name="Tschangyr" pageId="118" pageNumber="119">Tschangyr</location>
, northwest
<collectingMunicipality id="864DACFDFFCE4C6DFD273F69D648E194" box="[717,819,625,651]" pageId="118" pageNumber="119">Bukhara</collectingMunicipality>
,
<date id="12281047FFCE4C6DFCA83F69D698E194" box="[834,995,625,651]" pageId="118" pageNumber="119" value="1930-06-13">
<collectingDate id="026CE9AFFFCE4C6DFCA83F69D698E194" box="[834,995,625,651]" pageId="118" pageNumber="119" value="1930-06-13">13 June 1930</collectingDate>
</date>
<specimenCount id="7090FD0EFFCE4C6DFC073F69D160E194" box="[1005,1051,625,651]" pageId="118" pageNumber="119" type="male">1 ♂</specimenCount>
(leg.
<collectorName id="CB635351FFCE4C6DFB8E3F69D193E193" box="[1124,1256,625,652]" pageId="118" pageNumber="119">L.S. Zimin</collectorName>
)/
<taxonomicName id="A1964D04FFCE4C6DFB123F6AD47EE1B1" authority="Gibbs, 2023" authorityName="Gibbs" authorityYear="2023" class="Insecta" family="Bombyliidae" genus="Parageron" kingdom="Animalia" order="Diptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="zimini" status="sp. nov.">
<emphasis id="54E2EA95FFCE4C6DFB123F6AD47EE1B1" italics="true" pageId="118" pageNumber="119">Parageron zimini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFCE4C6DFEE63F8CD41CE1B1" box="[268,359,660,686]" pageId="118" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
<specimenCount id="7090FD0EFFCE4C6DFE843F8CD4FFE1B1" box="[366,388,660,686]" pageId="118" pageNumber="119" type="male">♂</specimenCount>
<determinerName id="93F5DA43FFCE4C6DFE613F8CD74BE1B1" box="[395,560,660,686]" pageId="118" pageNumber="119">S. Paramonov</determinerName>
det.
<typeStatus id="B92D8825FFCE4C6DFD833F8CD7ACE1B1" box="[617,727,660,686]" pageId="118" pageNumber="119" type="syntype">
<determinerName id="93F5DA43FFCE4C6DFD833F8CD7ACE1B1" box="[617,727,660,686]" pageId="118" pageNumber="119">Syntypus</determinerName>
</typeStatus>
/Zool. Mus. Berlin.”
<collectionCode id="0087AE42FFCE4C6DFC273F8CD15DE1B1" box="[973,1062,660,686]" pageId="118" pageNumber="119">ZMHB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="66293687FFCE4C6DFF003FCDD4D9E1F0" blockId="118.[189,1398,725,858]" box="[234,418,725,751]" pageId="118" pageNumber="119">
<emphasis id="54E2EA95FFCE4C6DFF003FCDD4D9E1F0" bold="true" box="[234,418,725,751]" pageId="118" pageNumber="119">
<typeStatus id="B92D8825FFCE4C6DFF003FCDD4D9E1F0" box="[234,418,725,751]" pageId="118" pageNumber="119" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="66293687FFCE4C6DFF573FE1D770E045" blockId="118.[189,1398,725,858]" pageId="118" pageNumber="119">
<materialsCitation id="D6FE3CDAFFCE4C6DFF573FE1D17EE00C" ID-GBIF-Occurrence="4401245547" box="[189,1029,760,787]" collectingDate="1930-06-10" collectionCode="ZMHB" collectorName="L. S. Zimin" country="Uzbekistan" determinerName="S. Paramonov &amp; Syntypus" location="Tschangyr" municipality="Bukhara" pageId="118" pageNumber="119" specimenCount="2" specimenCount-female="2" typeStatus="paralectotype">
<collectingCountry id="1E817617FFCE4C6DFF573FE1D408E00C" box="[189,371,761,787]" name="Uzbekistan" pageId="118" pageNumber="119">UZBEKISTAN</collectingCountry>
•
<specimenCount id="7090FD0EFFCE4C6DFE7B3FE1D4ACE00C" box="[401,471,761,787]" pageId="118" pageNumber="119" type="female">2 ♀♀</specimenCount>
; “locality as lectotype
<date id="12281047FFCE4C6DFD053FE1D6EDE00C" box="[751,918,761,787]" pageId="118" pageNumber="119" value="1930-06-10">
<collectingDate id="026CE9AFFFCE4C6DFD053FE1D6EDE00C" box="[751,918,761,787]" pageId="118" pageNumber="119" value="1930-06-10">10 June 1930</collectingDate>
</date>
”;
<collectionCode id="0087AE42FFCE4C6DFC5A3FE1D17EE00C" box="[944,1029,761,787]" pageId="118" pageNumber="119">ZMHB</collectionCode>
</materialsCitation>
•
<materialsCitation id="D6FE3CDAFFCE4C6DFBC93FE1D751E029" ID-GBIF-Occurrence="4401245496" collectingDate="1930-06-13" collectionCode="ZMHB" collectorName="L. S. Zimin" country="Uzbekistan" determinerName="S. Paramonov &amp; Syntypus" location="Tschangyr" municipality="Bukhara" pageId="118" pageNumber="119" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="7090FD0EFFCE4C6DFBC93FE1D129E00C" box="[1059,1106,761,787]" pageId="118" pageNumber="119" type="male">1 ♂</specimenCount>
[dissected],
<specimenCount id="7090FD0EFFCE4C6DFB063FE1D067E00C" box="[1260,1308,761,787]" pageId="118" pageNumber="119" type="female">1 ♀</specimenCount>
; “same locality
<date id="12281047FFCE4C6DFEF63E04D4C5E029" box="[284,446,796,822]" pageId="118" pageNumber="119" value="1930-06-13">
<collectingDate id="026CE9AFFFCE4C6DFEF63E04D4C5E029" box="[284,446,796,822]" pageId="118" pageNumber="119" value="1930-06-13">13 June 1930</collectingDate>
</date>
”;
<collectionCode id="0087AE42FFCE4C6DFE3F3E04D751E029" box="[469,554,796,822]" pageId="118" pageNumber="119">ZMHB</collectionCode>
</materialsCitation>
•
<materialsCitation id="D6FE3CDAFFCE4C6DFDA93E04D16EE029" ID-GBIF-Occurrence="4401245487" box="[579,1045,796,822]" collectingDate="1930-06-17" collectionCode="ZMHB" collectorName="L. S. Zimin" country="Uzbekistan" determinerName="S. Paramonov &amp; Syntypus" location="Tschangyr" municipality="Bukhara" pageId="118" pageNumber="119" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<specimenCount id="7090FD0EFFCE4C6DFDA93E04D70BE029" box="[579,624,796,822]" pageId="118" pageNumber="119" type="female">1 ♀</specimenCount>
; “same data
<date id="12281047FFCE4C6DFCED3E04D6D2E029" box="[775,937,796,822]" pageId="118" pageNumber="119" value="1930-06-17">
<collectingDate id="026CE9AFFFCE4C6DFCED3E04D6D2E029" box="[775,937,796,822]" pageId="118" pageNumber="119" value="1930-06-17">17 June 1930</collectingDate>
</date>
”;
<collectionCode id="0087AE42FFCE4C6DFC2A3E04D16EE029" box="[960,1045,796,822]" pageId="118" pageNumber="119">ZMHB</collectionCode>
</materialsCitation>
•
<materialsCitation id="D6FE3CDAFFCE4C6DFBC73E04D77CE045" ID-GBIF-Occurrence="4401245303" collectingDate="1930-06-20" collectionCode="ZMHB" collectorName="L. S. Zimin" country="Uzbekistan" determinerName="S. Paramonov &amp; Syntypus" location="Tschangyr" municipality="Bukhara" pageId="118" pageNumber="119" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paralectotype">
<specimenCount id="7090FD0EFFCE4C6DFBC73E04D13CE029" box="[1069,1095,796,822]" pageId="118" pageNumber="119" type="male">♂</specimenCount>
<specimenCount id="7090FD0EFFCE4C6DFBAD3E04D122E029" box="[1095,1113,796,822]" pageId="118" pageNumber="119" type="female">♀</specimenCount>
in copula,
<specimenCount id="7090FD0EFFCE4C6DFB363E04D064E029" box="[1244,1311,796,822]" pageId="118" pageNumber="119" type="female">2 ♀♀</specimenCount>
; “same data
<date id="12281047FFCE4C6DFF1F3E27D4ECE046" box="[245,407,831,858]" pageId="118" pageNumber="119" value="1930-06-20">
<collectingDate id="026CE9AFFFCE4C6DFF1F3E27D4ECE046" box="[245,407,831,858]" pageId="118" pageNumber="119" value="1930-06-20">20 June 1930</collectingDate>
</date>
”;
<collectionCode id="0087AE42FFCE4C6DFE443E58D77CE045" box="[430,519,832,858]" pageId="118" pageNumber="119">ZMHB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="66293687FFCE4C6DFF573E98D482E085" blockId="118.[189,505,896,922]" box="[189,505,896,922]" pageId="118" pageNumber="119">
<heading id="3D6181EBFFCE4C6DFF573E98D482E085" bold="true" box="[189,505,896,922]" fontSize="11" level="3" pageId="118" pageNumber="119" reason="3">
<emphasis id="54E2EA95FFCE4C6DFF573E98D482E085" bold="true" box="[189,505,896,922]" pageId="118" pageNumber="119">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="66293687FFCE4C6DFF573EB7D736E0F3" blockId="118.[189,1398,943,1005]" pageId="118" pageNumber="119">
<materialsCitation id="D6FE3CDAFFCE4C6DFF573EB7D732E0F3" ID-GBIF-Occurrence="4401245607" collectingDate="2009-06-28" collectingDateMax="2009-07-07" collectingDateMin="2009-06-28" collectionCode="PCDG" collectorName="Leg. B. Gharali" country="Iran" elevation="1682" location="Ghazvin" municipality="Zereshk" pageId="118" pageNumber="119" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingCountry id="1E817617FFCE4C6DFF573EB7D47EE0D6" box="[189,261,943,969]" name="Iran" pageId="118" pageNumber="119">IRAN</collectingCountry>
•
<specimenCount id="7090FD0EFFCE4C6DFEF53EB7D435E0D6" box="[287,334,943,969]" pageId="118" pageNumber="119" type="male">1 ♂</specimenCount>
,
<specimenCount id="7090FD0EFFCE4C6DFEB63EB7D4F1E0D6" box="[348,394,943,969]" pageId="118" pageNumber="119" type="female">1 ♀</specimenCount>
; “
<location id="6349605CFFCE4C6DFE493EB7D777E0D6" LSID="urn:lsid:plazi:treatment:EE3F8791FFCE4C63FE263C11D76FE5B0:6349605CFFCE4C6DFE493EB7D777E0D6" box="[419,524,943,969]" country="Iran" municipality="Zereshk" name="Ghazvin" pageId="118" pageNumber="119">Ghazvin</location>
, road to
<collectingMunicipality id="864DACFDFFCE4C6DFD9E3EB7D7A9E0D6" box="[628,722,943,969]" pageId="118" pageNumber="119">Zereshk</collectingMunicipality>
<quantity id="A16E9B62FFCE4C6DFD313EB7D64CE0D6" box="[731,823,943,969]" metricMagnitude="3" metricUnit="m" metricValue="1.682" pageId="118" pageNumber="119" unit="m" value="1682.0">
<elevation id="EDBBD1B4FFCE4C6DFD313EB7D64CE0D6" box="[731,823,943,969]" metricMagnitude="3" metricUnit="m" metricValue="1.682" pageId="118" pageNumber="119" unit="m" value="1682.0">1682 m</elevation>
</quantity>
, N36°24’128” E50°5’434”
<date id="12281047FFCE4C6DFB953EB7D009E0D6" box="[1151,1394,943,969]" pageId="118" pageNumber="119" value="2009-06-28" valueMax="2009-07-07" valueMin="2009-06-28">
<collectingDate id="026CE9AFFFCE4C6DFB953EB7D009E0D6" box="[1151,1394,943,969]" pageId="118" pageNumber="119" value="2009-06-28" valueMax="2009-07-07" valueMin="2009-06-28">28 June-7 July 2009</collectingDate>
</date>
,
<collectingMethod id="BFD74E90FFCE4C6DFF573ECBD466E0F2" box="[189,285,979,1005]" pageId="118" pageNumber="119">pan trap</collectingMethod>
<collectorName id="CB635351FFCE4C6DFECE3ECBD49BE0F3" box="[292,480,978,1005]" pageId="118" pageNumber="119">Leg. B. Gharali</collectorName>
”;
<collectionCode id="0087AE42FFCE4C6DFE1D3ECAD732E0F3" box="[503,585,978,1004]" pageId="118" pageNumber="119">PCDG</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2E8C650CFFCE4C63FF57390BD62CE747" lastPageId="120" lastPageNumber="121" pageId="118" pageNumber="119" type="description">
<paragraph id="66293687FFCE4C6DFF57390BD410E732" blockId="118.[189,363,1043,1069]" box="[189,363,1043,1069]" pageId="118" pageNumber="119">
<heading id="3D6181EBFFCE4C6DFF57390BD410E732" bold="true" box="[189,363,1043,1069]" fontSize="11" level="3" pageId="118" pageNumber="119" reason="3">
<emphasis id="54E2EA95FFCE4C6DFF57390BD410E732" bold="true" box="[189,363,1043,1069]" pageId="118" pageNumber="119">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="66293687FFCE4C6DFF57395BD696E742" blockId="118.[189,1005,1090,1117]" box="[189,1005,1090,1117]" pageId="118" pageNumber="119">
MEASUREMENTS. Body length;
<quantity id="A16E9B62FFCE4C6DFDF7395AD7D6E742" box="[541,685,1090,1117]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.1" metricValueMin="3.9" pageId="118" pageNumber="119" unit="mm" value="4.5" valueMax="5.1" valueMin="3.9">3.9–5.1 mm</quantity>
. Wing length:
<quantity id="A16E9B62FFCE4C6DFCB2395AD693E742" box="[856,1000,1090,1117]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="5.2" metricValueMin="4.1" pageId="118" pageNumber="119" unit="mm" value="4.65" valueMax="5.2" valueMin="4.1">4.1–5.2 mm</quantity>
.
</paragraph>
<paragraph id="66293687FFCE4C6DFF00399CD451E781" blockId="118.[189,1400,1156,1569]" box="[234,298,1156,1182]" pageId="118" pageNumber="119">
<emphasis id="54E2EA95FFCE4C6DFF00399CD451E781" bold="true" box="[234,298,1156,1182]" pageId="118" pageNumber="119">Male</emphasis>
</paragraph>
<paragraph id="66293687FFCE4C6DFF5739BFD4E6E53E" blockId="118.[189,1400,1156,1569]" pageId="118" pageNumber="119">HEAD. Frons and gena pale yellow in ground colour, the frons slightly tumescent covered with silky off-white dusting, lacking longer hairs. Gena moderately broad, at narrowest point about as broad as length of pedicel, more shining mouth margin a little narrower, together almost as broad as scape length. Occiput and ocellar tubercle dark in ground colour densely covered with grey dust and outstanding white hairs, ocelli contiguous with eyes. Eyes confluent for slightly longer than the length of vertex. Ommatidia conspicuously enlarged in the upper half of the eyes, sharply contrasting with the smaller ones occupying the lower half. Antennae blackish, tip of pedicel can be paler, postpedicel rather long with a small point dorsally immediately beyond the subapical sulcus. Palps small and slender, yellowish to brown, darker apically, the white apical setae shorter than the length of the palps. Proboscis very long, about equal to body length, naked dorsally, the basoventral membrane yellow.</paragraph>
<paragraph id="66293687FFCE4C6DFF573B51D4CDE429" blockId="118.[187,1401,1608,1846]" pageId="118" pageNumber="119">
THORAX. Dark ground colour obscured by dove-grey dust, in some becoming more yellowish on posterior of mesonotum and scutellum, very faint darker paramedian vittae on anterior half or slightly more of mesonotum, antehumeral vittae may be indicated. Whole of mesonotum and scutellum evenly covered with moderately long, silky white hair, largely absent on darker vittae, on the front of the mesonotum hairs longer than tibia is thick at its mid-point. Pleura with similar hairs on pronotum, posterior twothirds of the anepisternum, and a few in upper middle of katepisternum. Metepimeron naked, can be yellow (
<collectingCountry id="1E817617FFCE4C6DFEF53A03D4D1E429" box="[287,426,1819,1846]" name="Uzbekistan" pageId="118" pageNumber="119">Uzbekistan</collectingCountry>
).
</paragraph>
<paragraph id="66293687FFCE4C6DFF573A45D7FBE4A2" blockId="118.[188,1398,1885,1982]" pageId="118" pageNumber="119">WING. Membrane hyaline, the veins pale yellow, or becoming brown apically. Crossvein r-m from basal quarter to third of the discal cell, opposite or a little beyond m-cu. Anal lobe broad with conspicuously convex margin, broader than anal cell.</paragraph>
<caption id="32E9660FFFCF4C6CFF573A90D482E4F7" ID-DOI="http://doi.org/10.5281/zenodo.7768061" ID-Zenodo-Dep="7768061" httpUri="https://zenodo.org/record/7768061/files/figure.png" pageId="119" pageNumber="120" startId="119.[189,232,1928,1954]" targetBox="[256,1331,275,1877]" targetPageId="119">
<paragraph id="66293687FFCF4C6CFF573A90D482E4F7" blockId="119.[189,1399,1928,2025]" pageId="119" pageNumber="120">
<emphasis id="54E2EA95FFCF4C6CFF573A90D45AE4BD" bold="true" box="[189,289,1928,1954]" pageId="119" pageNumber="120">Fig. 59.</emphasis>
<taxonomicName id="A1964D04FFCF4C6CFEC53A91D782E4BD" authority="(Paramonov, 1947)" authorityName="Gibbs" authorityYear="2023" baseAuthorityName="Paramonov" baseAuthorityYear="1947" box="[303,761,1928,1954]" class="Insecta" family="Bombyliidae" genus="Protypusia" kingdom="Animalia" order="Diptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="zimini" status="gen. et comb. nov.">
<emphasis id="54E2EA95FFCF4C6CFEC53A91D77FE4BD" box="[303,516,1929,1954]" italics="true" pageId="119" pageNumber="120">Protypusia zimini</emphasis>
(
<bibRefCitation id="02074B76FFCF4C6CFDF03A90D78AE4BD" author="Paramonov S. J." box="[538,753,1928,1954]" pageId="119" pageNumber="120" pagination="207 - 220" refId="ref74939" refString="Paramonov S. J. 1947. Uebersicht der mit der Gattung Usia Latr. (Bombyliidae, Diptera) naechstverwandten Gattungen. Eos 23: 207 - 220. Available from http: // hdl. handle. net / 10261 / 149924 [accessed 25 Jan. 2023]." type="journal article" year="1947">Paramonov, 1947</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFFCF4C6CFCED3A90D688E4BD" box="[775,1011,1928,1954]" pageId="119" pageNumber="120" rank="species">gen. et comb. nov.</taxonomicNameLabel>
<emphasis id="54E2EA95FFCF4C6CFBEA3A90D174E4BD" bold="true" box="[1024,1039,1928,1954]" pageId="119" pageNumber="120">a</emphasis>
. Epiphallic complex ventral.
<emphasis id="54E2EA95FFCF4C6CFF573AB3D5B6E4DA" bold="true" box="[189,205,1963,1989]" pageId="119" pageNumber="120">b</emphasis>
. Epiphallic complex lateral.
<emphasis id="54E2EA95FFCF4C6CFDD03AB3D73CE4DA" bold="true" box="[570,583,1963,1989]" pageId="119" pageNumber="120">c</emphasis>
. Gonocoxite ventral.
<emphasis id="54E2EA95FFCF4C6CFCB03AB3D611E4DA" bold="true" box="[858,874,1963,1989]" pageId="119" pageNumber="120">d</emphasis>
. Gonocoxite lateral.
<emphasis id="54E2EA95FFCF4C6CFB9F3AB3D1F9E4DA" bold="true" box="[1141,1154,1963,1989]" pageId="119" pageNumber="120">e</emphasis>
. Epandrium dorsal.
<emphasis id="54E2EA95FFCF4C6CFF573AD6D5BCE4F7" bold="true" box="[189,199,1998,2024]" pageId="119" pageNumber="120">f</emphasis>
. Female genitalia ventral.
</paragraph>
</caption>
<paragraph id="66293687FFC04C63FF573C12D644E23C" blockId="120.[189,831,265,292]" box="[189,831,265,292]" pageId="120" pageNumber="121">HALTERE. Pale yellow, base of stem slightly infuscated.</paragraph>
<paragraph id="66293687FFC04C63FF573C53D07BE2EE" blockId="120.[189,1399,330,498]" pageId="120" pageNumber="121">
LEGS. Coxae dark with a dense coating of grey dust. Femora predominantly dark with a covering of grey dust, up to apical quarter clear yellow (
<collectingCountry id="1E817617FFC04C63FD3D3C76D626E297" box="[727,861,366,392]" name="Uzbekistan" pageId="120" pageNumber="121">Uzbekistan</collectingCountry>
specimens) or just narrowly yellow (Iranian specimens). Front and mid-tibia clear yellow, hind tibia infuscated apically (
<collectingCountry id="1E817617FFC04C63FB8B3C89D19CE2B4" box="[1121,1255,401,427]" name="Uzbekistan" pageId="120" pageNumber="121">Uzbekistan</collectingCountry>
specimens) or infuscation more extensive (Iranian specimens). Tarsi black with the basitarsi more or less yellow basally in some. Legs covered with silky white hairs, rather variable, longer on Iranian male.
</paragraph>
<paragraph id="66293687FFC04C63FF573F01D6B0E1A0" blockId="120.[187,1399,537,704]" pageId="120" pageNumber="121">
ABDOMEN. Tergite one entirely yellow (
<collectingCountry id="1E817617FFC04C63FD953F01D67EE12C" box="[639,773,537,563]" name="Uzbekistan" pageId="120" pageNumber="121">Uzbekistan</collectingCountry>
specimens) or with just apical margin yellow (Iranian specimens), remaining tergites black with dense grey dusting and sharply demarcated apical yellow margins (
<collectingCountry id="1E817617FFC04C63FEC63F47D4C9E166" box="[300,434,607,633]" name="Uzbekistan" pageId="120" pageNumber="121">Uzbekistan</collectingCountry>
specimens) or poorly defined pale margins (Iranian specimens), these becoming narrower rearwards. Sternites similar but yellow apical margins clearer. Tergites and sternites all covered with fairly long, erect white to very pale yellow, rather wavy hairs.
</paragraph>
<paragraph id="66293687FFC04C63FF573FFFD1C9E058" blockId="120.[189,1400,743,840]" pageId="120" pageNumber="121">GENITALIA. Relatively large when compared to other holoptic species. Gonocoxites black, grey dusted with broad yellow tips, covered in long whitish hairs. Epandrium basally black, grey dusted, with very broad yellow apical margin and narrow lateral margins, covered in short whitish hairs.</paragraph>
<paragraph id="66293687FFC04C63FF003E76D43EE097" blockId="120.[187,1399,878,1113]" box="[234,325,878,904]" pageId="120" pageNumber="121">
<emphasis id="54E2EA95FFC04C63FF003E76D43EE097" bold="true" box="[234,325,878,904]" pageId="120" pageNumber="121">Female</emphasis>
</paragraph>
<paragraph id="66293687FFC04C63FF573E89D62CE747" blockId="120.[187,1399,878,1113]" pageId="120" pageNumber="121">
Differs from the male in its broadly separated eyes, the frons pale yellow and hairless anteriorly, densely pale-grey dusted on hind half, laterally with 1–2 rows of tiny hairs. Legs yellow except for coxae basally (
<collectingCountry id="1E817617FFC04C63FF2E3ECED431E0EF" box="[196,330,982,1008]" name="Uzbekistan" pageId="120" pageNumber="121">Uzbekistan</collectingCountry>
specimens) or similar to the male, grey dusted, more or less infuscated tarsi largely blackish. Abdomen much more extensively yellow than in the male with less well defined blackish-brown bases to the tergites (except T1) and sternites (
<collectingCountry id="1E817617FFC04C63FD733903D664E72A" box="[665,799,1051,1077]" name="Uzbekistan" pageId="120" pageNumber="121">Uzbekistan</collectingCountry>
specimens) or similar to male. Hairs on the thorax and abdomen a little shorter, most clearly on the tergites.
</paragraph>
</subSubSection>
<subSubSection id="2E8C650CFFC04C63FF573967D40DE55F" pageId="120" pageNumber="121" type="discussion">
<paragraph id="66293687FFC04C63FF573967D454E786" blockId="120.[189,303,1151,1177]" box="[189,303,1151,1177]" pageId="120" pageNumber="121">
<heading id="3D6181EBFFC04C63FF573967D454E786" bold="true" box="[189,303,1151,1177]" fontSize="11" level="3" pageId="120" pageNumber="121" reason="3">
<emphasis id="54E2EA95FFC04C63FF573967D454E786" bold="true" box="[189,303,1151,1177]" pageId="120" pageNumber="121">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="66293687FFC04C63FF5739B5D4D5E66B" blockId="120.[188,1401,1197,1397]" pageId="120" pageNumber="121">
Although
<specimenCount id="7090FD0EFFC04C63FEDC39B5D772E7D8" box="[310,521,1197,1223]" pageId="120" pageNumber="121" type="generic">twelve specimens</specimenCount>
were available, they originate from just two widely separated localities, 10 from the
<typeStatus id="B92D8825FFC04C63FEBC39C8D4F2E7F5" box="[342,393,1232,1258]" pageId="120" pageNumber="121">type</typeStatus>
locality and two from
<collectingCountry id="1E817617FFC04C63FD4839C8D7AEE7F5" box="[674,725,1232,1258]" name="Iran" pageId="120" pageNumber="121">Iran</collectingCountry>
. There is quite a degree of variation between these two locations notably in the extent of yellow on tergites and legs.
<collectingCountry id="1E817617FFC04C63FC6B39EAD17EE613" box="[897,1029,1266,1292]" name="Uzbekistan" pageId="120" pageNumber="121">Uzbekistan</collectingCountry>
specimens have much yellower tergites and legs compared to the
<specimenCount id="7090FD0EFFC04C63FDB8380ED679E630" box="[594,770,1301,1328]" pageId="120" pageNumber="121" type="generic">two specimens</specimenCount>
from
<collectingCountry id="1E817617FFC04C63FCA4380ED6FAE62F" box="[846,897,1302,1328]" name="Iran" pageId="120" pageNumber="121">Iran</collectingCountry>
. However, in most features, including the structure of both male and female genitalia, they differ very little so I am as confident as I can be that they are conspecific.
</paragraph>
<paragraph id="66293687FFC04C63FF573883D40DE55F" blockId="120.[189,1399,1435,1600]" pageId="120" pageNumber="121">
The illustrations provided by
<bibRefCitation id="02074B76FFC04C63FDC23883D7A2E6A9" author="Zaitzev V. F." box="[552,729,1435,1462]" pageId="120" pageNumber="121" refId="ref75364" refString="Zaitzev V. F. 1966. [Parasitic Flies of the Family Bombyliidae (Diptera) in the Fauna of Transcaucasia.] Nauka, Moscow &amp; Leningrad. [In Russian.]" type="book" year="1966">Zaitzev (1966)</bibRefCitation>
and ascribed to
<taxonomicName id="A1964D04FFC04C63FC403884D1AEE6AA" authority="(Paramonov)" baseAuthorityName="Paramonov" box="[938,1237,1435,1462]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="grisea">
<emphasis id="54E2EA95FFC04C63FC403884D6AEE6AA" box="[938,981,1436,1461]" italics="true" pageId="120" pageNumber="121">Pro</emphasis>
.
<emphasis id="54E2EA95FFC04C63FC0D3884D14BE6AA" box="[999,1072,1436,1461]" italics="true" pageId="120" pageNumber="121">grisea</emphasis>
(Paramonov)
</taxonomicName>
are certainly not
<taxonomicName id="A1964D04FFC04C63FF0138A7D414E6C7" box="[235,367,1471,1497]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="grisea">
<emphasis id="54E2EA95FFC04C63FF0138A7D46DE6C7" box="[235,278,1471,1496]" italics="true" pageId="120" pageNumber="121">Pro</emphasis>
.
<emphasis id="54E2EA95FFC04C63FECC38A7D414E6C7" box="[294,367,1471,1496]" italics="true" pageId="120" pageNumber="121">grisea</emphasis>
</taxonomicName>
and appear very similar to this species or, perhaps the closely related
<taxonomicName id="A1964D04FFC04C63FB5938A7D041E6C7" authorityName="Gibbs" authorityYear="2023" baseAuthorityName="Zaitzev" baseAuthorityYear="1996" box="[1203,1338,1471,1496]" class="Insecta" family="Bombyliidae" genus="Protypusia" kingdom="Animalia" order="Diptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="negevi">
<emphasis id="54E2EA95FFC04C63FB5938A7D041E6C7" box="[1203,1338,1471,1496]" italics="true" pageId="120" pageNumber="121">Pro. negevi</emphasis>
</taxonomicName>
. The description, while brief, could fit either species, closer to
<taxonomicName id="A1964D04FFC04C63FC8A38F9D69DE6E5" authorityName="Gibbs" authorityYear="2023" baseAuthorityName="Zaitzev" baseAuthorityYear="1996" box="[864,998,1505,1531]" class="Insecta" family="Bombyliidae" genus="Protypusia" kingdom="Animalia" order="Diptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="negevi">
<emphasis id="54E2EA95FFC04C63FC8A38F9D6F0E6E5" box="[864,907,1505,1530]" italics="true" pageId="120" pageNumber="121">Pro</emphasis>
.
<emphasis id="54E2EA95FFC04C63FC7338F9D69DE6E5" box="[921,998,1505,1530]" italics="true" pageId="120" pageNumber="121">negevi</emphasis>
</taxonomicName>
on size but more likely to be
<taxonomicName id="A1964D04FFC04C63FAAE38F9D47EE502" authorityName="Gibbs" authorityYear="2023" baseAuthorityName="Paramonov" baseAuthorityYear="1947" class="Insecta" family="Bombyliidae" genus="Protypusia" kingdom="Animalia" order="Diptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="zimini">
<emphasis id="54E2EA95FFC04C63FAAE38F9D014E6E5" box="[1348,1391,1505,1530]" italics="true" pageId="120" pageNumber="121">Pro</emphasis>
.
<emphasis id="54E2EA95FFC04C63FF573B1CD47EE502" box="[189,261,1540,1565]" italics="true" pageId="120" pageNumber="121">zimini</emphasis>
</taxonomicName>
on range. Without seeing the specimens it is not possible to be confident what species Zaitzev had before him.
</paragraph>
</subSubSection>
<subSubSection id="2E8C650CFFC04C63FF573B7FD76FE5B0" box="[189,532,1639,1712]" pageId="120" pageNumber="121" type="distribution">
<paragraph id="66293687FFC04C63FF573B7FD42DE59E" blockId="120.[189,342,1639,1665]" box="[189,342,1639,1665]" pageId="120" pageNumber="121">
<heading id="3D6181EBFFC04C63FF573B7FD42DE59E" bold="true" box="[189,342,1639,1665]" fontSize="11" level="3" pageId="120" pageNumber="121" reason="3">
<emphasis id="54E2EA95FFC04C63FF573B7FD42DE59E" bold="true" box="[189,342,1639,1665]" pageId="120" pageNumber="121">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="66293687FFC04C63FF573B8DD76FE5B0" blockId="120.[189,532,1685,1712]" box="[189,532,1685,1712]" pageId="120" pageNumber="121">
(?
<collectingCountry id="1E817617FFC04C63FF3A3B8DD44DE5AF" box="[208,310,1685,1712]" name="Georgia" pageId="120" pageNumber="121">Georgia</collectingCountry>
),
<collectingCountry id="1E817617FFC04C63FEA03B8DD407E5B0" box="[330,380,1685,1711]" name="Iran" pageId="120" pageNumber="121">Iran</collectingCountry>
,
<collectingCountry id="1E817617FFC04C63FE623B8DD76BE5B0" box="[392,528,1685,1711]" name="Uzbekistan" pageId="120" pageNumber="121">Uzbekistan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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