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<document id="269A85D9C3DC28A80BFFB0B256E35BB4" ID="10.11646/zootaxa.4144.3.1" ID-DOI="10.11646/zootaxa.4144.3.1" ID-GBIF-Dataset="ea093250-c3c4-403c-a05c-98c49d0e0aa5" ID-ISSN="1175-5326" ID-Zenodo-Dep="255192" ID-ZooBank="B73CFE90-BDF1-47EA-BBD6-52A8DB2B144C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469685944659" checkinUser="plazi" docAuthor="Costa, Daniel N. R., Savaris, Marcoandre, Marinoni, Luciane &amp; Mathis, Wayne N." docDate="2016" docId="EA2A612CD03B1310FF65FF2A970B0ADA" docLanguage="en" docName="zootaxa.4144.3.1.pdf" docOrigin="Zootaxa 4144 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Limnellia Malloch" docType="treatment" docVersion="14" lastPageNumber="304" masterDocId="16131954D0381313FFF2FFB295790F50" masterDocTitle="Two new, brachypterous Limnellia species from the Venezuelan Andes (Diptera: Ephydridae)" masterLastPageNumber="315" masterPageNumber="301" pageNumber="304" updateTime="1698366400071" updateUser="plazi">
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<mods:title id="F2AF4F3FE029195CBADA2926EC542BEF">Two new, brachypterous Limnellia species from the Venezuelan Andes (Diptera: Ephydridae)</mods:title>
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<mods:namePart id="D6FCFF0E1BEF328FB6D75D1EDD244CC6">Costa, Daniel N. R.</mods:namePart>
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<mods:namePart id="086BE6E6A6B0C927AF8FB1FFF49B1B71">Marinoni, Luciane</mods:namePart>
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<mods:namePart id="781A18372292C639560C5516BBCACAFF">Mathis, Wayne N.</mods:namePart>
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<mods:date id="9A83E98F0057C7F14A286230E249CCD5">2016</mods:date>
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<treatment id="EA2A612CD03B1310FF65FF2A970B0ADA" ID-DOI="http://doi.org/10.5281/zenodo.3513763" ID-GBIF-Taxon="123208023" ID-Zenodo-Dep="3513763" LSID="urn:lsid:plazi:treatment:EA2A612CD03B1310FF65FF2A970B0ADA" httpUri="http://treatment.plazi.org/id/EA2A612CD03B1310FF65FF2A970B0ADA" lastPageNumber="304" pageId="3" pageNumber="304">
<subSubSection id="2A9983B1D03B1310FF65FF2A94A80FE2" box="[151,465,151,178]" pageId="3" pageNumber="304" type="nomenclature">
<paragraph id="623CD03AD03B1310FF65FF2A94A80FE2" blockId="3.[151,465,151,178]" box="[151,465,151,178]" pageId="3" pageNumber="304">
<heading id="39746756D03B1310FF65FF2A94A80FE2" bold="true" box="[151,465,151,178]" fontSize="11" level="1" pageId="3" pageNumber="304" reason="1">
<emphasis id="50F70C28D03B1310FF65FF2A94A80FE2" bold="true" box="[151,465,151,178]" pageId="3" pageNumber="304">
Genus
<taxonomicName id="A583ABB9D03B1310FF1CFF2594A80FE2" ID-CoL="62X6G" ID-ENA="92593" authority="Malloch" authorityName="Malloch" box="[238,465,151,178]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="50F70C28D03B1310FF1CFF25941A0FE1" bold="true" box="[238,355,151,177]" italics="true" pageId="3" pageNumber="304">Limnellia</emphasis>
Malloch
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2A9983B1D03B1310FF65FF6C94CC0EDE" pageId="3" pageNumber="304" type="reference_group">
<paragraph id="623CD03AD03B1310FF65FF6C90260E00" blockId="3.[151,1437,222,520]" pageId="3" pageNumber="304">
<treatmentCitationGroup id="4293F714D03B1310FF65FF6C90260E00" pageId="3" pageNumber="304">
<taxonomicName id="A583ABB9D03B1310FF65FF6C97780FA4" ID-CoL="62X6G" authority="Malloch 1925: 331" authorityName="Malloch" authorityPageNumber="331" authorityYear="1925" box="[151,513,222,244]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="50F70C28D03B1310FF65FF6C95810FA4" box="[151,248,222,244]" italics="true" pageId="3" pageNumber="304">Limnellia</emphasis>
<treatmentCitation id="E322F62BD03B1310FEE6FF6C97780FA4" author="Malloch" box="[276,513,222,244]" page="331" pageId="3" pageNumber="304" year="1925">
<bibRefCitation id="0612ADCBD03B1310FEE6FF6C94C20FA4" author="Malloch" box="[276,443,222,244]" pageId="3" pageNumber="304" refString="Malloch, J. R. (1925) Notes on Australian Diptera. No. vii. Proceedings of the Linnean Society of New South Wales, 50 (4), 311 - 340." type="journal article" year="1925">Malloch 1925</bibRefCitation>
: 331
</treatmentCitation>
</taxonomicName>
(feminine).
<typeStatus id="BD386E98D03B1310FD59FF6D97A40FA4" box="[683,733,223,244]" pageId="3" pageNumber="304">Type</typeStatus>
species:
<taxonomicName id="A583ABB9D03B1310FC94FF6C90540FA4" ID-CoL="72BRZ" authority="Malloch 1925" authorityName="Malloch" authorityYear="1925" box="[870,1325,222,244]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="50F70C28D03B1310FC94FF6C91120FA4" box="[870,1131,222,244]" italics="true" pageId="3" pageNumber="304">Limnellia maculipennis</emphasis>
<bibRefCitation id="0612ADCBD03B1310FB75FF6C90540FA4" author="Malloch" box="[1159,1325,222,244]" pageId="3" pageNumber="304" refString="Malloch, J. R. (1925) Notes on Australian Diptera. No. vii. Proceedings of the Linnean Society of New South Wales, 50 (4), 311 - 340." type="journal article" year="1925">Malloch 1925</bibRefCitation>
</taxonomicName>
, original designation.─
<treatmentCitation id="E322F62BD03B1310FEA6FF4F974D0E43" author="Andersson" box="[340,564,253,275]" page="53" pageId="3" pageNumber="304" year="1971">
<bibRefCitation id="0612ADCBD03B1310FEA6FF4F977E0E43" author="Andersson" box="[340,519,253,275]" pageId="3" pageNumber="304" refString="Andersson, H. (1971) The European species of Limnellia (Dipt., Ephydridae). Entomologica Scandinavica, 2 (1), 53 - 59. http: // dx. doi. org / 10.1163 / 187631271 X 00059" type="journal article" year="1971">Andersson 1971</bibRefCitation>
: 53
</treatmentCitation>
59 [review, European species].─
<treatmentCitation id="E322F62BD03B1310FC5BFF4F910B0E43" author="Mathis" box="[937,1138,253,275]" page="250" pageId="3" pageNumber="304" year="1978">
<bibRefCitation id="0612ADCBD03B1310FC5BFF4F914E0E43" author="Mathis" box="[937,1079,253,275]" pageId="3" pageNumber="304" refString="Mathis, W. N. (1978) A revision of the Nearctic species of Limnellia Malloch (Diptera: Ephydridae). Proceedings of the Biological Society of Washington, 91 (1), 250 - 293." type="journal article" year="1978">Mathis 1978</bibRefCitation>
: 250
</treatmentCitation>
293 [revision of Nearctic species].─
<bibRefCitation id="0612ADCBD03B1310FEC2FEAE94BC0E62" author="Mathis" box="[304,453,284,306]" pageId="3" pageNumber="304" refString="Mathis, W. N. (1980) Studies of Ephydrinae (Diptera: Ephydridae), III: Revision of some Neotropical genera and species. Smithsonian Contributions to Zoology, 303, 50 + iv pp." type="book" year="1980">Mathis 1980</bibRefCitation>
[review, Neotropical fauna].─
<treatmentCitation id="E322F62BD03B1310FCCAFEAE91E50E62" ID-DOI="http://doi.org/10.5281/zenodo.3507222" ID-Zenodo-Dep="3507222" author="Zhang" box="[824,1180,284,306]" httpUri="http://treatment.plazi.org/id/5B0587C9FF90FF8719A3D079FBE2E8AA" page="558" pageId="3" pageNumber="304" year="2009">
<bibRefCitation id="0612ADCBD03B1310FCCAFEAE91210E62" author="Zhang" box="[824,1112,284,306]" pageId="3" pageNumber="304" refString="Zhang, J. &amp; Yang, D. (2009) Species of the genus Limnellia from China (Diptera: Ephydridae). Zootaxa, 2308, 58 - 64." type="journal article" year="2009">Zhang and Yang 2009</bibRefCitation>
: 558
</treatmentCitation>
64 [review, Chinese fauna].─
<treatmentCitation id="E322F62BD03B1310FEEDFE8894920E00" author="Krivosheina" box="[287,491,314,336]" page="1" pageId="3" pageNumber="304" year="2012">
<bibRefCitation id="0612ADCBD03B1310FEEDFE8894A00E00" author="Krivosheina" box="[287,473,314,336]" pageId="3" pageNumber="304" refString="Krivosheina, M. G. (2012) Review of the shore-fly genus Limnellia Malloch, 1925 (Diptera, Ephydridae) of Russia. Far Eastern Entomologist, 246, 1 - 7." type="journal article" year="2012">Krivosheina 2012</bibRefCitation>
: 1
</treatmentCitation>
7 [review, Russian species].─
<treatmentCitation id="E322F62BD03B1310FCD4FE8891680E00" author="Mathis" box="[806,1041,314,336]" page="563" pageId="3" pageNumber="304" year="2014">
<bibRefCitation id="0612ADCBD03B1310FCD4FE88969A0E00" author="Mathis" box="[806,995,314,336]" pageId="3" pageNumber="304" refString="Mathis, W. N., Marinoni, L. &amp; Costa, D. N. R. (2014) A review of Scatellini Wirth and Stone (Diptera: Ephydridae) from Brazil. Zoologia. An International Journal of Zoology, 28, 1 - 16. http: // dx. doi. org / 10.1590 / S 1984 - 46702014000600005" type="journal article" year="2014">
Mathis
<emphasis id="50F70C28D03B1310FC81FE8996DF0E00" box="[883,934,314,336]" italics="true" pageId="3" pageNumber="304">et al.</emphasis>
2014
</bibRefCitation>
: 563
</treatmentCitation>
564 [review, Brazilian species].
</treatmentCitationGroup>
</paragraph>
<paragraph id="623CD03AD03B1310FF65FEE894CC0EDE" blockId="3.[151,1437,222,520]" pageId="3" pageNumber="304">
<emphasis id="50F70C28D03B1310FF65FEE894630E3F" box="[151,282,346,367]" italics="true" pageId="3" pageNumber="304">Eustigoptera</emphasis>
<bibRefCitation id="0612ADCBD03B1310FED6FEEB94CF0E3F" author="Cresson" box="[292,438,345,367]" pageId="3" pageNumber="304" refString="Cresson, E. T. Jr. (1930) Studies in the dipterous family Ephydridae. Paper III. Transactions of the American Entomological Society, 56, 93 - 131." type="journal article" year="1930">Cresson 1930</bibRefCitation>
: 126 (feminine). Type species:
<taxonomicName id="A583ABB9D03B1310FCF4FEEB912F0E3F" ID-CoL="88SC2" authority="Fallen 1813" authorityName="Fallen" authorityYear="1813" box="[774,1110,345,367]" class="Insecta" family="Ephydridae" genus="Notiphila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="50F70C28D03B1310FCF4FEEB96B40E3F" box="[774,973,345,367]" italics="true" pageId="3" pageNumber="304">Notiphila quadrata</emphasis>
Fallén 1813
</taxonomicName>
, original designation.─
<bibRefCitation id="0612ADCBD03B1310FAB9FEEB95840EDE" author="Cresson" pageId="3" pageNumber="304" refString="Cresson, E. T. Jr. (1935) Descriptions of genera and species of the dipterous family Ephydridae. Transactions of the American Entomological Society, 61, 345 - 372." type="journal article" year="1935">Cresson 1935</bibRefCitation>
: 362 [synonymy].
</paragraph>
</subSubSection>
<subSubSection id="2A9983B1D03B1310FF65FE2497B70D58" pageId="3" pageNumber="304" type="description">
<paragraph id="623CD03AD03B1310FF65FE24940C0E9B" blockId="3.[151,1437,222,520]" pageId="3" pageNumber="304">
<emphasis id="50F70C28D03B1310FF65FE2494660EFC" box="[151,287,406,428]" italics="true" pageId="3" pageNumber="304">Stictoscatella</emphasis>
<bibRefCitation id="0612ADCBD03B1310FED6FE2494D90EFC" author="Collin" box="[292,416,406,428]" pageId="3" pageNumber="304" refString="Collin, J. E. (1930) Some new species of the dipterous genus Scatella Dsv. and the differentiation of Stictoscatella gen. nov. (Ephydridae). The Entomologist's Monthly Magazine, 66, 133 - 39." type="journal article" year="1930">Collin 1930</bibRefCitation>
: 133 (feminine). Type species:
<taxonomicName id="A583ABB9D03B1310FD2EFE2491660EFC" authority="Fallen 1813" authorityName="Fallen" authorityYear="1813" box="[732,1055,406,428]" class="Insecta" family="Ephydridae" genus="Notiphila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="50F70C28D03B1310FD2EFE2496E70EFC" box="[732,926,406,428]" italics="true" pageId="3" pageNumber="304">Notiphila quadrata</emphasis>
Fallén 1813
</taxonomicName>
, original designation.─
<bibRefCitation id="0612ADCBD03B1310FAF9FE2490E10EFC" author="Cresson" box="[1291,1432,406,428]" pageId="3" pageNumber="304" refString="Cresson, E. T. Jr. (1935) Descriptions of genera and species of the dipterous family Ephydridae. Transactions of the American Entomological Society, 61, 345 - 372." type="journal article" year="1935">Cresson 1935</bibRefCitation>
: 362 [synonymy].
</paragraph>
<paragraph id="623CD03AD03B1310FF65FE6697B70D58" blockId="3.[151,1437,222,520]" pageId="3" pageNumber="304">
<emphasis id="50F70C28D03B1310FF65FE6694770EBA" box="[151,270,468,490]" italics="true" pageId="3" pageNumber="304">Stranditella</emphasis>
<bibRefCitation id="0612ADCBD03B1310FEEAFE6694E90EBA" author="Duda" box="[280,400,468,490]" pageId="3" pageNumber="304" refString="Duda, O. (1942) Neue oder ungenugend bekannte Zweiflugler der palaarktischen Region aus meiner Sammlung. 2. Fortsetzung. Deutsche Entomologische Zeitschrift, 1942 (1 - 4), 1 - 39. http: // dx. doi. org / 10.1002 / mmnd. 194219420102" type="journal article" year="1942">Duda 1942</bibRefCitation>
: 30 (as a subgenus of
<taxonomicName id="A583ABB9D03B1310FD79FE6696500EBA" box="[651,809,468,490]" class="Insecta" family="Ephydridae" genus="Lamproscatella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="50F70C28D03B1310FD79FE6696500EBA" box="[651,809,468,490]" italics="true" pageId="3" pageNumber="304">Lamproscatella</emphasis>
</taxonomicName>
; feminine). Type species:
<taxonomicName id="A583ABB9D03B1310FBB4FE6690E10EBA" authority="Fallen 1813" authorityName="Fallen" authorityYear="1813" box="[1094,1432,468,490]" class="Insecta" family="Ephydridae" genus="Notiphila" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="50F70C28D03B1310FBB4FE6690770EBA" box="[1094,1294,468,490]" italics="true" pageId="3" pageNumber="304">Notiphila quadrata</emphasis>
Fallén 1813
</taxonomicName>
, original designation.─
<bibRefCitation id="0612ADCBD03B1310FE5BFE40976F0D58" author="Dahl" box="[425,534,498,520]" pageId="3" pageNumber="304" refString="Dahl, R. G. (1959) Studies on Scandinavian Ephydridae (Diptera Brachycera). Opuscula Entomologica, 15 (Supplementum), 1 - 224." type="journal article" year="1959">Dahl 1959</bibRefCitation>
: 126 [synonymy].
</paragraph>
</subSubSection>
<subSubSection id="2A9983B1D03B1310FF65FD8794870B16" pageId="3" pageNumber="304" type="diagnosis">
<paragraph id="623CD03AD03B1310FF65FD8794870B16" blockId="3.[151,1437,565,1418]" pageId="3" pageNumber="304">
<emphasis id="50F70C28D03B1310FF65FD8794730D1E" bold="true" box="[151,266,565,590]" pageId="3" pageNumber="304">Diagnosis</emphasis>
.
<taxonomicName id="A583ABB9D03B1310FEEAFD8794FA0D1E" box="[280,387,565,590]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="50F70C28D03B1310FEEAFD8794FA0D1E" box="[280,387,565,590]" italics="true" pageId="3" pageNumber="304">Limnellia</emphasis>
</taxonomicName>
is distinguished from other genera of the tribe
<taxonomicName id="A583ABB9D03B1310FC7CFD8796810D1E" box="[910,1016,565,590]" class="Insecta" family="Ephydridae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="tribe" tribe="Scatellini">Scatellini</taxonomicName>
by the following characters: Small to moderately small shore flies, body length 1.00
<quantity id="A57B7DDFD03B1310FD4FFDEB96510D21" box="[701,808,601,626]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="304" unit="mm" value="2.5">2.50 mm</quantity>
; mostly dark brown to black, microtomentose to bare, shiny, frequently with cinereous guttate and vittate maculae; wing maculate, generally dark with white spots.
<emphasis id="50F70C28D03B1310FAA4FDCF90E50DC6" box="[1366,1436,637,662]" italics="true" pageId="3" pageNumber="304">Head:</emphasis>
Mesofrons distinct from duller parafrons, sometimes subshiny; lateroclinate fronto-orbital seta 1 (rarely 2); inner and outer vertical setae both well developed; paravertical setae either reduced or absent; ocelli arranged in isosceles or equilateral triangle or absent. Arista at most macropubescent. Face with shallow antennal grooves; interfoveal carina not projected or creased dorsally; facial setae mostly small, hair-like. Eye nearly round. Gena relatively short, bearing 1 well-developed seta.
<emphasis id="50F70C28D03B1310FDB9FC8397DC0C1A" box="[587,677,817,842]" italics="true" pageId="3" pageNumber="304">Thorax:</emphasis>
Acrostichal setae uniform in size, small, arranged in 2 rows that extend to base of scutellum; dorsocentral setae 2 (0+2); supra-alar seta either reduced or lacking; disc of scutellum bare; lateral scutellar setae usually 2, sometimes 1 or lacking. Wing maculate, mostly dark brown with white, generally guttate markings.
<emphasis id="50F70C28D03B1310FE24FC2F97300CE6" box="[470,585,925,950]" italics="true" pageId="3" pageNumber="304">Abdomen:</emphasis>
Tergites black, becoming shiny and polished posteriorly. Male terminalia: Epandrium a closed plate around cercal cavity, bearing articulated surstyli on anterior margin; gonites elongate with broad base, bearing setae on ventral portion; phallapodeme in lateral view nearly straight or deflected medially; dorsal aedeagal opening with a fold along ventral margin; aedeagus without distiphallus; ejaculatory apodeme usually present, small.
</paragraph>
</subSubSection>
<subSubSection id="2A9983B1D03B1310FF35FBE3970B0ADA" pageId="3" pageNumber="304" type="discussion">
<paragraph id="623CD03AD03B1310FF35FBE390E40A4E" blockId="3.[151,1437,565,1418]" pageId="3" pageNumber="304">
<materialsCitation id="D2EBDA67D03B1310FF35FBE390E40A4E" ID-GBIF-Occurrence="1288188035" country="Colombia, Jamaica" county="Mathis" location="Only" municipality="Zatwarnicki" pageId="3" pageNumber="304" specimenCount="1" typeStatus="holotype">
<emphasis id="50F70C28D03B1310FF35FBE3943C0B3A" bold="true" box="[199,325,1105,1130]" pageId="3" pageNumber="304">Discussion</emphasis>
. With description of the two new species below,
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<emphasis id="50F70C28D03B1310FC78FBE3968F0B3A" box="[906,1014,1105,1130]" italics="true" pageId="3" pageNumber="304">Limnellia</emphasis>
</taxonomicName>
now includes 24 species worldwide (
<bibRefCitation id="0612ADCBD03B1310FF6CFBC797890BDE" author="Mathis" box="[158,752,1141,1166]" pageId="3" pageNumber="304" refString="Mathis, W. N. &amp; Zatwarnicki, T. (1995) A world catalog of the shore flies (Diptera: Ephydridae). Memoirs on Entomology, International 4, vi + 423 pp." type="book" year="1995">
<collectingCounty id="8B5DA8B6D03B1310FF6CFBC795940BDE" box="[158,237,1141,1166]" pageId="3" pageNumber="304">Mathis</collectingCounty>
and
<collectingMunicipality id="82584A40D03B1310FED7FBC794D50BDE" box="[293,428,1141,1166]" pageId="3" pageNumber="304">Zatwarnicki</collectingMunicipality>
1995 and electronic updates
</bibRefCitation>
) that occur in most temperate regions or at higher elevations in subtropical and tropical zones.
<location id="675C86E1D03B1310FDE0FB2B97330BE2" LSID="urn:lsid:plazi:treatment:EA2A612CD03B1310FF65FF2A970B0ADA:675C86E1D03B1310FDE0FB2B97330BE2" box="[530,586,1177,1202]" country="Colombia, Jamaica" county="Mathis" municipality="Zatwarnicki" name="Only" pageId="3" pageNumber="304">Only</location>
two species were known previously from
<location id="675C86E1D03B1310FBD5FB2B907F0BE2" LSID="urn:lsid:plazi:treatment:EA2A612CD03B1310FF65FF2A970B0ADA:675C86E1D03B1310FBD5FB2B907F0BE2" box="[1063,1286,1177,1202]" country="Colombia, Jamaica" county="Mathis" municipality="Zatwarnicki" name="Neotropical Region" pageId="3" pageNumber="304">Neotropical Region</location>
,
<taxonomicName id="A583ABB9D03B1310FAE3FB28959C0B86" authority="Mathis" authorityName="Mathis" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="huachuca">
<emphasis id="50F70C28D03B1310FAE3FB2890E50BE2" box="[1297,1436,1177,1202]" italics="true" pageId="3" pageNumber="304">L. huachuca</emphasis>
Mathis
</taxonomicName>
(
<collectingCountry id="1A9490AAD03B1310FF07FB0F94AB0B86" box="[245,466,1213,1238]" isEnumeration="true" pageId="3" pageNumber="304">Colombia, Jamaica</collectingCountry>
,
<collectingCountry id="1A9490AAD03B1310FE2DFB0F974E0B86" box="[479,567,1213,1238]" name="Mexico" pageId="3" pageNumber="304">Mexico</collectingCountry>
, and
<location id="675C86E1D03B1310FD84FB0C978B0B86" LSID="urn:lsid:plazi:treatment:EA2A612CD03B1310FF65FF2A970B0ADA:675C86E1D03B1310FD84FB0C978B0B86" box="[630,754,1213,1238]" country="Colombia, Jamaica" county="Mathis" municipality="Zatwarnicki" name="Porto Rico" pageId="3" pageNumber="304">Porto Rico</location>
) and
<taxonomicName id="A583ABB9D03B1310FCC6FB0C91220B86" authority="Mathis" authorityName="Mathis" box="[820,1115,1213,1238]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="itatiaia">
<emphasis id="50F70C28D03B1310FCC6FB0C96DD0B85" box="[820,932,1214,1237]" italics="true" pageId="3" pageNumber="304">L. itatiaia</emphasis>
Mathis (
<collectingCountry id="1A9490AAD03B1310FBF9FB0F912B0B86" box="[1035,1106,1213,1238]" name="Brazil" pageId="3" pageNumber="304">Brazil</collectingCountry>
)
</taxonomicName>
(
<bibRefCitation id="0612ADCBD03B1310FB99FB0F91860B86" author="Mathis" box="[1131,1279,1213,1238]" pageId="3" pageNumber="304" refString="Mathis, W. N. (1980) Studies of Ephydrinae (Diptera: Ephydridae), III: Revision of some Neotropical genera and species. Smithsonian Contributions to Zoology, 303, 50 + iv pp." type="book" year="1980">
<location id="675C86E1D03B1310FB99FB0F91C30B86" LSID="urn:lsid:plazi:treatment:EA2A612CD03B1310FF65FF2A970B0ADA:675C86E1D03B1310FB99FB0F91C30B86" box="[1131,1210,1213,1238]" country="Colombia, Jamaica" county="Mathis" municipality="Zatwarnicki" name="Mathis" pageId="3" pageNumber="304">Mathis</location>
1980
</bibRefCitation>
,
<bibRefCitation id="0612ADCBD03B1310FAFEFB0F95AD0BAA" author="Mathis" pageId="3" pageNumber="304" refString="Mathis, W. N., Marinoni, L. &amp; Costa, D. N. R. (2014) A review of Scatellini Wirth and Stone (Diptera: Ephydridae) from Brazil. Zoologia. An International Journal of Zoology, 28, 1 - 16. http: // dx. doi. org / 10.1590 / S 1984 - 46702014000600005" type="journal article" year="2014">
<location id="675C86E1D03B1310FAFEFB0F90230B86" LSID="urn:lsid:plazi:treatment:EA2A612CD03B1310FF65FF2A970B0ADA:675C86E1D03B1310FAFEFB0F90230B86" box="[1292,1370,1213,1238]" country="Colombia, Jamaica" county="Mathis" municipality="Zatwarnicki" name="Mathis" pageId="3" pageNumber="304">Mathis</location>
<emphasis id="50F70C28D03B1310FA90FB0C90E50B86" box="[1378,1436,1213,1238]" italics="true" pageId="3" pageNumber="304">et al.</emphasis>
2014
</bibRefCitation>
).
<location id="675C86E1D03B1310FF14FB50946E0BA9" LSID="urn:lsid:plazi:treatment:EA2A612CD03B1310FF65FF2A970B0ADA:675C86E1D03B1310FF14FB50946E0BA9" box="[230,279,1250,1273]" country="Colombia, Jamaica" county="Mathis" municipality="Zatwarnicki" name="Two" pageId="3" pageNumber="304">Two</location>
species in the genus exhibit brachyptery of two different
<typeStatus id="BD386E98D03B1310FC69FB5096A30BAA" box="[923,986,1250,1274]" pageId="3" pageNumber="304">types</typeStatus>
:
<taxonomicName id="A583ABB9D03B1310FC16FB5094670A4E" authority="Hollmann-Schirrmacher and Zatwarnicki" authorityName="Hollmann-Schirrmacher and Zatwarnicki" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="helmuti">
<emphasis id="50F70C28D03B1310FC16FB50912F0BAA" box="[996,1110,1249,1274]" italics="true" pageId="3" pageNumber="304">L. helmuti</emphasis>
Hollmann-Schirrmacher and Zatwarnicki
</taxonomicName>
has a short and broad wing, and
<taxonomicName id="A583ABB9D03B1310FD7FFAB4965B0A4E" box="[653,802,1285,1310]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="abbreviata">
<emphasis id="50F70C28D03B1310FD7FFAB4965B0A4E" box="[653,802,1285,1310]" italics="true" pageId="3" pageNumber="304">L. abbreviata</emphasis>
</taxonomicName>
, has a narrow wing. Both species are incapable of flight.
</materialsCitation>
</paragraph>
<paragraph id="623CD03AD03B1310FF35FA9B970B0ADA" blockId="3.[151,1437,565,1418]" pageId="3" pageNumber="304">
The two new species described herein are closely related (see “Remarks” sections below) and form a lineage within
<taxonomicName id="A583ABB9D03B1310FF1BFAFF942C0A36" box="[233,341,1357,1382]" class="Insecta" family="Ephydridae" genus="Limnellia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="50F70C28D03B1310FF1BFAFF942C0A36" box="[233,341,1357,1382]" italics="true" pageId="3" pageNumber="304">Limnellia</emphasis>
</taxonomicName>
that seems to have arisen and diversified due to evolutionary pressures within the Paramo or during the development of this ecosystem.
</paragraph>
</subSubSection>
</treatment>
</document>