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<mods:title id="37034427E67EF208F20F3ABA9E6A5CDA">Taxonomic review of the planthopper genus Orthopagus (Hemiptera, Fulgoromorpha, Dictyopharidae), with descriptions of two new species</mods:title>
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<taxonomicName id="00BBF9ED1D71DED597CCD3823910900C" ID-CoL="6SX24" authority="Germar, 1830" authorityName="Germar" authorityYear="1830" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus splendens" order="Hemiptera" pageId="8" pageNumber="377" phylum="Arthropoda" rank="species" species="splendens">Orthopagus splendens (Germar, 1830)</taxonomicName>
Figs 1B, 2
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, 3F, 10
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</paragraph>
</subSubSection>
<subSubSection id="945019182CF882A0BF77373F5E16957D" pageId="9" pageNumber="378" type="reference_group">
<paragraph id="2FD721F7C642F0C17F441C06745EEF07" pageId="9" pageNumber="378">
<taxonomicName id="321EB08053667D00693DB885BC8D5B54" class="Insecta" family="Dictyopharidae" genus="Flata" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Flata splendens" order="Hemiptera" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="splendens">
<pageBreakToken id="36966D54993C2DB68B1BA9BDDE0CC1AF" pageId="9" pageNumber="378" start="start">Flata</pageBreakToken>
splendens
</taxonomicName>
Germar, 1830: 48. Syntype(s) (number of specimens and sex unknown), Java, Indonesia (not examined).
</paragraph>
<paragraph id="6C11138BF026BEC9601C391C0C8EADB9" pageId="9" pageNumber="378">
<taxonomicName id="A005148611F9671FA29417F82686343D" class="Insecta" family="Dictyopharidae" genus="Pseudophana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophana splendens" order="Hemiptera" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="splendens">Pseudophana splendens</taxonomicName>
:
<bibRefCitation id="06E6DD20E9FC869FA8FB526C94B76287" author="Westwood, JO" journalOrPublisher="Transactions of the Linnean Society of London, Second series, Zoology" pageId="13" pageNumber="382" pagination="133 - 153" title="On the family Fulgoridae, with a monograph of the genus Fulgora of Linnaeus." volume="18" year="1839">Westwood 1839</bibRefCitation>
: 151.
</paragraph>
<paragraph id="5CE96D4229B43F8E6F21B73EADBCF93F" pageId="9" pageNumber="378">
<taxonomicName id="01DE817FE2427F36A481B0C9552952B9" class="Insecta" family="Dictyopharidae" genus="Dictyophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dictyophora" order="Hemiptera" pageId="9" pageNumber="378" phylum="Arthropoda" rank="genus">Dictyophora</taxonomicName>
[sic]
<taxonomicName id="571F68EAF09189B99CFC7E47CEFB1DDB" family="Dictyopharidae" genus="Fulgoromorpha" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Fulgoromorpha splendens" order="Hemiptera" pageId="9" pageNumber="378" rank="species" species="splendens">splendens</taxonomicName>
:
<bibRefCitation id="CBF75758876447A08E96D98ABB49195A" author="Walker, F" journalOrPublisher="Printed by order of the Trustees, London" pageId="13" pageNumber="382" title="List of the specimens of Homopterous insects in the collection of the British Museum. Part I. II." year="1851">Walker 1851</bibRefCitation>
: 310.
</paragraph>
<paragraph id="AE6DA79F09F7B19E1BBE6E7329C2357A" pageId="9" pageNumber="378">
<taxonomicName id="2AD5CFB688450433D35EDF678F10EAFD" class="Insecta" family="Dictyopharidae" genus="Anagnia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anagnia splendens" order="Hemiptera" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="splendens">Anagnia splendens</taxonomicName>
:
<bibRefCitation id="9BD85AB4F818C5199467448A78BA1225" author="Stal, C" journalOrPublisher="Herausgegeben von dem entomologischen Vereine zu Stettin" pageId="13" pageNumber="382" pagination="129 - 153" title="Miscellanea hemopterologica. Entomologische Zeitung." url="https://doi.org/10.1111/zoj.12401" volume="22" year="1861">
<normalizedToken id="C897B86A7E9C4C2880DCB5829D319DE8" originalValue="Stål">Stal</normalizedToken>
1861
</bibRefCitation>
: 149.
</paragraph>
<paragraph id="E9F7A1B1EE77D74E09C003AD2EAFE2A5" pageId="9" pageNumber="378">
<taxonomicName id="7A3B5E1947CCEB2830D61A5D461E491C" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udugama splendens" order="Hemiptera" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="splendens">Udugama splendens</taxonomicName>
:
<bibRefCitation id="F68CE0A93DBCE7AE4C5EE662F9D31132" author="Distant, WL" journalOrPublisher="Taylor &amp; Francis, London" pageId="12" pageNumber="381" title="The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera)." year="1906">Distant 1906</bibRefCitation>
): 249.
</paragraph>
<paragraph id="9D9428EBB19C3F3FFA577529FE320CBA" pageId="9" pageNumber="378">
<taxonomicName id="A684CAABD36FD1DA636FCC96B74710A7" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udugama flavocarinata" order="Hemiptera" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="flavocarinata">Udugama flavocarinata</taxonomicName>
Bierman, 1907: 161;
<bibRefCitation id="1A8D79182CE9915CAD998DE1B63313C0" author="Bierman, CJH" journalOrPublisher="Notes from the Leyden Museum Leyden" pageId="12" pageNumber="381" pagination="151 - 169" title="Homopteren aus Semarang (Java) gesammelt von Herrn E. Jacobson." volume="29" year="1908">Bierman (1908)</bibRefCitation>
: 151, Pl. 3, fig. 1. Syntypes 2 males, 1 female, Semarang, Java, Indonesia (not examined). Synonymized under
<taxonomicName id="E63BF027DE93F172C5A2D43A35BE01AE" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus splendens" order="Hemiptera" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="splendens">Orthopagus splendens</taxonomicName>
(Germar) by
<bibRefCitation id="D549C692A21531DEDA66E1B3EE7A9C2C" author="Melichar, L" journalOrPublisher="Zoologisch-Botanischen Gesellschaft in Wien" pageId="12" pageNumber="381" pagination="1 - 221" title="Monographie der Dictyophorinen (Homoptera). Abhandlungen der K. K." volume="7" year="1912">Melichar 1912</bibRefCitation>
: 59.
</paragraph>
<paragraph id="63B74C01A086C4C680413574CD721B24" pageId="9" pageNumber="378">
<taxonomicName id="58A0CB81D3B1A09B5B366E5B42CCF917" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus splendens" order="Hemiptera" pageId="9" pageNumber="378" phylum="Arthropoda" rank="species" species="splendens">Orthopagus splendens</taxonomicName>
:
<bibRefCitation id="5E425C7153A3DE8FEBAC7830EABD6F7D" author="Oshanin, VT" journalOrPublisher="- Petersbourg" pageId="12" pageNumber="381" pagination="385 - 492" title="Verzeichnis der palaearktischen Hemipteren mit besonderer Beruecksichtung ihrer Verteilung im Russischen Reiche. II. Band. Homoptera. III. Lieferung. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St." volume="13" year="1908">Oshanin 1908</bibRefCitation>
: 444;
<bibRefCitation id="6CE85B69CE780628F09246FD48C68448" author="Melichar, L" journalOrPublisher="Zoologisch-Botanischen Gesellschaft in Wien" pageId="12" pageNumber="381" pagination="1 - 221" title="Monographie der Dictyophorinen (Homoptera). Abhandlungen der K. K." volume="7" year="1912">Melichar 1912</bibRefCitation>
: 59;
<bibRefCitation id="16C58959D6597F6D5398F69A9633968A" author="Yang, CT" journalOrPublisher="Chinese Journal of Entomology Special Publication" pageId="13" pageNumber="382" pagination="1 - 189" title="Nymphs of Fulgoroidea (Homoptera: Auchenorrhyncha) with descriptions of two new species and notes on adults of Dictyopharidae." volume="8" year="1994">Yang and Yeh 1994</bibRefCitation>
: 108, 116, figs 71, 76.
</paragraph>
<paragraph id="754FE01B01103C57E133CB3B49DE2161" pageId="9" pageNumber="378">
<taxonomicName id="985F8F6D0AA1150E3BFFB3E72B5D28FB" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Orthopagus splendens var. tibialis" order="Hemiptera" pageId="9" pageNumber="378" phylum="Arthropoda" rank="variety" species="splendens" variety="tibialis">Orthopagus splendens var. tibialis</taxonomicName>
Kirkaldy in Kirkaldy &amp; Muir, 1913: 12. Syntypes (number of specimens and sex unknown), Macao, China (not examined).
</paragraph>
</subSubSection>
<subSubSection id="B68C89CC2FBEAF1381EC3E6F7AB323B4" pageId="9" pageNumber="378" type="materials_examined">
<paragraph id="637A73FA1AAB46E75B385E4338F97474" pageId="9" pageNumber="378">Material examined.</paragraph>
<paragraph id="052C204842F91C3E66324A184116A487" pageId="9" pageNumber="378">
CHINA: Hainan island: 1 male, Hainan, 5.v.1934, C. Ho leg.; 2 females, Shuiman, 640 m, 25.v.1960, C. Q. Li leg.; 2 males, 5 females, Tongshi, 340 m, 23. and 25.vi.1960, C. Q. Li leg.; 1 female, Yinggen, 200 m, 6.vii.1960, S. F. Li leg.; Yunnan province: 1 male, 1 female, Hekou, 80 m, 7.vi.1956, K. R. Huang leg.; 3 females, Hekou, Xioananxi, 200 m, 8.vi.1956, K. R. Huang leg.; 1 male, Gaoligong Mountain, 1000 m, 20.viii.1958, C. L. Li leg.; 1 male, Malipo, 20.vii.1958; 1 female, Xishuangbanna, Menga, 1050 m, 18.ix.1957, F. J. Pu leg.; 1 male, 1 female, Xishuangbanna, Menglun, 600 m, 9.ix.1993, H. L. Xu &amp; L. L. Yang leg.; 1 male, Xishuangbanna, Menglun, 10.ix.1993, X. Y. Cheng leg. (all IZCAS). VIETNAM: 1 female, Lao Kay, 31.v.1960, at light, K. Galewski leg. (MZPW); 1 male, 1 female, Ninh Binh, Cuo-phuong, 5., 7, and 8.vi.1966, R. Bielawski &amp; B. Pisarski leg. (MZPW); 2 males, 22 km S of Nha Trang, 20-26.xi.1960, C. M. Yoshimoto leg. (BPBM); 1 male, DaiLanh, N of Nha Trang, 30.xi.-5.xii.1960, C. M. Yoshimoto leg. (BPBM). THAILAND: Trang province: 1 male, Khaophappha Khaochang, 200-400 m, 3.i.1964, G. A. Samuelson leg. (BPBM). INDIA: Assam: 1 female, Mazbat near Mangaldai, 11-15.x.1910 (coll. Distant, BMNH). MALAYSIA: Penang: 1 male, Island of Penang, Baker leg. (USNM). INDONESIA: Java: 1 male, Java (West), Djasinga, 5.i.1966, J. Stusak (BPBM); 1 female, Samarang, iv.1909, E. Jacobson; 1 female, same but vi.1909 (both coll. Melichar, MMBC); 1 female, Wied. in CW. Java, GW [handwriting] (coll. Zool. Mus. Leipzig
<normalizedToken id="FB16264E0F5D55F1C6D4A42FB33F486D" originalValue="Übernahme">Uebernahme</normalizedToken>
1971, SNSD); 1 male, Java [handwriting] (MFNB).
</paragraph>
</subSubSection>
<subSubSection id="26E12122625238411807CF488DF8D0E2" lastPageId="10" lastPageNumber="379" pageId="9" pageNumber="378" type="redescription">
<paragraph id="0A6D0D379F678763BA021CEDC9FB3C5F" pageId="9" pageNumber="378">Redescription.</paragraph>
<paragraph id="CA8EF164859AF38C9A8F30CF4E64D2F9" pageId="9" pageNumber="378">Measurements (3 males, 11 females). Body length (from apex of head to tip of forewings): male 10.3-11.8 mm, female 12.7-13.6 mm; head length (from apex of cephalic process to base of eyes): male 1.28-1.36 mm, female 1.35-1.48 mm; head width (including eyes): male 1.30-1.49 mm, female 1.50-1.59 mm; forewing length: male 8.5-9.1 mm, female 10.4-11.3 mm.</paragraph>
<paragraph id="FEE656E5748692BFD06A9D18333D4E22" lastPageId="10" lastPageNumber="379" pageId="9" pageNumber="378">
Coloration. General coloration as in generic description (Figs 1B, 2
<normalizedToken id="D605DE81657CDED25C45C6C5FEF0EC21" originalValue="KL">K-L</normalizedToken>
). Vertex predominantly dark brown, lateral margins with a pale greenish or ochraceous S-shaped streak in posterior three quarters, median ca
<pageBreakToken id="929976CA4873134960EC5FCA5098430B" pageId="10" pageNumber="379" start="start">rina</pageBreakToken>
anteriorly light in some specimens (Figs 1B, 10A). Frons light ochraceous with small dark brown spots along intermediate and lateral carinae and indistinct infuscation at base (Fig. 10C). Forewing membrane pattern as in Fig. 3F. Hind wing membrane with a narrow dark brown infuscation along the apical portion of CuA1 vein, extending into a narrow infuscation along hind wing apical margin.
</paragraph>
<paragraph id="BE9073B13CA0CE72DA84C4F75231A3FB" pageId="10" pageNumber="379">
Structure. Head with cephalic process relatively short (Fig. 10
<normalizedToken id="AD9BB15587520655BB84A4EB6E43EB65" originalValue="AB">A-B</normalizedToken>
). Vertex (Fig. 10A) with ratio of length at midline to width between eyes 1.5-1.6. Transition of vertex to frons narrowly rounded in lateral view, cephalic process not inflated apically (Fig. 10B). Frons relatively narrow, with ratio of length at midline to maximum width 2.8-3.1.
</paragraph>
<paragraph id="585FB2BF48842AB3B4488CC33E797DA8" pageId="10" pageNumber="379">
Male genitalia. Pygofer, in lateral view, similar to
<taxonomicName id="E6CEACE378E2366E03B9CE271204F98F" lsidName="O. philippinus" pageId="10" pageNumber="379" rank="species" species="philippinus">O. philippinus</taxonomicName>
, with dorso-posterior margin with a large and relatively narrow, tooth-like, apically obtuse process (Fig. 10E); pygofer with ratio of ventral to dorsal width about 2.6 (Fig. 10D, F). Gonostyles (Fig. 10E) large, broad medially, with dorsal margin weakly sinuate. Aedeagus (Fig. 10
<normalizedToken id="C0381385596B4AE3D22D056A09F57E64" originalValue="GI">G-I</normalizedToken>
) with endosomal processes very long, extended posteriad and strongly curved dorso-anteriad, reaching the base of phallobase; phallobase with one pair of relatively small, inflated dorsolateral lobes, their apex elongate, thumb-like, directed posteriad (Fig. 10H), and one pair of relatively large ventral lobes, convergent and tapering posteriad (Fig. 10
<normalizedToken id="6FABC854C57C0C01DD39BAA762E53E8C" originalValue="HI">H-I</normalizedToken>
). Segment X, in lateral view, narrow basally, widening to apex beyond middle (Fig. 10E); in dorsal view, elongate, widest at apical third, ratio of length to maximum width 1.5-1.6 (Fig. 10D).
</paragraph>
<paragraph id="64FCF5B9E44DC0C6CB71EB5320C9CB33" pageId="10" pageNumber="379">Female genitalia as in generic description.</paragraph>
</subSubSection>
<subSubSection id="009240036CF830DA6D2CC0BD33840DF3" pageId="10" pageNumber="379" type="distribution">
<paragraph id="8C7A974AA69B388B7CA84528199A91BD" pageId="10" pageNumber="379">Distribution.</paragraph>
<paragraph id="3FEC326504E991D1E41A0B55083BE523" pageId="10" pageNumber="379">
Confirmed records are from Indonesia (Java), West Malaysia, Vietnam, Thailand, southern China (Hainan, Yunnnan) and north-eastern India (Assam). Based on the description and illustrations in
<bibRefCitation id="C508F023E192FC15A435099BF9A3C473" author="Yang, CT" journalOrPublisher="Chinese Journal of Entomology Special Publication" pageId="13" pageNumber="382" pagination="1 - 189" title="Nymphs of Fulgoroidea (Homoptera: Auchenorrhyncha) with descriptions of two new species and notes on adults of Dictyopharidae." volume="8" year="1994">Yang and Yeh (1994)</bibRefCitation>
,
<taxonomicName id="5E846EB7D87F5F049F0266CA8FA9882F" lsidName="O. splendens" pageId="10" pageNumber="379" rank="species" species="splendens">O. splendens</taxonomicName>
probably also occurs in Taiwan (see also
<bibRefCitation id="E9FC8A2DB4ECB3138C1DAAE0E65BCAB3" author="Tsaur, SC" journalOrPublisher="Zoological Studies" pageId="13" pageNumber="382" pagination="1 - 4" title="Some Fulgoroids (Insecta: Hemiptera) collected on Turtle Island, Taiwan." volume="44" year="2005">Tsaur 2005</bibRefCitation>
). Published records from the Philippines (Luzon;
<bibRefCitation id="91E0A26222367FA3C8E592F67C70B121" author="Stal, C" journalOrPublisher="Herausgegeben von dem entomologischen Vereine zu Stettin" pageId="13" pageNumber="382" pagination="129 - 153" title="Miscellanea hemopterologica. Entomologische Zeitung." url="https://doi.org/10.1111/zoj.12401" volume="22" year="1861">
<normalizedToken id="A6FE5FEE0409526623B04BC4CA2C4209" originalValue="Stål">Stal</normalizedToken>
1861
</bibRefCitation>
,
<bibRefCitation id="0B87DBA940F13D4336A1D1932CFFF5A0" author="Distant, WL" journalOrPublisher="Taylor &amp; Francis, London" pageId="12" pageNumber="381" title="The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera)." year="1906">Distant 1906</bibRefCitation>
) probably refer to
<taxonomicName id="DDF9183523086C00995417CD90EEF0F4" lsidName="O. philippinus" pageId="10" pageNumber="379" rank="species" species="philippinus">O. philippinus</taxonomicName>
, while the records from Sri Lanka (
<bibRefCitation id="7051D2491D2EDEDF25EC9036A75CFF94" pageId="10" pageNumber="379">Melichar 1903</bibRefCitation>
,
<bibRefCitation id="E6532DD193ED0EB04A39C661EC33428E" author="Melichar, L" journalOrPublisher="Zoologisch-Botanischen Gesellschaft in Wien" pageId="12" pageNumber="381" pagination="1 - 221" title="Monographie der Dictyophorinen (Homoptera). Abhandlungen der K. K." volume="7" year="1912">1912</bibRefCitation>
;
<bibRefCitation id="DA3AD742659C341C9FD6EFBCB740E2ED" author="Distant, WL" journalOrPublisher="Taylor &amp; Francis, London" pageId="12" pageNumber="381" title="The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera)." year="1906">Distant 1906</bibRefCitation>
) concern
<taxonomicName id="19B77254BFEAD11D6824D9695FBCB407" lsidName="O. exoletus" pageId="10" pageNumber="379" rank="species" species="exoletus">O. exoletus</taxonomicName>
and the ones from Japan (
<bibRefCitation id="3AE5E2CF4E460C04BFD28CE5615EC9D5" pageId="10" pageNumber="379">Matsumura 1905a</bibRefCitation>
,
<bibRefCitation id="BD65B451D2DA082B0A3DE0DBF6D783E3" author="Matsumura, S" journalOrPublisher="Transactions of the Sapporo Natural History Society" pageId="12" pageNumber="381" pagination="15 - 38" title="Die Hemipterenfauna von Riukiu (Okinawa)." volume="1" year="1905 b">b</bibRefCitation>
,
<bibRefCitation id="59FC9F8559B1A29656A3AB19D0DE0A2A" author="Matsumura, S" journalOrPublisher="Transactions of the Sapporo Natural History Society" pageId="12" pageNumber="381" url="https://doi.org/10.5962/bhl.title.35666" year="1910">1910</bibRefCitation>
) refer to
<taxonomicName id="45C3BC8A7656F4138984E7FF9CAA5C41" lsidName="O. lunulifer" pageId="10" pageNumber="379" rank="species" species="lunulifer">O. lunulifer</taxonomicName>
. Records from western India (
<bibRefCitation id="4593E5146108274AAD23ACB592CCC74E" author="Stal, C" journalOrPublisher="Herausgegeben von dem entomologischen Vereine zu Stettin" pageId="13" pageNumber="382" pagination="129 - 153" title="Miscellanea hemopterologica. Entomologische Zeitung." url="https://doi.org/10.1111/zoj.12401" volume="22" year="1861">
<normalizedToken id="F3E4C4BF8A66E5D7D616072A706688D6" originalValue="Stål">Stal</normalizedToken>
1861
</bibRefCitation>
), Myanmar (
<bibRefCitation id="B78F6F7FAFBF3A6675C177ABC2791086" author="Distant, WL" journalOrPublisher="Taylor &amp; Francis, London" pageId="12" pageNumber="381" title="The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera)." year="1906">Distant 1906</bibRefCitation>
), Singapore, and Indonesia: Sumatra (
<bibRefCitation id="F2369721AF80A46EEFD2FAEBD0807ADB" author="Bierman, CJH" journalOrPublisher="Notes from the Leyden Museum Leyden" pageId="12" pageNumber="381" pagination="151 - 169" title="Homopteren aus Semarang (Java) gesammelt von Herrn E. Jacobson." volume="29" year="1908">Bierman 1908</bibRefCitation>
) still need to be checked.
</paragraph>
</subSubSection>
<subSubSection id="E12460C25835654A631786A0381AAB35" pageId="10" pageNumber="379" type="remarks">
<paragraph id="2455144F833A28F948DB6F6293D847DA" pageId="10" pageNumber="379">Remarks.</paragraph>
<paragraph id="0780E2198BBBEED975763E002152104B" pageId="10" pageNumber="379">
<taxonomicName id="3205AB690DFF4466EAF7A70BDC43F52F" class="Insecta" family="Flatidae" genus="Flata" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Flata splendens" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="splendens">Flata splendens</taxonomicName>
was described by
<bibRefCitation id="0BC12AABD06876902EB38B2CC34110AD" author="Germar, EF" journalOrPublisher="Thon's Entomologisches Archiv" pageId="12" pageNumber="381" pagination="1 - 57" title="Species Cicadarium enumeratae et sub genera distributae." volume="2" year="1830">Germar (1830)</bibRefCitation>
from Java, and was designated as the type species of
<taxonomicName id="8D82D829A7176E2ACF50166F8A6CDE53" class="Insecta" family="Noctuidae" genus="Anagnia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anagnia" order="Lepidoptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="genus">Anagnia</taxonomicName>
by
<bibRefCitation id="8E1A8CF23434A12E8D4C1F845A12A3F4" author="Stal, C" journalOrPublisher="Herausgegeben von dem entomologischen Vereine zu Stettin" pageId="13" pageNumber="382" pagination="129 - 153" title="Miscellanea hemopterologica. Entomologische Zeitung." url="https://doi.org/10.1111/zoj.12401" volume="22" year="1861">
<normalizedToken id="83DA815FB11D006FE17FD0A6B130F01D" originalValue="Stål">Stal</normalizedToken>
(1861)
</bibRefCitation>
.
<taxonomicName id="7B424F59C2C96B40A2D530F6CC46E1A4" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udugama flavocarinata" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="flavocarinata">Udugama flavocarinata</taxonomicName>
Bierman, 1907 from Java was considered as a junior synonym of
<taxonomicName id="3385ED4F66DE606D030B9895371A0466" lsidName="O. splendens" pageId="10" pageNumber="379" rank="species" species="splendens">O. splendens</taxonomicName>
by
<bibRefCitation id="AEA1ED3D749EDE82473ABED3B3D6B6A7" author="Melichar, L" journalOrPublisher="Zoologisch-Botanischen Gesellschaft in Wien" pageId="12" pageNumber="381" pagination="1 - 221" title="Monographie der Dictyophorinen (Homoptera). Abhandlungen der K. K." volume="7" year="1912">Melichar (1912)</bibRefCitation>
.
<taxonomicName id="C16B59C044ABEA9EA891EE725B75FE88" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Orthopagus splendens var. tibialis" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="variety" species="splendens" variety="tibialis">Orthopagus splendens var. tibialis</taxonomicName>
Kirkaldy was differentiated from the typical form by having &quot;the fore and middle tibiae distinctly bi- or tri-angulate with brownish&quot; (
<bibRefCitation id="738D27ECB755183605199EF4103C3DF1" author="Kirkaldy, GW" journalOrPublisher="Report of the work of the Experiment Station of the Hawaiian Sugar Plantersʼ Association, Entomological series, Bulletin Honolulu" pageId="12" pageNumber="381" pagination="1 - 90" title="On some new species of leaf-hoppers." volume="12" year="1913">Kirkaldy and Muir 1913</bibRefCitation>
). However, this pattern is typical for all
<taxonomicName id="9B8EFCA16C967E3E92323CCAD14E07E4" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="genus">Orthopagus</taxonomicName>
species.
</paragraph>
<paragraph id="5BCE070E524F5CF71B2847A62613F46E" pageId="10" pageNumber="379">
The synonymy of
<taxonomicName id="FDB3FD7981C8D03EA556579B813814F0" class="Insecta" family="Dictyopharidae" genus="Dictyophara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dictyophara indiana" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="indiana">Dictyophara indiana</taxonomicName>
Walker, 1851 with
<taxonomicName id="3E644EE0AADD3BFC753404DC71420577" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus splendens" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="splendens">Orthopagus splendens</taxonomicName>
proposed by
<bibRefCitation id="9F2EDA162336DDA6B59BA92FE7A6E992" author="Stal, C" journalOrPublisher="Herausgegeben von dem entomologischen Vereine zu Stettin" pageId="13" pageNumber="382" pagination="129 - 153" title="Miscellanea hemopterologica. Entomologische Zeitung." url="https://doi.org/10.1111/zoj.12401" volume="22" year="1861">
<normalizedToken id="EA2FAE292A5222F54186DCB544563B53" originalValue="Stål">Stal</normalizedToken>
(1861)
</bibRefCitation>
is considered doubtful (see above).
</paragraph>
<paragraph id="762AA5344877902B6976171C261EA73E" pageId="10" pageNumber="379">
<taxonomicName id="09C8B126B0481FA64C75FC026B2001A4" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus splendens" order="Hemiptera" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="splendens">Orthopagus splendens</taxonomicName>
has been frequently confused in literature and museum collections with other species of the genus. It can be differentiated from them by the combination of the following characters: a relatively small body size, a moderately short head, an extensive dark pattern on vertex, the pygofer with dorso-posterior margin bearing a relatively large but narrow, simply tooth-like process, the endosomal processes of the aedeagus relatively long and robust, covered with minute superficial spines and phallotheca bearing two pairs of membranous lobes.
</paragraph>
</subSubSection>
</treatment>
</document>