476 lines
63 KiB
XML
476 lines
63 KiB
XML
<document id="4E9F0D2EACD717D92F8D1F27DDFC9515" ID="10.11646/zootaxa.4273.3.4SLASH11253" ID-CLB-Dataset="32904" ID-DOI="10.11646/zootaxa.4273.3.4" ID-GBIF-Dataset="c029f78e-c026-48ac-8c61-8b168fef59ea" ID-ISSN="1175-5326" ID-Zenodo-Dep="803109" ID-ZooBank="ADC167F6-C3A0-43EC-8636-F65F5ABE85AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1496736238170" checkinUser="plazi" docAuthor="Rodriguez, Lily O. & Catenazzi, Alessandro" docDate="2017" docId="ED0DF709FFB675439EB473EB95EFF932" docLanguage="en" docName="zootaxa.4273.3.4SLASH11253.pdf" docOrigin="Zootaxa 4273 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phrynopus capitalis Rodriguez & Catenazzi, 2017, new species" docType="treatment" docVersion="8" lastPageNumber="392" masterDocId="11348F71FFBE75489E23770F9474FFF9" masterDocTitle="Four new species of terrestrial-breeding frogs of the genus Phrynopus (Anura: Terrarana: Craugastoridae) from Río Abiseo National Park, Peru" masterLastPageNumber="406" masterPageNumber="381" pageNumber="389" updateTime="1698440199461" updateUser="ExternalLinkService">
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<mods:mods id="ED3322A3DEA57507E5F4B3E9ABB3DA04" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9C34900CFC4DC1E7EF1AD14DAC2FC9BA">
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<mods:title id="78A47D6E38E4769A09794DC6C4DE706B">Four new species of terrestrial-breeding frogs of the genus Phrynopus (Anura: Terrarana: Craugastoridae) from Río Abiseo National Park, Peru</mods:title>
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<mods:name id="0ACFC8F73C29C950FA5BC51D508B1F8C" type="personal">
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<mods:roleTerm id="011071D45ABEC4F264911216427D98D7">Author</mods:roleTerm>
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<mods:namePart id="15E085BFA0B317554F8A1517340EE6B6">Rodriguez, Lily O.</mods:namePart>
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</mods:name>
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<mods:name id="31B9F68A4D45475A4C9E588D35A3D6FA" type="personal">
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<mods:roleTerm id="CE75EC3CC427B986C8D917B10CC8AC8C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="440847531132693B68565A746E45CD67">Catenazzi, Alessandro</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4022F9ABCA69ABFDACA9D40AC649B70C">text</mods:typeOfResource>
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<mods:titleInfo id="CDA9E5D06148DB3920D7136382D40D2B">
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<mods:title id="BE41A1F0F485053682C0A2D07CDC29C2">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="DA4FC1AD68E8646847B5733307954EA7">
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<mods:date id="8F4AFEF993EA81326CB75FA6322F0347">2017</mods:date>
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<mods:detail id="A48F624E6737E77B65FB5E6F5CC3A2BA" type="volume">
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<mods:number id="F2EFC19937A2BA520EA4A04C3F0CFB73">4273</mods:number>
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<mods:detail id="AE7A9FFDFDEB9390496F27DA698F7220" type="issue">
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<mods:number id="1A1C95CF63264B1AEEF7AF482D1222EF">3</mods:number>
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<mods:classification id="B0140B5A0100D9239024ABCCF553BB2B">journal article</mods:classification>
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<mods:identifier id="72AB2043C01A42321597A2975C0D8CA1" type="CLB-Dataset">32904</mods:identifier>
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<mods:identifier id="A1A4C412251F1603720ADBA9634E5286" type="DOI">10.11646/zootaxa.4273.3.4</mods:identifier>
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<mods:identifier id="A524B1F53AEF91520C4E71EB4EBB3999" type="GBIF-Dataset">c029f78e-c026-48ac-8c61-8b168fef59ea</mods:identifier>
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<mods:identifier id="0761766F71FD2874C21D9BFF19353E4C" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="ED0DF709FFB675439EB473EB95EFF932" ID-DOI="http://doi.org/10.5281/zenodo.6033268" ID-GBIF-Taxon="131812331" ID-Zenodo-Dep="6033268" LSID="urn:lsid:plazi:treatment:ED0DF709FFB675439EB473EB95EFF932" httpUri="http://treatment.plazi.org/id/ED0DF709FFB675439EB473EB95EFF932" lastPageId="11" lastPageNumber="392" pageId="8" pageNumber="389">
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<subSubSection id="2DBE1594FFB675409EB473EB95F9FAE6" pageId="8" pageNumber="389" type="nomenclature">
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<paragraph id="651B461FFFB675409EB473EB966AFB07" blockId="8.[151,542,1252,1311]" box="[151,542,1252,1278]" pageId="8" pageNumber="389">
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<heading id="3E53F173FFB675409EB473EB966AFB07" bold="true" box="[151,542,1252,1278]" fontSize="11" level="1" pageId="8" pageNumber="389" reason="1">
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<emphasis id="57D09A0DFFB675409EB473EB966AFB07" bold="true" box="[151,542,1252,1278]" pageId="8" pageNumber="389">
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<taxonomicName id="A2A43D9CFFB675409EB473EB95F3FB07" box="[151,391,1252,1278]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="389" phylum="Chordata" rank="species" species="capitalis" status="sp. nov.">
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<emphasis id="57D09A0DFFB675409EB473EB95F3FB07" bold="true" box="[151,391,1252,1278]" italics="true" pageId="8" pageNumber="389">Phrynopus capitalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="4CE32776FFB675409FAD73EB966AFB07" box="[398,542,1252,1278]" pageId="8" pageNumber="389" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="651B461FFFB675409EB4720995F9FAE6" blockId="8.[151,542,1252,1311]" box="[151,397,1286,1311]" pageId="8" pageNumber="389">
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(
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<figureCitation id="FD9F5A9AFFB675409EBD7209957EFAE6" box="[158,266,1286,1311]" captionStart="FIGURE 6" captionStartId="7.[151,250,1854,1876]" captionTargetBox="[165,1411,981,1831]" captionTargetId="figure@7.[151,1436,972,1832]" captionTargetPageId="7" captionText="FIGURE 6. Palmar and plantar surfaces of the new species of Phrynopus: (A) P. anancites sp. nov. (MUSM 33168), (B) P. capitalis sp. nov. (AMNH 134158), (C) P. dumicola sp. nov. (MUSM 33102), and (D) P. personatus sp. nov. (MUSM 33096). Photographs by M. Flecks (A, C and D) and A. Catenazzi (B)." httpUri="https://zenodo.org/record/803121/files/figure.png" pageId="8" pageNumber="389">Figures 6</figureCitation>
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B, 7, 8, 9A)
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</paragraph>
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</subSubSection>
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<subSubSection id="2DBE1594FFB675409EB472419574FA0E" pageId="8" pageNumber="389" type="materials_examined">
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<paragraph id="651B461FFFB675409EB4724196FDFA56" blockId="8.[151,1436,1358,2031]" pageId="8" pageNumber="389">
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<materialsCitation id="D5CC4C42FFB675409EB4724196FDFA56" ID-GBIF-Occurrence="1563404871" collectingDate="1989-07-03" collectionCode="AMNH" collectorName="NE Pataz & Provincia Pataz & Departamento & L. O. Rodriguez." country="Peru" county="Figure" elevation="3600" latitude="-7.6963887" location="Figures" longLatPrecision="21" longitude="-77.51556" municipality="Region" pageId="8" pageNumber="389" specimenCode="AMNH 134158" specimenCount="1" stateProvince="La Libertad" typeStatus="holotype">
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<emphasis id="57D09A0DFFB675409EB47241957FFA9E" bold="true" box="[151,267,1358,1383]" pageId="8" pageNumber="389">
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<typeStatus id="BA1FF8BDFFB675409EB472419572FA9E" box="[151,262,1358,1383]" pageId="8" pageNumber="389" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="3502EE64FFB675409F32724095B0FA9E" box="[273,452,1358,1383]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="389">
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<collectionCode id="03B5DEDAFFB675409F3272409512FA9E" box="[273,358,1359,1383]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="389">AMNH</collectionCode>
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134158
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</specimenCode>
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, an adult male (
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<figureCitation id="FD9F5A9AFFB675409C57724196ABFA9E" box="[628,735,1358,1383]" captionStart="FIGURE 6" captionStartId="7.[151,250,1854,1876]" captionTargetBox="[165,1411,981,1831]" captionTargetId="figure@7.[151,1436,972,1832]" captionTargetPageId="7" captionText="FIGURE 6. Palmar and plantar surfaces of the new species of Phrynopus: (A) P. anancites sp. nov. (MUSM 33168), (B) P. capitalis sp. nov. (AMNH 134158), (C) P. dumicola sp. nov. (MUSM 33102), and (D) P. personatus sp. nov. (MUSM 33096). Photographs by M. Flecks (A, C and D) and A. Catenazzi (B)." httpUri="https://zenodo.org/record/803121/files/figure.png" pageId="8" pageNumber="389">
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<location id="607B10C4FFB675409C57724196BEFA9E" LSID="urn:lsid:plazi:treatment:ED0DF709FFB675439EB473EB95EFF932:607B10C4FFB675409C57724196BEFA9E" box="[628,714,1358,1383]" country="Peru" county="Figure" latitude="-7.6963887" longLatPrecision="21" longitude="-77.51556" municipality="Region" name="Figures" pageId="8" pageNumber="389" stateProvince="La Libertad">Figures</location>
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6
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</figureCitation>
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B, 7) obtained at
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<location id="607B10C4FFB675409DBD72419013FA9E" LSID="urn:lsid:plazi:treatment:ED0DF709FFB675439EB473EB95EFF932:607B10C4FFB675409DBD72419013FA9E" box="[926,1127,1358,1383]" country="Peru" county="Figure" latitude="-7.6963887" longLatPrecision="21" longitude="-77.51556" municipality="Region" name="Lake Manachaqui" pageId="8" pageNumber="389" stateProvince="La Libertad">Lake Manachaqui</location>
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(
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<geoCoordinate id="009020D8FFB675409A5572409081FA9E" box="[1142,1269,1359,1383]" direction="south" orientation="latitude" pageId="8" pageNumber="389" precision="15" value="-7.6963887">7°41’47” S</geoCoordinate>
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,
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<geoCoordinate id="009020D8FFB675409B21724191ECFA9F" box="[1282,1432,1358,1383]" direction="west" orientation="longitude" pageId="8" pageNumber="389" precision="15" value="-77.51556">77°30’56” W</geoCoordinate>
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,
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<quantity id="A25CEBFAFFB675409EB4727D9480FA72" box="[151,244,1394,1419]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="389" unit="m" value="3600.0">
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<elevation id="EE89A12CFFB675409EB4727D9480FA72" box="[151,244,1394,1419]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="389" unit="m" value="3600.0">3600 m</elevation>
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</quantity>
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), ca.
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||
<quantity id="A25CEBFAFFB675409F1B727C95F0FA72" box="[312,388,1394,1419]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="389" unit="km" value="14.0">14 km</quantity>
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airline
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<collectorName id="C85123C9FFB675409FC7727C9620FA73" box="[484,596,1395,1418]" pageId="8" pageNumber="389">NE Pataz</collectorName>
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||
,
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<collectorName id="C85123C9FFB675409C47727D976FFA73" box="[612,795,1394,1419]" pageId="8" pageNumber="389">Provincia Pataz</collectorName>
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,
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<collectorName id="C85123C9FFB675409D08727C97BEFA73" box="[811,970,1395,1418]" pageId="8" pageNumber="389">Departamento</collectorName>
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||
(
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<collectingMunicipality id="857FDC65FFB675409DFE727D9045FA72" box="[989,1073,1394,1419]" pageId="8" pageNumber="389">Región</collectingMunicipality>
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||
)
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<collectingRegion id="A76088FDFFB675409A66727C90A5FA72" box="[1093,1233,1394,1419]" country="Peru" name="La Libertad" pageId="8" pageNumber="389">La Libertad</collectingRegion>
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||
,
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<collectingCountry id="1DB3068FFFB675409AC3727C9167FA73" box="[1248,1299,1395,1418]" name="Peru" pageId="8" pageNumber="389">Peru</collectingCountry>
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||
(
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<figureCitation id="FD9F5A9AFFB675409B04727D91FFFA72" box="[1319,1419,1394,1419]" captionStart="FIGURE 1" captionStartId="2.[151,250,1830,1852]" captionTargetBox="[189,1411,235,1774]" captionTargetId="figure@2.[175,1427,193,1813]" captionTargetPageId="2" captionText="FIGURE 1. Map of the Andes of central and northern Peru, with type localities of new species Phrynopus capitalis sp. nov., P. dumicola sp. nov., P. personatus sp. nov. (star) and P. anancites sp. nov. (asterisk) and of congeneric forms: P. valquii (asterisk), P. thompsoni (1), P. daemon (2), P. lechriorhynchus and P. vestigiatus (3), P. horstpauli (4), P. kauneorum and P. kotosh (5), P. dagmarae and P. interstinctus (6), P. heimorum (7), P. barthlenae and P. tautzorum (8), P. miroslawae and P. nicoleae (9), P. badius and P. curator (10), P. auriculatus, P. bracki and P. tribulosus (11), P. pesantesi (12), P. bufoides and P. paucari (13), P. juninensis and P. montium (14), P. oblivious and P. peruanus (15), and P. chaparroi (16)." httpUri="https://zenodo.org/record/803111/files/figure.png" pageId="8" pageNumber="389">
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<collectingCounty id="8C7A3E93FFB675409B04727D9106FA72" box="[1319,1394,1394,1419]" pageId="8" pageNumber="389">Figure</collectingCounty>
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1
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</figureCitation>
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), collected on
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<date id="111A60DFFFB675409F05729895DDFA56" box="[294,425,1430,1455]" pageId="8" pageNumber="389" value="1989-07-03">
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<collectingDate id="015E9937FFB675409F05729895DDFA56" box="[294,425,1430,1455]" pageId="8" pageNumber="389" value="1989-07-03">3 July 1989</collectingDate>
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</date>
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by
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<collectorName id="C85123C9FFB675409FF0729896FDFA56" box="[467,649,1430,1455]" pageId="8" pageNumber="389">L.O. Rodríguez.</collectorName>
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</materialsCitation>
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</paragraph>
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<paragraph id="651B461FFFB675409EE472B59574FA0E" blockId="8.[151,1436,1358,2031]" pageId="8" pageNumber="389">
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<emphasis id="57D09A0DFFB675409EE472B5950FFA2A" bold="true" box="[199,379,1466,1491]" pageId="8" pageNumber="389">Paratopotypes.</emphasis>
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MUSM 8959, a gravid female (
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<figureCitation id="FD9F5A9AFFB675409CD072B59723FA2A" box="[755,855,1466,1491]" captionStart="FIGURE 8" captionStartId="11.[151,250,930,952]" captionTargetBox="[340,1247,303,908]" captionTargetId="figure@11.[340,1247,303,908]" captionTargetPageId="11" captionText="FIGURE 8. Female paratopotype of Phrynopus capitalis sp. nov. in life (MUSM 8959, SVL = 35.6 mm) from around Lake Manachaqui, 3600 m, near Rio Abiseo National Park, San Martín, Peru. Photograph by L. O. Rodriguez." httpUri="https://zenodo.org/record/803125/files/figure.png" pageId="8" pageNumber="389">Figure 8</figureCitation>
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), and MUSM 8957, a juvenile, collected with the holotype.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="2DBE1594FFB675419EE4710D9025F8B5" lastPageId="9" lastPageNumber="390" pageId="8" pageNumber="389" type="diagnosis">
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||
<paragraph id="651B461FFFB675419EE4710D9120FF2D" blockId="8.[151,1436,1358,2031]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="390" pageId="8" pageNumber="389">
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<emphasis id="57D09A0DFFB675409EE4710D9535F9E2" bold="true" box="[199,321,1538,1563]" pageId="8" pageNumber="389">Diagnosis.</emphasis>
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A moderately-sized
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<taxonomicName id="A2A43D9CFFB675409C11710D96DFF9E2" box="[562,683,1538,1563]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="389" phylum="Chordata" rank="genus">
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<emphasis id="57D09A0DFFB675409C11710D96DFF9E2" box="[562,683,1538,1563]" italics="true" pageId="8" pageNumber="389">Phrynopus</emphasis>
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</taxonomicName>
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with the following characteristics: (1) skin on dorsum and flanks fleshy and tuberculated, densely covered with minute round tubercles, bearing low warts especially above the insertion of the forelimbs and dorsolaterally; skin on throat, chest and belly coarsely areolate, ventral surface of thighs coarsely areolate; discoidal fold absent, thoracic fold evident; dorsolateral folds absent; (2) tympanic membrane and tympanic annulus absent; supratympanic fold swollen (similar to a parotoid gland); (3) snout short, nearly rounded in dorsal view, sloping and rounded in profile; nasals large, frontoparietals in contact; (4) upper eyelid without tubercles; width of upper eyelid narrower than IOD; cranial crests absent; (5) dentigerous processes of vomers absent; (6) males with gray nuptial pad on thumb, lacking vocal slits and vocal sac; (7) Finger I slightly shorter than Finger II; tips of digits bulbous, terminal phalanges T-shaped (
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||
<figureCitation id="FD9F5A9AFFB675409DD7702D9023F8C2" box="[1012,1111,1826,1851]" captionStart="FIGURE 9" captionStartId="12.[151,250,1242,1264]" captionTargetBox="[357,1211,197,1207]" captionTargetId="figure@12.[340,1247,193,1220]" captionTargetPageId="12" captionText="FIGURE 9. Condition of terminal phalanges of Finger III, left hand: T-shaped in (A) Phrynopus capitalis sp. nov. (MUSM 8959), and knob-shaped in (B) P. dumicola sp. nov. (USNM 299788), (C) P. personatus sp. nov. (MUSM 3813), (D) P. valquii (MUSM 3823). Drawings by F. Chang." httpUri="https://zenodo.org/record/803127/files/figure.png" pageId="8" pageNumber="389">Figure 9</figureCitation>
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A); (8) fingers bearing basal webbing and fleshy lateral fringes; (9) ulnar and tarsal tubercles absent; (10) heels lacking tubercles; inner tarsal fold absent; (11) inner metatarsal tubercle rounded, about 1.5 times larger than rounded outer metatarsal tubercle; supernumerary plantar tubercles absent (12) toes bearing lateral fringes; basal webbing present; Toe V longer than Toe III; toe tips rounded, bulbous, about as large as those on fingers; terminal phalanges with lateral processes (as in
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<figureCitation id="FD9F5A9AFFB675409E9470D9956DF816" box="[183,281,2006,2031]" captionStart="FIGURE 9" captionStartId="12.[151,250,1242,1264]" captionTargetBox="[357,1211,197,1207]" captionTargetId="figure@12.[340,1247,193,1220]" captionTargetPageId="12" captionText="FIGURE 9. Condition of terminal phalanges of Finger III, left hand: T-shaped in (A) Phrynopus capitalis sp. nov. (MUSM 8959), and knob-shaped in (B) P. dumicola sp. nov. (USNM 299788), (C) P. personatus sp. nov. (MUSM 3813), (D) P. valquii (MUSM 3823). Drawings by F. Chang." httpUri="https://zenodo.org/record/803127/files/figure.png" pageId="8" pageNumber="389">Figure 9</figureCitation>
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A) (13) in life, dorsal surfaces are brownish tan with small, dark brown reticulate spots having the appearance of a labyrinth; areolation on ventral surfaces are grayish cream in a darker ground color; in preservative, flanks are blueish black or black; (14) SVL in single female
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<quantity id="A25CEBFAFFB775419DF077B3904DFF2D" box="[979,1081,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.56" pageId="9" pageNumber="390" unit="mm" value="35.6">35.6 mm</quantity>
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, in single male
|
||
<quantity id="A25CEBFAFFB775419ACA77B39124FF2D" box="[1257,1360,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="9" pageNumber="390" unit="mm" value="23.2">23.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="651B461FFFB775419EE477D09025F8B5" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="390">
|
||
The new species cannot be confused with any other described species of
|
||
<taxonomicName id="A2A43D9CFFB775419DDF77D09001FF01" box="[1020,1141,223,248]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="genus">
|
||
<emphasis id="57D09A0DFFB775419DDF77D09001FF01" box="[1020,1141,223,248]" italics="true" pageId="9" pageNumber="390">Phrynopus</emphasis>
|
||
</taxonomicName>
|
||
because of its very robust and prominent head (HW/SVL = 40 %), swollen tympanic area, absence of tympanic annulus, thick lateral fringes on all digits, and its distinctive dorsal coloration pattern with small brown reticulated spots.
|
||
<taxonomicName id="A2A43D9CFFB775419A9A762891E8FEB9" box="[1209,1436,295,320]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="capitalis">
|
||
<emphasis id="57D09A0DFFB775419A9A762891E8FEB9" box="[1209,1436,295,320]" italics="true" pageId="9" pageNumber="390">Phrynopus capitalis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<emphasis id="57D09A0DFFB775419F127642961FFE9D" box="[305,619,332,357]" italics="true" pageId="9" pageNumber="390">
|
||
<taxonomicName id="A2A43D9CFFB775419F12764295A1FE9C" box="[305,469,332,357]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="auriculatus">P. auriculatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419FC47642961FFE9D" box="[487,619,333,356]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="montium">P. montium</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="A2A43D9CFFB775419C8D7642974CFE9D" box="[686,824,333,356]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="peruanus">
|
||
<emphasis id="57D09A0DFFB775419C8D7642974CFE9D" box="[686,824,333,356]" italics="true" pageId="9" pageNumber="390">P. peruanus</emphasis>
|
||
</taxonomicName>
|
||
by lacking a visible tympanic annulus.
|
||
<emphasis id="57D09A0DFFB775419B0076439640FE55" italics="true" pageId="9" pageNumber="390">
|
||
<taxonomicName id="A2A43D9CFFB775419B0076439571FE71" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="anancites">Phrynopus anancites</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419F31767E95C4FE71" box="[274,432,367,392]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="auriculatus">P. auriculatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419F9E767E9650FE71" box="[445,548,367,392]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="bracki">P. bracki</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419C12767E96B7FE71" box="[561,707,367,392]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="dagmarae">P. dagmarae</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419CEC767E970DFE71" box="[719,889,369,392]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="interstinctus">P. interstinctus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419DA5767E9050FE71" box="[902,1060,367,392]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="kauneorum">P. kauneorum</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419A13767E90EDFE71" box="[1072,1177,367,392]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="kotosh">P. kotosh</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419A85767E910FFE71" box="[1190,1403,367,392]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="lechriorhynchus">P. lechriorhynchus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419BAB767E948DFE54" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="nicoleae">P. nicoleae</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419F25769A95E4FE55" box="[262,400,405,428]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="peruanus">P. peruanus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419FBE769A9640FE55" box="[413,564,405,428]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="vestigiatus">P. vestigiatus</taxonomicName>
|
||
</emphasis>
|
||
all differ (character condition for
|
||
<taxonomicName id="A2A43D9CFFB775419DE2769A9034FE54" box="[961,1088,404,429]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="capitalis">
|
||
<emphasis id="57D09A0DFFB775419DE2769A9034FE54" box="[961,1088,404,429]" italics="true" pageId="9" pageNumber="390">P. capitalis</emphasis>
|
||
</taxonomicName>
|
||
in parentheses) by having the dentigerous processes of vomers (absent).
|
||
<emphasis id="57D09A0DFFB775419CAA76B890FCFE29" box="[649,1160,439,464]" italics="true" pageId="9" pageNumber="390">
|
||
<taxonomicName id="A2A43D9CFFB775419CAA76B8972EFE29" box="[649,858,439,464]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="badius">Phrynopus badius</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419D4876B697ABFE29" box="[875,991,441,464]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="curator">P. curator</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419DCD76B690F0FE29" box="[1006,1156,439,464]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="horstpauli">P. horstpauli</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="A2A43D9CFFB775419AE476B69128FE29" box="[1223,1372,439,464]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="thompsoni">
|
||
<emphasis id="57D09A0DFFB775419AE476B69128FE29" box="[1223,1372,439,464]" italics="true" pageId="9" pageNumber="390">P. thompsoni</emphasis>
|
||
</taxonomicName>
|
||
have discontinuous or ridge-like dorsolateral folds (absent). Of the species lacking a tympanum, the dentigerous processes of vomers and dorsolateral folds or ridges,
|
||
<taxonomicName id="A2A43D9CFFB775419D27750E97E8FDE1" box="[772,924,511,536]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="barthlenae">
|
||
<emphasis id="57D09A0DFFB775419D27750E97E8FDE1" box="[772,924,511,536]" italics="true" pageId="9" pageNumber="390">P. barthlenae</emphasis>
|
||
</taxonomicName>
|
||
has similar size, coarsely tuberculate dorsal skin, and lateral fringes on digits, but differs from
|
||
<taxonomicName id="A2A43D9CFFB775419CEF752A973CFDC4" box="[716,840,548,573]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="capitalis">
|
||
<emphasis id="57D09A0DFFB775419CEF752A973CFDC4" box="[716,840,548,573]" italics="true" pageId="9" pageNumber="390">P. capitalis</emphasis>
|
||
</taxonomicName>
|
||
by lacking warts (present), by bearing a dark canthal and postorbital stripe (absent), by having small nasals (large) and widely separated frontoparietals (large nasals, frontoparietals in contact), and distinctive black dorsal blotches (labyrinth-like short spots on dorsal surfaces).
|
||
<taxonomicName id="A2A43D9CFFB775419EB475809508FD51" box="[151,380,655,680]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="bufoides">
|
||
<emphasis id="57D09A0DFFB775419EB475809508FD51" box="[151,380,655,680]" italics="true" pageId="9" pageNumber="390">Phrynopus bufoides</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<emphasis id="57D09A0DFFB775419C4F759E9681FD51" box="[620,757,655,680]" italics="true" pageId="9" pageNumber="390">
|
||
<taxonomicName id="A2A43D9CFFB775419C4F759E9685FD51" box="[620,753,655,680]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="capitalis">P. capitalis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but differs by having a discoidal fold (absent), a broad supratympanic fold (swollen), by lacking lateral fringes on toes (present), and by bearing prominent dorsal isolated and elongated warts forming discontinuous ridges (coarsely areolate dorsal skin with warts and minute tubercles), and black mottling on venter (uniform ventral coloration).
|
||
<taxonomicName id="A2A43D9CFFB775419D6A75F39048FCEC" box="[841,1084,764,789]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="chaparroi">
|
||
<emphasis id="57D09A0DFFB775419D6A75F39048FCEC" box="[841,1084,764,789]" italics="true" pageId="9" pageNumber="390">Phrynopus chaparroi</emphasis>
|
||
</taxonomicName>
|
||
lacks lateral fringes on digits (present), has a much darker coloration without dorsal pattern (tan with brown pattern);
|
||
<taxonomicName id="A2A43D9CFFB775419A8F742E9155FCC1" box="[1196,1313,799,824]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="daemon">
|
||
<emphasis id="57D09A0DFFB775419A8F742E9155FCC1" box="[1196,1313,799,824]" italics="true" pageId="9" pageNumber="390">P. daemon</emphasis>
|
||
</taxonomicName>
|
||
is readily distinguished by having subacuminate snout in lateral view (rounded), short supratympanic fold (swollen), no dorsal pattern (short labyrinth-like lines) and males having prominent subgular vocal sac (absent), and by lacking nuptial pads (present).
|
||
<taxonomicName id="A2A43D9CFFB775419FBD748396E6FC5C" box="[414,658,908,933]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="heimorum">
|
||
<emphasis id="57D09A0DFFB775419FBD748396E6FC5C" box="[414,658,908,933]" italics="true" pageId="9" pageNumber="390">Phrynopus heimorum</emphasis>
|
||
</taxonomicName>
|
||
has a slightly tuberculate dorsum (fleshy and tuberculate) and light reticulations especially on flanks and venter (dorsum with labyrinth-like small spots, venter uniform), and lacks lateral fringes on all digits (present);
|
||
<taxonomicName id="A2A43D9CFFB775419C4074DA968CFC15" box="[611,760,981,1004]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="juninensis">
|
||
<emphasis id="57D09A0DFFB775419C4074DA968CFC15" box="[611,760,981,1004]" italics="true" pageId="9" pageNumber="390">P. juninensis</emphasis>
|
||
</taxonomicName>
|
||
has smooth dorsal and ventral skin (coarsely areolate).
|
||
<taxonomicName id="A2A43D9CFFB775419EB474F895EEFBE9" box="[151,410,1015,1040]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="miroslawae">
|
||
<emphasis id="57D09A0DFFB775419EB474F895EEFBE9" box="[151,410,1015,1040]" italics="true" pageId="9" pageNumber="390">Phrynopus miroslawae</emphasis>
|
||
</taxonomicName>
|
||
differs by lacking lateral fringes on digits and basal webbing on toes (present), by bearing dorsolateral, occipital and supratympanic folds (absent, supratympanic fold swollen), by having a row of subconical tubercles on outer edge of tarsus (absent), and by being smaller with the single known female having SVL =
|
||
<quantity id="A25CEBFAFFB775419EC9736B953BFB85" box="[234,335,1124,1148]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="9" pageNumber="390" unit="mm" value="29.2">29.2 mm</quantity>
|
||
(SVL =
|
||
<quantity id="A25CEBFAFFB775419F90736B966FFB85" box="[435,539,1124,1149]" metricMagnitude="-2" metricUnit="m" metricValue="3.56" pageId="9" pageNumber="390" unit="mm" value="35.6">35.6 mm</quantity>
|
||
).
|
||
<taxonomicName id="A2A43D9CFFB775419C0D736B9763FB85" box="[558,791,1124,1149]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="pesantesi">
|
||
<emphasis id="57D09A0DFFB775419C0D736B9763FB85" box="[558,791,1124,1149]" italics="true" pageId="9" pageNumber="390">Phrynopus pesantesi</emphasis>
|
||
</taxonomicName>
|
||
lacks lateral fringes on digits (present), has a blunt snout (sloping to rounded in lateral profile) and smaller head with HW/SVL = 36% (40% in
|
||
<taxonomicName id="A2A43D9CFFB775419A5773869086FB59" box="[1140,1266,1159,1184]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="capitalis">
|
||
<emphasis id="57D09A0DFFB775419A5773869086FB59" box="[1140,1266,1159,1184]" italics="true" pageId="9" pageNumber="390">P. capitalis</emphasis>
|
||
</taxonomicName>
|
||
), a tuberculate dorsum (tuberculated bearing low warts), and olive and gray mottled and yellow blotches on groin, ventral surfaces of throat, upper arms, forearms, thighs, shanks and venter.
|
||
<taxonomicName id="A2A43D9CFFB775419D1573C09072FB11" box="[822,1030,1231,1256]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="valquii">
|
||
<emphasis id="57D09A0DFFB775419D1573C09072FB11" box="[822,1030,1231,1256]" italics="true" pageId="9" pageNumber="390">Phrynopus valquii</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="A2A43D9CFFB775419ABC73DE9169FB11" box="[1183,1309,1231,1256]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="capitalis">
|
||
<emphasis id="57D09A0DFFB775419ABC73DE9169FB11" box="[1183,1309,1231,1256]" italics="true" pageId="9" pageNumber="390">P. capitalis</emphasis>
|
||
</taxonomicName>
|
||
by lacking lateral fringes or basal webbing in toes and fingers, by bearing canthal and supratympanic stripes (absent), and by having dorsal skin shagreen with scattered tubercles (coarsely areolate).
|
||
<taxonomicName id="A2A43D9CFFB775419DDA72189085FAC9" box="[1017,1265,1303,1328]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="tautzorum">
|
||
<emphasis id="57D09A0DFFB775419DDA72189085FAC9" box="[1017,1265,1303,1328]" italics="true" pageId="9" pageNumber="390">Phrynopus tautzorum</emphasis>
|
||
</taxonomicName>
|
||
has a smooth dorsum with scattered tubercles (fleshy and tuberculate), lacks lateral fringes on digits (present), and has Toe V slightly shorter than Toe III (Toe V longer than Toe III). The following species of the
|
||
<taxonomicName id="A2A43D9CFFB775419A51726E9126FA81" box="[1138,1362,1377,1400]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="orestes">
|
||
<emphasis id="57D09A0DFFB775419A51726E9126FA81" box="[1138,1362,1377,1400]" italics="true" pageId="9" pageNumber="390">Pristimantis orestes</emphasis>
|
||
</taxonomicName>
|
||
group with reduced finger tips may externally resemble
|
||
<taxonomicName id="A2A43D9CFFB775419CCE728B97A0FA64" box="[749,980,1412,1437]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="capitalis">
|
||
<emphasis id="57D09A0DFFB775419CCE728B97A0FA64" box="[749,980,1412,1437]" italics="true" pageId="9" pageNumber="390">Phrynopus capitalis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="57D09A0DFFB775419DCA728A94B9FA1D" italics="true" pageId="9" pageNumber="390">
|
||
<taxonomicName id="A2A43D9CFFB775419DCA728A9088FA64" box="[1001,1276,1412,1437]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="atrabracus">Pristimantis atrabracus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419B2E728A9102FA64" box="[1293,1398,1412,1437]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="chimu">P. chimu</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419BA4728A9575FA39" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="cordovae">P. cordovae</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419F2C72A695CDFA39" box="[271,441,1447,1472]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="mariaelenae">P mariaelenae</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419FE472A6960AFA39" box="[455,638,1447,1472]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="melanogaster">P. melanogaster</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419CAE72A69779FA39" box="[653,781,1447,1472]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="pataikos">P. pataikos</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419D3872A697FBFA39" box="[795,911,1449,1472]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="pinguis">P. pinguis</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419DBE72A69067FA39" box="[925,1043,1449,1472]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="seorsus">P. seorsus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419A0272A690D4FA39" box="[1057,1184,1449,1472]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="simonsii">P. simonsii</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419A8D72A6910DFA39" box="[1198,1401,1447,1472]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="stictoboubonus">P. stictoboubonus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419BAB72A694B9FA1D" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="stipa">P. stipa</taxonomicName>
|
||
</emphasis>
|
||
. All of these except
|
||
<emphasis id="57D09A0DFFB775419F9972C29741FA1C" box="[442,821,1484,1509]" italics="true" pageId="9" pageNumber="390">
|
||
<taxonomicName id="A2A43D9CFFB775419F9972C29647FA1D" box="[442,563,1485,1508]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="simonsii">P. simonsii</taxonomicName>
|
||
and
|
||
<taxonomicName id="A2A43D9CFFB775419C4E72C29741FA1C" box="[621,821,1484,1509]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="attenbouroughi">P. attenbouroughi</taxonomicName>
|
||
</emphasis>
|
||
, have a visible tympanum (
|
||
<emphasis id="57D09A0DFFB775419A5272C29644F9F1" italics="true" pageId="9" pageNumber="390">
|
||
<taxonomicName id="A2A43D9CFFB775419A5272C29178FA1C" box="[1137,1292,1484,1509]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="atrabracus">P. atrabracus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419B3B72C2910FFA1C" box="[1304,1403,1484,1509]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="chimu">P. chimu</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419BA472C29575F9F1" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="cordovae">P. cordovae</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419F2F72FE95C3F9F1" box="[268,439,1519,1544]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="mariaelenae">P. mariaelenae</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419FE172FE9644F9F1" box="[450,560,1521,1544]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="pinguis">P. pinguis</taxonomicName>
|
||
</emphasis>
|
||
), or the tympanic annulus is weakly defined or concealed beneath the skin (
|
||
<emphasis id="57D09A0DFFB775419BA472FE9714F9D5" italics="true" pageId="9" pageNumber="390">
|
||
<taxonomicName id="A2A43D9CFFB775419BA472FE955BF9D4" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="melanogaster">P. melanogaster</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419F19711A95C3F9D4" box="[314,439,1556,1581]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="pataikos">P. pataikos</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419FE2711A96FCF9D4" box="[449,648,1556,1581]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="stictoboubonus">P. stictoboubonus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419CB0711A9771F9D5" box="[659,773,1557,1580]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="seorsus">P. seorsus</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419D2C711A9714F9D5" box="[783,864,1557,1580]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="stipa">P. stipa</taxonomicName>
|
||
</emphasis>
|
||
). The other two members of the group occurring in
|
||
<collectingCountry id="1DB3068FFFB775419EB4713694B9F9A9" box="[151,205,1593,1616]" name="Peru" pageId="9" pageNumber="390">Peru</collectingCountry>
|
||
,
|
||
<taxonomicName id="A2A43D9CFFB775419EFF71369503F9A9" box="[220,375,1593,1616]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="corrugatus">
|
||
<emphasis id="57D09A0DFFB775419EFF71369503F9A9" box="[220,375,1593,1616]" italics="true" pageId="9" pageNumber="390">P. corrugatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A2A43D9CFFB775419F9671369605F9A9" box="[437,625,1593,1616]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="ventriguttatus">
|
||
<emphasis id="57D09A0DFFB775419F9671369605F9A9" box="[437,625,1593,1616]" italics="true" pageId="9" pageNumber="390">P. ventriguttatus</emphasis>
|
||
</taxonomicName>
|
||
have well developed pads on all digits and cannot be confused with
|
||
<taxonomicName id="A2A43D9CFFB775419EB47153950DF98C" box="[151,377,1628,1653]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="capitalis">
|
||
<emphasis id="57D09A0DFFB775419EB47153950DF98C" box="[151,377,1628,1653]" italics="true" pageId="9" pageNumber="390">Phrynopus capitalis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="A2A43D9CFFB775419FAA7152960CF98D" box="[393,632,1629,1652]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="simonsii">
|
||
<emphasis id="57D09A0DFFB775419FAA7152960CF98D" box="[393,632,1629,1652]" italics="true" pageId="9" pageNumber="390">Pristimantis simonsii</emphasis>
|
||
</taxonomicName>
|
||
lacks a tympanum, dentigerous processes of vomers, and bears lateral fringes on digits, as does
|
||
<emphasis id="57D09A0DFFB775419F9E717096D3F961" box="[445,679,1663,1688]" italics="true" pageId="9" pageNumber="390">
|
||
<taxonomicName id="A2A43D9CFFB775419F9E717096D7F961" box="[445,675,1663,1688]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="capitalis">Phrynopus capitalis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but differs by having relatively larger nasal bones, by lacking the swollen supratympanic fold, by having smoother dorsal skin (coarsely tuberculate with warts) and by bearing some dorsal dark blotches (absent).
|
||
<taxonomicName id="A2A43D9CFFB775419FCF71C6975DF919" box="[492,809,1735,1760]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="390" phylum="Chordata" rank="species" species="attenbouroughi">
|
||
<emphasis id="57D09A0DFFB775419FCF71C6975DF919" box="[492,809,1735,1760]" italics="true" pageId="9" pageNumber="390">Pristimantis attenbouroughi</emphasis>
|
||
</taxonomicName>
|
||
differs by bearing the dentigerous processes of vomers (absent), by having dorsal skin shagreened with low scattered tubercles (coarsely areolate with large rounded warts), a dark canthal and supratympanic stripe (absent, supratympanic fold swollen), and by being smaller with males SVL =
|
||
<quantity id="A25CEBFAFFB775419F11703B95A1F8B5" box="[306,469,1844,1869]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" metricValueMax="1.92" metricValueMin="1.46" pageId="9" pageNumber="390" unit="mm" value="16.9" valueMax="19.2" valueMin="14.6">14.6–19.2 mm</quantity>
|
||
(single male SVL =
|
||
<quantity id="A25CEBFAFFB775419C9C703B9752F8B5" box="[703,806,1844,1868]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="9" pageNumber="390" unit="mm" value="23.2">23.2 mm</quantity>
|
||
; Lehr and von
|
||
<bibRefCitation id="01353BEEFFB775419DEE703A9031F8B5" author="May" box="[973,1093,1844,1869]" pageId="9" pageNumber="390" refString="Lehr, E. & von May, R. (2017) A new species of terrestrial-breeding frog (Amphibia, Craugastoridae, Pristimantis) from high elevations of the Pui Pui Protected Forest in central Peru. ZooKeys, 660, 17 - 42." type="journal article" year="2017">
|
||
<date id="111A60DFFFB775419DEE703A9031F8B5" box="[973,1093,1844,1869]" pageId="9" pageNumber="390" value="2017-05">May 2017</date>
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2DBE1594FFB775439EE4705890C6FA51" lastPageId="11" lastPageNumber="392" pageId="9" pageNumber="390" type="description">
|
||
<paragraph id="651B461FFFB775429EE470589059FEB9" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1437,151,320]" lastPageId="10" lastPageNumber="391" pageId="9" pageNumber="390">
|
||
<emphasis id="57D09A0DFFB775419EE470589592F889" bold="true" box="[199,486,1879,1904]" pageId="9" pageNumber="390">
|
||
Description of
|
||
<typeStatus id="BA1FF8BDFFB775419F5470589594F889" box="[375,480,1879,1904]" pageId="9" pageNumber="390" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Head distinct, narrower than body, wider than long, (HW 133% of HL; HW 40% of SVL; snout nearly rounded in dorsal view, sloping to rounded tip in lateral profile; snout moderately short; E-N 70% of eye diameter; nostrils small, weakly protuberant, dorsolateraly directed; canthus rostralis slightly curved in dorsal view, rounded in profile; loreal region slightly concave, lips weakly flared; interorbital area slightly convex, no cranial crests; upper eyelid without enlarged tubercles; EW 90% of IOD in male specimen. Tympanic membrane and annulus absent; tympanic region with several irregular fused tubercles or warts, some of which are arranged in a swollen fold (similar to a parotoid gland), extending from the posterior margin of the eye to the shoulders. Oval to elongate, large tongue, weakly notched; choanae small, round, well separated and anterolaterally situated; dentigerous processes of vomers absent. Vocal sac and vocal slits absent.
|
||
</paragraph>
|
||
<caption id="31DB1697FFB475429EB471A59711F904" httpUri="https://zenodo.org/record/803123/files/figure.png" pageId="10" pageNumber="391" targetBox="[475,1085,382,1678]" targetPageId="10">
|
||
<paragraph id="651B461FFFB475429EB471A59711F904" blockId="10.[151,1435,1706,1789]" pageId="10" pageNumber="391">
|
||
<emphasis id="57D09A0DFFB475429EB471A59561F946" bold="true" box="[151,277,1706,1728]" pageId="10" pageNumber="391">FIGURE 7.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="A2A43D9CFFB475429F8271A5961AF939" box="[417,622,1706,1728]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="391" phylum="Chordata" rank="species" species="capitalis" status="sp. nov.">
|
||
<emphasis id="57D09A0DFFB475429F8271A5961AF939" box="[417,622,1706,1728]" italics="true" pageId="10" pageNumber="391">Phrynopus capitalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="57D09A0DFFB475429C5471A596BDF946" bold="true" box="[631,713,1706,1727]" pageId="10" pageNumber="391">
|
||
<taxonomicNameLabel id="4CE32776FFB475429C5471A596BDF946" box="[631,713,1706,1727]" pageId="10" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male AMNH 134158 (SVL= 23.2 mm), showing (
|
||
<emphasis id="57D09A0DFFB475429AC671A59083F946" bold="true" box="[1253,1271,1706,1727]" pageId="10" pageNumber="391">A</emphasis>
|
||
) head in lateral view and (
|
||
<emphasis id="57D09A0DFFB475429F2071C69547F927" bold="true" box="[259,307,1736,1758]" pageId="10" pageNumber="391">B, C</emphasis>
|
||
) body in dorsal and ventral views. Specimen collected near Lake Manachaqui, 3550 m, near the border of Río Abiseo National Park, San Martín, Peru. Photographs by A. Catenazzi.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="651B461FFFB475439EE4702B903CFF0E" blockId="10.[151,1436,1828,2033]" lastBlockId="11.[151,1436,151,248]" lastPageId="11" lastPageNumber="392" pageId="10" pageNumber="391">
|
||
Skin of dorsum irregularly granular bearing some warts and small ridges, especially in the scapular area and on the lower back, bearing scattered very small tubercles; skin of venter and throat coarsely areolate; two large postrictal tubercles under the swollen supratympanic fold; a low elongate tubercle on the interorbital area; metacarpal tubercles large, oval and low; outer areas of forelimbs bearing some elongate warts; inner palmar tubercle nearly rhomboidal, thenar round, small slightly shorter than palmar; fingers bearing thick lateral fringes, finger tips rounded and slightly bulbous; first finger slightly shorter than second; digital pads present; subarticular tubercles round, conical in profile (
|
||
<figureCitation id="FD9F5A9AFFB575439C00779896FBFF49" box="[547,655,151,176]" captionStart="FIGURE 6" captionStartId="7.[151,250,1854,1876]" captionTargetBox="[165,1411,981,1831]" captionTargetId="figure@7.[151,1436,972,1832]" captionTargetPageId="7" captionText="FIGURE 6. Palmar and plantar surfaces of the new species of Phrynopus: (A) P. anancites sp. nov. (MUSM 33168), (B) P. capitalis sp. nov. (AMNH 134158), (C) P. dumicola sp. nov. (MUSM 33102), and (D) P. personatus sp. nov. (MUSM 33096). Photographs by M. Flecks (A, C and D) and A. Catenazzi (B)." httpUri="https://zenodo.org/record/803121/files/figure.png" pageId="11" pageNumber="392">Figures 6</figureCitation>
|
||
B, 9A); THL 38% of SVL; two metatarsal tubercles, round and large, inner one and a half the size of outer; some supernumerary plantar tubercles (
|
||
<figureCitation id="FD9F5A9AFFB575439A0477B390F8FF2C" box="[1063,1164,188,213]" captionStart="FIGURE 6" captionStartId="7.[151,250,1854,1876]" captionTargetBox="[165,1411,981,1831]" captionTargetId="figure@7.[151,1436,972,1832]" captionTargetPageId="7" captionText="FIGURE 6. Palmar and plantar surfaces of the new species of Phrynopus: (A) P. anancites sp. nov. (MUSM 33168), (B) P. capitalis sp. nov. (AMNH 134158), (C) P. dumicola sp. nov. (MUSM 33102), and (D) P. personatus sp. nov. (MUSM 33096). Photographs by M. Flecks (A, C and D) and A. Catenazzi (B)." httpUri="https://zenodo.org/record/803121/files/figure.png" pageId="11" pageNumber="392">Figure 6</figureCitation>
|
||
B). Toes bearing lateral fringes and basal webbing; toe tips bulbous, emarginated; Toe V longer than Toe III.
|
||
</paragraph>
|
||
<caption id="31DB1697FFB575439EB474AD90C8FC2F" httpUri="https://zenodo.org/record/803125/files/figure.png" pageId="11" pageNumber="392" targetBox="[340,1247,303,908]" targetPageId="11">
|
||
<paragraph id="651B461FFFB575439EB474AD90C8FC2F" blockId="11.[151,1436,930,982]" pageId="11" pageNumber="392">
|
||
<emphasis id="57D09A0DFFB575439EB474AD957AFC4E" bold="true" box="[151,270,930,952]" pageId="11" pageNumber="392">FIGURE 8</emphasis>
|
||
. Female paratopotype of
|
||
<taxonomicName id="A2A43D9CFFB575439C3474AD9697FC41" box="[535,739,930,952]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="392" phylum="Chordata" rank="species" species="capitalis" status="sp. nov.">
|
||
<emphasis id="57D09A0DFFB575439C3474AD9697FC41" box="[535,739,930,952]" italics="true" pageId="11" pageNumber="392">Phrynopus capitalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="57D09A0DFFB575439CC874AD9749FC4E" bold="true" box="[747,829,930,951]" pageId="11" pageNumber="392">
|
||
<taxonomicNameLabel id="4CE32776FFB575439CC874AD9749FC4E" box="[747,829,930,951]" pageId="11" pageNumber="392" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life (MUSM 8959, SVL = 35.6 mm) from around Lake Manachaqui, 3600 m, near Rio Abiseo National Park, San Martín, Peru. Photograph by L. O. Rodriguez.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="651B461FFFB575439EE4730C90B0FBB9" blockId="11.[151,1437,1027,1739]" pageId="11" pageNumber="392">
|
||
<emphasis id="57D09A0DFFB575439EE4730C9516FBE5" bold="true" box="[199,354,1027,1052]" pageId="11" pageNumber="392">Color in life.</emphasis>
|
||
Dorsal surfaces were brownish-gray on the scapular region on the
|
||
<typeStatus id="BA1FF8BDFFB575439A48730C90B9FBE5" box="[1131,1229,1027,1052]" pageId="11" pageNumber="392" type="holotype">holotype</typeStatus>
|
||
and brownish-tan bluish black on the flanks; the venter was yellowish gray. Iris cocoa brown. Nuptial pads black.
|
||
</paragraph>
|
||
<paragraph id="651B461FFFB575439EE4734495F2FB29" blockId="11.[151,1437,1027,1739]" pageId="11" pageNumber="392">
|
||
<emphasis id="57D09A0DFFB575439EE4734495ACFB9D" bold="true" box="[199,472,1099,1124]" pageId="11" pageNumber="392">Color in preservative.</emphasis>
|
||
Dorsal surfaces are brownish-tan with small dark brown spots appearing like a labyrinth; flanks are black with some brown on the warts (
|
||
<figureCitation id="FD9F5A9AFFB575439D00736097F1FB71" box="[803,901,1135,1160]" captionStart="FIGURE 7" captionStartId="10.[151,250,1706,1728]" captionTargetBox="[475,1085,382,1678]" captionTargetId="figure@10.[475,1112,373,1684]" captionTargetPageId="10" captionText="FIGURE 7. Holotype of Phrynopus capitalis sp. nov., male AMNH 134158 (SVL = 23.2 mm), showing (A) head in lateral view and (B, C) body in dorsal and ventral views. Specimen collected near Lake Manachaqui, 3550 m, near the border of Río Abiseo National Park, San Martín, Peru. Photographs by A. Catenazzi." httpUri="https://zenodo.org/record/803123/files/figure.png" pageId="11" pageNumber="392">Figure 7</figureCitation>
|
||
). Groin with black reticulations; posterior thigh black or brownish-tan, with the same dorsal pattern. Ventral surfaces grayish cream on the areoles with darker grayish ground color.
|
||
</paragraph>
|
||
<paragraph id="651B461FFFB575439EE473D496C3FAC5" blockId="11.[151,1437,1027,1739]" pageId="11" pageNumber="392">
|
||
<emphasis id="57D09A0DFFB575439EE473D49122FB0D" bold="true" box="[199,1366,1243,1268]" pageId="11" pageNumber="392">
|
||
Measurements in mm (data for male
|
||
<typeStatus id="BA1FF8BDFFB575439CB773D4968FFB0D" box="[660,763,1243,1268]" pageId="11" pageNumber="392" type="holotype">holotype</typeStatus>
|
||
followed by female paratopotype in parentheses)
|
||
</emphasis>
|
||
. SVL 23.2 (35.9), HL 7.0 (13.6), HW 9.3 (14.3), TL (9.0 (12.0), THL 9.1 (13.0), FL 9.4 (13.9), IOD 2.9 (3.6), EW 2.6 (3.3), E-N 2.1 (2.8), IND 1.6 (2.7), ED 3.0 (3.7).
|
||
</paragraph>
|
||
<paragraph id="651B461FFFB575439EE4724890C6FA51" blockId="11.[151,1437,1027,1739]" pageId="11" pageNumber="392">
|
||
<emphasis id="57D09A0DFFB575439EE472489548FA99" bold="true" box="[199,316,1351,1376]" pageId="11" pageNumber="392">Variation.</emphasis>
|
||
Female MUSM 8959 is larger in size (
|
||
<quantity id="A25CEBFAFFB575439CDC7248971EFA99" box="[767,874,1351,1376]" metricMagnitude="-2" metricUnit="m" metricValue="3.56" pageId="11" pageNumber="392" unit="mm" value="35.6">35.6 mm</quantity>
|
||
), and EW is narrower than the IOD (wider in the holotype). The two postrictal tubercles are also present. Coloration is the same as in the holotype. In life (
|
||
<figureCitation id="FD9F5A9AFFB575439B0F726491F9FA7D" box="[1324,1421,1387,1412]" captionStart="FIGURE 8" captionStartId="11.[151,250,930,952]" captionTargetBox="[340,1247,303,908]" captionTargetId="figure@11.[340,1247,303,908]" captionTargetPageId="11" captionText="FIGURE 8. Female paratopotype of Phrynopus capitalis sp. nov. in life (MUSM 8959, SVL = 35.6 mm) from around Lake Manachaqui, 3600 m, near Rio Abiseo National Park, San Martín, Peru. Photograph by L. O. Rodriguez." httpUri="https://zenodo.org/record/803125/files/figure.png" pageId="11" pageNumber="392">Figure 8</figureCitation>
|
||
), bluish black coloration under arms and above groin, more conspicuous than in male holotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2DBE1594FFB575439EE472BC963BFA09" pageId="11" pageNumber="392" type="etymology">
|
||
<paragraph id="651B461FFFB575439EE472BC963BFA09" blockId="11.[151,1437,1027,1739]" pageId="11" pageNumber="392">
|
||
<emphasis id="57D09A0DFFB575439EE472BC953AFA35" bold="true" box="[199,334,1459,1484]" pageId="11" pageNumber="392">Etymology.</emphasis>
|
||
The specific name
|
||
<taxonomicName id="A2A43D9CFFB575439C0B72BC96FDFA35" box="[552,649,1459,1484]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="392" phylum="Chordata" rank="species" species="capitalis">
|
||
<emphasis id="57D09A0DFFB575439C0B72BC96FDFA35" box="[552,649,1459,1484]" italics="true" pageId="11" pageNumber="392">capitalis</emphasis>
|
||
</taxonomicName>
|
||
is a Latin adjective meaning “pre-eminent” and is used in reference to the distinctive large head of the species
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2DBE1594FFB575439EE472F495EFF932" pageId="11" pageNumber="392" type="distribution">
|
||
<paragraph id="651B461FFFB575439EE472F495EFF932" blockId="11.[151,1437,1027,1739]" pageId="11" pageNumber="392">
|
||
<emphasis id="57D09A0DFFB575439EE472F49585F9ED" bold="true" box="[199,497,1531,1556]" pageId="11" pageNumber="392">Distribution and ecology.</emphasis>
|
||
<taxonomicName id="A2A43D9CFFB575439FDB72F496ACF9ED" box="[504,728,1531,1556]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="392" phylum="Chordata" rank="species" species="capitalis">
|
||
<emphasis id="57D09A0DFFB575439FDB72F496ACF9ED" box="[504,728,1531,1556]" italics="true" pageId="11" pageNumber="392">Phrynopus capitalis</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="BA1FF8BDFFB575439DCB72F3906CF9ED" box="[1000,1048,1532,1556]" pageId="11" pageNumber="392">type</typeStatus>
|
||
locality in the Manachaqui Valley, where it co-occurs with
|
||
<taxonomicName id="A2A43D9CFFB575439F8471109601F9C1" box="[423,629,1567,1592]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="392" phylum="Chordata" rank="species" species="valquii">
|
||
<emphasis id="57D09A0DFFB575439F8471109601F9C1" box="[423,629,1567,1592]" italics="true" pageId="11" pageNumber="392">Phrynopus valquii</emphasis>
|
||
</taxonomicName>
|
||
and an undescribed species of
|
||
<taxonomicName id="A2A43D9CFFB575439DF7712F9029F9CE" box="[980,1117,1568,1591]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="392" phylum="Chordata" rank="genus">
|
||
<emphasis id="57D09A0DFFB575439DF7712F9029F9CE" box="[980,1117,1568,1591]" italics="true" pageId="11" pageNumber="392">Pristimantis</emphasis>
|
||
</taxonomicName>
|
||
. This valley is covered with wet montane grasslands and contains a glacial lake that feeds a tributary of the Marañon river (
|
||
<figureCitation id="FD9F5A9AFFB575439AE3714C9158F9A5" box="[1216,1324,1603,1628]" captionStart="FIGURE 2" captionStartId="3.[151,250,1182,1204]" captionTargetBox="[159,1427,205,1151]" captionTargetId="figure@3.[151,1436,193,1161]" captionTargetPageId="3" captionText="FIGURE 2. Study area within and around Río Abiseo National Park (bold continuous line) in the Departaments (Regiones) San Martín and La Libertad, eastern slopes of the Cordillera Oriental of the Andes, Peru, containing type localities of (1) Phrynopus personatus, sp. nov., El Mirador; (2) P. dumicola, sp. nov., Pampa del Cuy; (3) P. capitalis sp. nov.., Manachaqui lake; and (4) P. anancites, sp. nov., Puesto de Vigilancia Ventanas." httpUri="https://zenodo.org/record/803113/files/figure.png" pageId="11" pageNumber="392">Figures 2</figureCitation>
|
||
,
|
||
<figureCitation id="FD9F5A9AFFB575439B1A714B9133F9A5" box="[1337,1351,1604,1628]" captionStart="FIGURE 3" captionStartId="4.[151,250,1498,1520]" captionTargetBox="[181,1413,663,1486]" captionTargetId="figure@4.[181,1413,663,1486]" captionTargetPageId="4" captionText="FIGURE 3. Habitats of species of Phrynopus in the Río Abiseo NP and surrounding area. (A, B) habitat of P. capitalis sp. nov. and P. valquii in the Manachaqui valley, 3500 m, Department of La Libertad, near the border of Río Abiseo NP. (C) Pampa del Cuy valley, showing forest patches on the slopes; habitat of P. dumicola sp. nov.. (D) View over the Montecristo Valley, from El Mirador, habitat of P. personatus sp. nov. Photographs by A. Catenazzi." httpUri="https://zenodo.org/record/803115/files/figure.png" pageId="11" pageNumber="392">3</figureCitation>
|
||
C). The three specimens were found within a
|
||
<quantity id="A25CEBFAFFB575439C79716496D1F97B" box="[602,677,1643,1667]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="392" unit="m" value="100.0">100 m</quantity>
|
||
2 area in the wettest flat area of this valley, among dense moss vegetation. This area lays outside of the Río Abiseo NP, but
|
||
<taxonomicName id="A2A43D9CFFB575439D7E718097A8F95E" box="[861,988,1678,1703]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="392" phylum="Chordata" rank="species" species="capitalis">
|
||
<emphasis id="57D09A0DFFB575439D7E718097A8F95E" box="[861,988,1678,1703]" italics="true" pageId="11" pageNumber="392">P. capitalis</emphasis>
|
||
</taxonomicName>
|
||
might be present in areas of the same habitat within the park.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |