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<mods:title id="604492A07E974B3D2B2EEAAF1C066DFB">Systematic revision and phylogeny of the endemic southeastern Asian Pristaulacus comptipennis species group (Hymenoptera: Aulacidae) 2959</mods:title>
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<mods:namePart id="17B4C75B30C2ED377C2EA9EDDCF9C96A">Turrisi, Giuseppe Fabrizio</mods:namePart>
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<paragraph id="6512F633D947FFC2FF47FA4B448005C6" blockId="42.[151,650,1483,1542]" box="[151,650,1483,1510]" pageId="42" pageNumber="43">
<heading id="3E5A415FD947FFC2FF47FA4B448005C6" bold="true" box="[151,650,1483,1510]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<taxonomicName id="A2AD8DB0D947FFC2FF47FA4B448005C6" authority="Sun &amp; Sheng 2007" authorityName="Sun &amp; Sheng" authorityYear="2007" box="[151,650,1483,1510]" class="Insecta" family="Aulacidae" genus="Pristaulacus" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="porcatus">
<emphasis id="57D92A21D947FFC2FF47FA4B448005C6" bold="true" box="[151,650,1483,1510]" pageId="42" pageNumber="43">
<emphasis id="57D92A21D947FFC2FF47FA4B479705C5" bold="true" box="[151,413,1484,1510]" italics="true" pageId="42" pageNumber="43">Pristaulacus porcatus</emphasis>
Sun &amp; Sheng 2007
</emphasis>
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<paragraph id="6512F633D947FFC2FF47FA6947480625" blockId="42.[151,650,1483,1542]" box="[151,322,1518,1542]" pageId="42" pageNumber="43">
(
<figureCitation id="FD96EAB6D947FFC2FF4FFA6947300625" box="[159,314,1518,1542]" captionStart-0="FIGURES 118122" captionStart-1="FIGURES 123126" captionStartId-0="43.[151,264,1929,1952]" captionStartId-1="44.[151,263,1482,1505]" captionTargetBox-0="[227,1363,301,1907]" captionTargetBox-1="[227,1363,157,1456]" captionTargetId-0="figure-65@43.[224,1363,301,1907]" captionTargetId-1="figure-249@44.[223,1363,157,1456]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="FIGURES 118122. Pristaulacus porcatus, paratype ♀. 118. Head, dorsal view. 119. Head, frontal view. 120. Head, lateral view. 121. Mesosoma, dorsal view. 122. Pronotum, lateral view." captionText-1="FIGURES 123126. Pristaulacus porcatus, paratype ♀. 123. Wings. 124. Mesosoma, lateral view. 125. Hind coxa ventral view. 126. Metasoma, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.5281496" figureDoi-1="http://doi.org/10.5281/zenodo.5281498" httpUri-0="https://zenodo.org/record/5281496/files/figure.png" httpUri-1="https://zenodo.org/record/5281498/files/figure.png" pageId="42" pageNumber="43">Figs 118126</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="2DB7A5B8D947FFC2FF47F9B245600668" box="[151,874,1589,1611]" pageId="42" pageNumber="43" type="reference_group">
<paragraph id="6512F633D947FFC2FF47F9B245600668" blockId="42.[151,874,1589,1611]" box="[151,874,1589,1611]" pageId="42" pageNumber="43">
<treatmentCitationGroup id="45BDD11DD947FFC2FF47F9B245600668" box="[151,874,1589,1611]" pageId="42" pageNumber="43">
<treatmentCitation id="E40CD022D947FFC2FF47F9B244760668" author="Sun, S. - P. &amp; Sheng, M. - L." box="[151,636,1589,1611]" page="217" pageId="42" pageNumber="43" year="2007">
<taxonomicName id="A2AD8DB0D947FFC2FF47F9B244760668" authority="Sun &amp; Sheng 2007 a: 217" authorityName="Sun &amp; Sheng" authorityPageNumber="217" authorityYear="2007" box="[151,636,1589,1611]" class="Insecta" family="Aulacidae" genus="Pristaulacus" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="porcatus">
<emphasis id="57D92A21D947FFC2FF47F9B2477F0668" box="[151,373,1589,1611]" italics="true" pageId="42" pageNumber="43">Pristaulacus porcatus</emphasis>
<bibRefCitation id="013C8BC2D947FFC2FEABF9B244760668" author="Sun &amp; Sheng" box="[379,636,1589,1611]" firstAuthor="Sun" page="217" pageId="42" pageNumber="43" pagination="216 - 220" refId="ref35635" refString="Sun, S. - P. &amp; Sheng, M. - L. (2007 a) Genus Pristaulacus (Hymenoptera, Aulacidae) in Henan Province, China. Acta Zootaxonomica Sinica, 32, 216 - 220." type="journal article" year="2007" yearSuffix="a">Sun &amp; Sheng 2007a: 217</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="E40CD022D947FFC2FD57F9B2456C0668" author="Turrisi, G. F. &amp; Jennings, J. T. &amp; Vilhelmsen, L." box="[647,870,1589,1611]" page="58" pageId="42" pageNumber="43" year="2009">
<bibRefCitation id="013C8BC2D947FFC2FD57F9B2456C0668" author="Turrisi" box="[647,870,1589,1611]" etAl="et al." firstAuthor="Turrisi" page="58" pageId="42" pageNumber="43" pagination="27 - 59" refId="ref36047" refString="Turrisi, G. F., Jennings, J. T. &amp; Vilhelmsen, L. (2009) Phylogeny and generic concepts of the parasitoid wasp family Aulacidae (Hymenoptera: Evanioidea). Invertebrate Systematics, 23, 27 - 59." type="journal article" year="2009">
Turrisi
<emphasis id="57D92A21D947FFC2FD01F9B1450F0668" box="[721,773,1589,1611]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
2009: 58
</bibRefCitation>
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.
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</paragraph>
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<emphasis id="57D92A21D947FFC2FF47F9FF47B506B3" box="[151,447,1655,1680]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">
<emphasis id="57D92A21D947FFC2FF47F9FF47B206B3" bold="true" box="[151,440,1655,1680]" pageId="42" pageNumber="43">Type material examined</emphasis>
.
</emphasis>
<materialsCitation id="D5C5FC6ED947FFC2FE17F9FF450206FB" collectingDate="1999-05-24" collectionCode="SFPS" elevation="450" location="The" pageId="42" pageNumber="43" specimenCount="2" specimenCount-female="1" stateProvince="Henan" typeStatus="holotype">
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type material consists of
<specimenCount id="73AB3DBAD947FFC2FCC3F9FE45B306B3" box="[787,953,1656,1680]" pageId="42" pageNumber="43" type="generic">two specimens</specimenCount>
(both
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), the
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(not examined) and the
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(examined), the latter labelled: “
<typeStatus id="BA164891D947FFC2FD63F91B451D0697" box="[691,791,1692,1716]" pageId="42" pageNumber="43" type="paratype">Paratype</typeStatus>
/
<taxonomicName id="A2AD8DB0D947FFC2FCCFF91B42C20697" authority="Sun &amp; Sheng" authorityName="Sun &amp; Sheng" authorityYear="2007" box="[799,1224,1692,1716]" class="Insecta" family="Aulacidae" genus="Pristaulacus" kingdom="Animalia" order="Hymenoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="porcatus">
<emphasis id="57D92A21D947FFC2FCCFF91B42100697" box="[799,1050,1692,1716]" italics="true" pageId="42" pageNumber="43">Pristaulacus porcatus</emphasis>
Sun &amp; Sheng
</taxonomicName>
/Lingshan,
<collectingRegion id="A76938D1D947FFC2FA9CF91A43920697" box="[1356,1432,1693,1716]" country="China" name="Henan" pageId="42" pageNumber="43">Henan</collectingRegion>
,
<date id="1113D0F3D947FFC2FF47F947470006FB" box="[151,266,1728,1752]" pageId="42" pageNumber="43" value="1999-05-24">
<collectingDate id="0157291BD947FFC2FF47F947470006FB" box="[151,266,1728,1752]" pageId="42" pageNumber="43" value="1999-05-24">1999.5.24</collectingDate>
</date>
, M-L. Sheng/
<quantity id="A2555BD6D947FFC2FE73F947442F06FB" box="[419,549,1728,1752]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="42" pageNumber="43" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">
<elevation id="EE801100D947FFC2FE73F947442F06FB" box="[419,549,1728,1752]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="42" pageNumber="43" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400500 m</elevation>
</quantity>
,
<date id="1113D0F3D947FFC2FDE1F94744A906FB" box="[561,675,1728,1752]" pageId="42" pageNumber="43" value="1999-05-24">
<collectingDate id="0157291BD947FFC2FDE1F94744A906FB" box="[561,675,1728,1752]" pageId="42" pageNumber="43" value="1999-05-24">1999.5.24</collectingDate>
</date>
” (
<collectionCode id="03BC6EF6D947FFC2FD6DF947450B06FB" box="[701,769,1728,1752]" pageId="42" pageNumber="43">SFPS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2DB7A5B8D947FFC5FF17F9634787042C" lastPageId="45" lastPageNumber="46" pageId="42" pageNumber="43" type="description">
<paragraph id="6512F633D947FFC2FF17F963439006DF" blockId="42.[151,1436,1655,2005]" box="[199,1434,1764,1789]" pageId="42" pageNumber="43">
<emphasis id="57D92A21D947FFC2FF17F963477E06DF" box="[199,372,1764,1789]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">
<emphasis id="57D92A21D947FFC2FF17F963476706DE" bold="true" box="[199,365,1764,1789]" pageId="42" pageNumber="43">Redescription</emphasis>
.
</emphasis>
FEMALE (
<typeStatus id="BA164891D947FFC2FE2AF963446B06DF" box="[506,609,1764,1788]" pageId="42" pageNumber="43" type="paratype">paratype</typeStatus>
). Length:
<quantity id="A2555BD6D947FFC2FD01F963453206DF" box="[721,824,1764,1788]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="42" pageNumber="43" unit="mm" value="10.3">10.3 mm</quantity>
; fore wing length:
<quantity id="A2555BD6D947FFC2FBDBF963426E06DF" box="[1035,1124,1764,1788]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="42" pageNumber="43" unit="mm" value="7.5">7.5 mm</quantity>
; ovipositor length:
<quantity id="A2555BD6D947FFC2FAEDF963439C06DF" box="[1341,1430,1764,1788]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="42" pageNumber="43" unit="mm" value="8.7">8.7 mm</quantity>
.
</paragraph>
<paragraph id="6512F633D947FFC3FF17F88F478900F7" blockId="42.[151,1436,1655,2005]" lastBlockId="43.[151,1435,151,212]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<emphasis id="57D92A21D947FFC2FF17F88F471C0703" box="[199,278,1800,1824]" italics="true" pageId="42" pageNumber="43">Colour</emphasis>
black, except: clypeus reddish black; mandible extensively dark reddish; maxillo-labial complex dark brown with palpomeres lighter; antenna with A1 light orange and A2 dark reddish; legs mostly reddish orange with coxae and hind femur (except articular parts) and tarsi (except articular base and apical fourth) dark brown and fore tarsus reddish brown; wings hyaline, yellowish with veins orange to light brown and stigma brownish; fore wing with a moderately wide irregular brown spot below stigma (as wide as more than half of stigma length), extended posteriorly to hind margin of cell SM2, and a very small brown spot on apex of cell D2; metasoma with side of T1, most of segment 2 and side of T3 irregularly orange. Setae: mostly whitish, light brown on frons and vertex, partly goldish on mandible.
</paragraph>
<caption id="31D2A6BBD946FFC3FF47F80E452A079D" ID-DOI="http://doi.org/10.5281/zenodo.5281496" ID-Zenodo-Dep="5281496" httpUri="https://zenodo.org/record/5281496/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,264,1929,1952]" targetBox="[227,1363,301,1907]" targetPageId="43">
<paragraph id="6512F633D946FFC3FF47F80E452A079D" blockId="43.[151,1435,1928,1982]" pageId="43" pageNumber="44">
<emphasis id="57D92A21D946FFC3FF47F80E47640783" bold="true" box="[151,366,1929,1952]" pageId="43" pageNumber="44">FIGURES 118122.</emphasis>
<taxonomicName id="A2AD8DB0D946FFC3FEA5F80E445C07BC" authorityName="Sun &amp; Sheng" authorityYear="2007" box="[373,598,1929,1951]" class="Insecta" family="Aulacidae" genus="Pristaulacus" kingdom="Animalia" order="Hymenoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="porcatus">
<emphasis id="57D92A21D946FFC3FEA5F80E445C07BC" box="[373,598,1929,1951]" italics="true" pageId="43" pageNumber="44">Pristaulacus porcatus</emphasis>
</taxonomicName>
, paratype ♀. 118. Head, dorsal view. 119. Head, frontal view. 120. Head, lateral view. 121.
<taxonomicName id="A2AD8DB0D946FFC3FED6F82E477C079D" box="[262,374,1961,1982]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="43" pageNumber="60" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
, dorsal view. 122. Pronotum, lateral view.
</paragraph>
</caption>
<caption id="31D2A6BBD941FFC4FF47FA4D47F20620" ID-DOI="http://doi.org/10.5281/zenodo.5281498" ID-Zenodo-Dep="5281498" httpUri="https://zenodo.org/record/5281498/files/figure.png" pageId="44" pageNumber="45" startId="44.[151,263,1482,1505]" targetBox="[227,1363,157,1456]" targetPageId="44">
<paragraph id="6512F633D941FFC4FF47FA4D47F20620" blockId="44.[151,1436,1482,1539]" pageId="44" pageNumber="45">
<emphasis id="57D92A21D941FFC4FF47FA4D477A05C2" bold="true" box="[151,368,1482,1505]" pageId="44" pageNumber="45">FIGURES 123126.</emphasis>
<taxonomicName id="A2AD8DB0D941FFC4FEA8FA4C445305C2" authorityName="Sun &amp; Sheng" authorityYear="2007" box="[376,601,1483,1505]" class="Insecta" family="Aulacidae" genus="Pristaulacus" kingdom="Animalia" order="Hymenoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="porcatus">
<emphasis id="57D92A21D941FFC4FEA8FA4C445305C2" box="[376,601,1483,1505]" italics="true" pageId="44" pageNumber="45">Pristaulacus porcatus</emphasis>
</taxonomicName>
, paratype ♀. 123. Wings. 124.
<taxonomicName id="A2AD8DB0D941FFC4FC77FA4C421D05C3" box="[935,1047,1483,1504]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="44" pageNumber="60" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
, lateral view. 125. Hind coxa ventral view. 126. Metasoma, lateral view.
</paragraph>
</caption>
<paragraph id="6512F633D941FFC4FF17F9D142B307F5" blockId="44.[151,1437,1622,2006]" pageId="44" pageNumber="45">
<emphasis id="57D92A21D941FFC4FF17F9D1470E064D" box="[199,260,1622,1646]" italics="true" pageId="44" pageNumber="45">Head</emphasis>
, from above, 1.1 × wider than long, shiny; occipital margin concave, with a moderately wide and deep medial groove; temple, from above, well developed, weakly convex, about as long as eye length; occipital carina moderately wide, 0.4 × diameter of an ocellus, absent along medial occipital groove; POL:OOL= 1.1; ocellar area 1.9 × wider than long; vertex, temple and upper frons with fine, superficial and dense punctures (distance between punctures 3.04.0 × puncture diameters); lower frons with fine, superficial and dense punctures; clypeus with coarse, deep and very dense punctures; malar area with fine, superficial and dense punctures on anterior half, polished on posterior half; occipital area with fine, superficial and dense punctures; antenna 0.8 × fore wing length; A3 5.3 × longer than wide; A4 7.4 × longer than wide, and 1.7 × longer than A3; A5 7.1 × longer than wide, and 1.6 × longer than A3. Setae: erect, long and scattered to dense on vertex; erect, long and dense on temple (length of setae 0.70.8 × diameter of an ocellus); erect, long and scattered to dense on upper frons; recumbent, long and dense on lower frons; recumbent, long and dense on clypeus; recumbent, short and dense on malar area.
</paragraph>
<paragraph id="6512F633D940FFC5FF17FF1E45A90337" blockId="45.[151,1436,151,1112]" pageId="45" pageNumber="46">
<taxonomicName id="A2AD8DB0D940FFC5FF17FF1E47350093" box="[199,319,153,176]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="57D92A21D940FFC5FF17FF1E47350093" box="[199,319,153,176]" italics="true" pageId="45" pageNumber="46">Mesosoma</emphasis>
</taxonomicName>
coarsely sculptured; pronotum mostly areolate-rugose, foveolate above lateroventral margin, with one acute tooth-like process above dorsal part of propleuron, and one posterior tooth like process on each lateroventral margin; propleuron polished and shiny with coarse, superficial and scattered to dense punctures on dorsal surface, fine, superficial and dense punctures on ventral surface (distance between punctures 3.04.0 × puncture diameters); prescutum triangular, narrow, long, very weakly concave, transverse-carinulate; mesoscutum mostly areolate-rugose, except anterior third, transverse-carinate; dorsally not prominent, anterior margin rounded (lateral view); notauli deep and wide; scutellum mostly transverse-carinate, areolate-rugose on anterior margin; mesopleuron areolate-rugose; metanotum scrobiculate; propodeum areolate-rugose, anterior margin longitudinally carinate, carinae irregular; venter of mesosoma mostly areolate-rugose to transverse-carinulate in middle; fore wing with vein
<emphasis id="57D92A21D940FFC5FF1EFE5B472801D7" box="[206,290,476,500]" italics="true" pageId="45" pageNumber="46">2-rs+m</emphasis>
short; coxa I polished-punctate with fine, deep and scattered to dense (outer half) punctures on ventral surface; coxa II polished-punctate on dorsal surface, with fine, superficial to deep and scattered punctures, transverse-carinate on ventral surface and on sides; coxa III transverse-carinate on dorsal surface, polished-punctate on ventral surface, with fine to coarse, deep and dense punctures on ventral surface (distance between punctures 2.0 3.0 × puncture diameters); outer spur of mid tibia longer than inner spur; inner spur of hind tibia slightly longer than outer spur; hind basitarsus 9.9 × longer than wide, and 1.3 × as tarsomeres 25; tarsal claw with four tooth-like processes. Setae: erect to semi erect, short and dense on dorsal surface; recumbent to erect, long and dense on sides and venter of mesosoma (especially on pronotum); erect, long and dense on hind surface of propodeum; erect, long and dense on propleuron (setae length at most 0.8 × pretarsus length).
</paragraph>
<paragraph id="6512F633D940FFC5FF17FCA6448A03CF" blockId="45.[151,1436,151,1112]" pageId="45" pageNumber="46">
<emphasis id="57D92A21D940FFC5FF17FCA64736031B" box="[199,316,801,824]" italics="true" pageId="45" pageNumber="46">Metasoma</emphasis>
pyriform (lateral view), strongly compressed laterally; petiole elongate, slender, 3.7 × longer than wide; segments 1 and 2 polished and shiny, impunctate; remaining segments finely microsculptured, with fine, superficial and dense punctures, except laterally on tergites; S7 polished with coarse, deep and dense punctures; T8 microsculptured with fine, deep and dense punctures; ovipositor 1.2 × fore wing length. Setae: segments 12 and sides of remaining tergites and sternites glabrous; recumbent, short and dense on middle of remaining tergites and sternites; semi erect, short and dense on S7.
</paragraph>
<paragraph id="6512F633D940FFC5FF17FC704787042C" blockId="45.[151,1436,151,1112]" box="[199,397,1015,1039]" pageId="45" pageNumber="46">MALE unknown.</paragraph>
</subSubSection>
<subSubSection id="2DB7A5B8D940FFC5FF17FB9C44FF0417" box="[199,757,1051,1076]" pageId="45" pageNumber="46" type="distribution">
<paragraph id="6512F633D940FFC5FF17FB9C44FF0417" blockId="45.[151,1436,151,1112]" box="[199,757,1051,1076]" pageId="45" pageNumber="46">
<emphasis id="57D92A21D940FFC5FF17FB9C476A0417" box="[199,352,1051,1076]" inLineHeading="true" pageId="45" pageNumber="46" reason="1">
<emphasis id="57D92A21D940FFC5FF17FB9C47530417" bold="true" box="[199,345,1051,1076]" pageId="45" pageNumber="46">Distribution</emphasis>
.
</emphasis>
<collectingCountry id="1DBAB6A3D940FFC5FEB7FB9B47A00417" box="[359,426,1052,1076]" name="China" pageId="45" pageNumber="46">China</collectingCountry>
(
<collectingRegion id="A76938D1D940FFC5FE69FB9B44650417" box="[441,623,1052,1076]" country="China" name="Henan" pageId="45" pageNumber="46">Henan Province</collectingRegion>
) (
<figureCitation id="FD96EAB6D940FFC5FD56FB9B44E30417" box="[646,745,1052,1076]" captionStart="FIGURE 186" captionStartId="68.[151,250,1664,1687]" captionTargetBox="[151,1436,193,1643]" captionTargetId="figure-176@68.[151,1436,193,1643]" captionTargetPageId="68" captionText="FIGURE 186. Distribution map of Pristaulacus comptipennis species group. The number of species is indicated for each subregions, north: Japanese islands, middle: China and surroundings, south: Laos, Thailand, Vietnam. Letters refer to species: a, P. asiaticus; b, P. boninensis; c, P. comptipennis; d, P. corellianus; e, P. dilleri; f, P. emarginaticeps; g, P. excisus; h, P. gusenleitneri; i, P. insularis; j, P. jenningsi; k, P.konishii; l, P. lagrecai; m, P. nobilei; n, P. porcatus; o, P. sharkeyi; p, P. thailandensis; q, P. vietnamensis; r, P. vilhelmseni; s, P. vivaldianus; t, P. watanabei." figureDoi="http://doi.org/10.5281/zenodo.5281552" httpUri="https://zenodo.org/record/5281552/files/figure.png" pageId="45" pageNumber="46">Fig. 186</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="2DB7A5B8D940FFC5FF17FBB847AE0474" box="[199,420,1087,1112]" pageId="45" pageNumber="46" type="biology_ecology">
<paragraph id="6512F633D940FFC5FF17FBB847AE0474" blockId="45.[151,1436,151,1112]" box="[199,420,1087,1112]" pageId="45" pageNumber="46">
<emphasis id="57D92A21D940FFC5FF17FBB8472C047B" box="[199,294,1087,1112]" inLineHeading="true" pageId="45" pageNumber="46" reason="1">
<emphasis id="57D92A21D940FFC5FF17FBB8472A047B" bold="true" box="[199,288,1087,1112]" pageId="45" pageNumber="46">Biology</emphasis>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>