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<document id="B994A5F624E30E9BDDC2B9EFEEA89106" ID-DOI="10.5281/zenodo.5276613" ID-ISSN="0253-116X" ID-Zenodo-Dep="5276613" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630016730227" checkinUser="carolina" docAuthor="Assing, V." docDate="2009" docId="EA1EF20AFFE8FFBFCEABFBC5FEB4FAE1" docLanguage="en" docName="LinzerbiolBeitr.2009.41.2.1285-1305.pdf" docOrigin="Linzer biologische Beiträge 41 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Leptusa (Roubaliusa) sengleti PACE 1984" docType="treatment" docVersion="4" lastPageNumber="1294" masterDocId="16278A72FFE1FFB6CE19FF9DFFE9FF92" masterDocTitle="New species, new synonymies, and additional records of Leptusa from Turkey and Iran (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="1305" masterPageNumber="1285" pageNumber="1294" updateTime="1699137003817" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="D67983399B30F3511675CB5F047031A5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="16016A5691205A2BF7F1AC0968EE3D4F">
<mods:title id="30B222AFEA66E10C5C90E20E196EC29A">New species, new synonymies, and additional records of Leptusa from Turkey and Iran (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="0FCC74A45929C520F2CAF54AB8355199">Assing, V.</mods:namePart>
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<mods:title id="533FA7B5B3B58A404AA1E4C9FDD2091D">Linzer biologische Beiträge</mods:title>
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<mods:date id="3FD31460EC91FDC6143EC2B55DFB5209">2009</mods:date>
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<mods:number id="2E4A8AE5BAF2FDF75D9FF15FEA64BA3B">2009-12-18</mods:number>
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<mods:number id="D1F693618E794EC8AC321B40BBB4CF54">41</mods:number>
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<treatment id="EA1EF20AFFE8FFBFCEABFBC5FEB4FAE1" ID-DOI="http://doi.org/10.5281/zenodo.5278776" ID-Zenodo-Dep="5278776" LSID="urn:lsid:plazi:treatment:EA1EF20AFFE8FFBFCEABFBC5FEB4FAE1" httpUri="http://treatment.plazi.org/id/EA1EF20AFFE8FFBFCEABFBC5FEB4FAE1" lastPageNumber="1294" pageId="9" pageNumber="1294">
<subSubSection id="2AAD1097FFE8FFBFCEABFBC5FCDCFBFC" box="[178,821,1111,1134]" pageId="9" pageNumber="1294" type="nomenclature">
<paragraph id="6208431CFFE8FFBFCEABFBC5FCDCFBFC" blockId="9.[178,821,1111,1134]" box="[178,821,1111,1134]" pageId="9" pageNumber="1294">
<heading id="3940F470FFE8FFBFCEABFBC5FCDCFBFC" box="[178,821,1111,1134]" fontSize="8" level="2" pageId="9" pageNumber="1294" reason="2">
<taxonomicName id="A5B7389FFFE8FFBFCEABFBC5FD8AFBFC" ID-CoL="722F8" authority="PACE 1984" authorityName="PACE" authorityYear="1984" box="[178,611,1111,1134]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1294" phylum="Arthropoda" rank="species" species="sengleti" subGenus="Roubaliusa">
<emphasis id="50C39F0EFFE8FFBFCEABFBC5FD8AFBFC" bold="true" box="[178,611,1111,1134]" pageId="9" pageNumber="1294">
<emphasis id="50C39F0EFFE8FFBFCEABFBC5FEECFBFF" bold="true" box="[178,261,1112,1133]" italics="true" pageId="9" pageNumber="1294">Leptusa</emphasis>
(
<emphasis id="50C39F0EFFE8FFBFCF0AFBCAFE63FBFF" bold="true" box="[275,394,1111,1133]" italics="true" pageId="9" pageNumber="1294">Roubaliusa</emphasis>
)
<emphasis id="50C39F0EFFE8FFBFCF83FBCAFE03FBFF" bold="true" box="[410,490,1111,1133]" italics="true" pageId="9" pageNumber="1294">sengleti</emphasis>
<bibRefCitation id="06263EEDFFE8FFBFCFE9FBC5FDC0FBFE" author="PACE R" box="[496,553,1112,1134]" pageId="9" pageNumber="1294" pagination="211 - 219" refId="ref11133" refString="PACE R. (1984): Nuove Aleocharinae microftalme mediterranee e dell'Iran, del Museum d'Histoire naturelle di Ginevra (Coleoptera, Staphylinidae). - Archives Scientifiques Geneve 37: 211 - 219." type="journal article" year="1984">PACE</bibRefCitation>
1984
</emphasis>
</taxonomicName>
(
<figureCitation id="FA8C5F99FFE8FFBFCC69FBC5FD08FBFC" box="[624,737,1112,1134]" captionStart="Figs 49-54" captionStartId="16.[178,218,1491,1511]" captionTargetBox="[186,1079,879,1436]" captionTargetId="figure-319@16.[178,1084,873,1476]" captionTargetPageId="16" captionText="Figs 49-54: Leptusa ionopolitana PACE (49-51), L. amisensis PACE (52), and L. sengleti PACE (53- 54): (49, 53) median lobe of aedeagus in lateral view; (50-52) spermatheca; (54) male sternite VII. Scale bars: 54: 0.2 mm; 49-53: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276633" httpUri="https://zenodo.org/record/5276633/files/figure.png" pageId="9" pageNumber="1294">Figs 53-54</figureCitation>
,
<figureCitation id="FA8C5F99FFE8FFBFCCF2FBC5FCC7FBFC" box="[747,814,1112,1134]" captionStart="Map 4" captionStartId="14.[178,223,731,751]" captionTargetBox="[178,1084,212,707]" captionTargetId="figure-1@14.[178,1084,212,707]" captionTargetPageId="14" captionText="Map 4: Distributions of the Leptusa species of the subgenus Roubaliusa SCHEERPELTZ in northern Iran: L. sengleti PACE (filled square), L. piciformis nov.sp. (open circle), L. flagrifera nov.sp. (open circle), L. delphiniformis nov.sp. (filled circle), and L. soriciformis nov.sp. (open square). Localities where only females were found are omitted." figureDoi="http://doi.org/10.5281/zenodo.5276631" httpUri="https://zenodo.org/record/5276631/files/figure.png" pageId="9" pageNumber="1294">Map 4</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2AAD1097FFE8FFBFCEABFBE3FD60FB48" pageId="9" pageNumber="1294" type="reference_group">
<paragraph id="6208431CFFE8FFBFCEABFBE3FD9CFB01" blockId="9.[178,1085,1150,1242]" box="[178,629,1150,1171]" pageId="9" pageNumber="1294">
<treatmentCitationGroup id="42A76432FFE8FFBFCEABFBE3FD9CFB01" box="[178,629,1150,1171]" pageId="9" pageNumber="1294">
<treatmentCitation id="E316650DFFE8FFBFCEABFBE3FDB0FB01" author="PACE R" box="[178,601,1150,1171]" page="212" pageId="9" pageNumber="1294" year="1984">
<taxonomicName id="A5B7389FFFE8FFBFCEABFBE3FDB0FB01" ID-CoL="722F8" authority="PACE 1984: 212" authorityName="PACE" authorityPageNumber="212" authorityYear="1984" box="[178,601,1150,1171]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1294" phylum="Arthropoda" rank="species" species="sengleti" subGenus="Roubaliusa">
<emphasis id="50C39F0EFFE8FFBFCEABFBE3FF13FB01" box="[178,250,1150,1171]" italics="true" pageId="9" pageNumber="1294">Leptusa</emphasis>
(
<emphasis id="50C39F0EFFE8FFBFCF1EFBE3FE87FB01" box="[263,366,1150,1171]" italics="true" pageId="9" pageNumber="1294">Roubaliusa</emphasis>
)
<emphasis id="50C39F0EFFE8FFBFCF62FBE3FE28FB01" box="[379,449,1150,1171]" italics="true" pageId="9" pageNumber="1294">sengleti</emphasis>
<bibRefCitation id="06263EEDFFE8FFBFCFDEFBE2FDB0FB01" author="PACE R" box="[455,601,1151,1171]" pageId="9" pageNumber="1294" pagination="211 - 219" refId="ref11133" refString="PACE R. (1984): Nuove Aleocharinae microftalme mediterranee e dell'Iran, del Museum d'Histoire naturelle di Ginevra (Coleoptera, Staphylinidae). - Archives Scientifiques Geneve 37: 211 - 219." type="journal article" year="1984">PACE 1984: 212</bibRefCitation>
</taxonomicName>
</treatmentCitation>
ff.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6208431CFFE8FFBFCEABFB05FD60FB48" blockId="9.[178,1085,1150,1242]" pageId="9" pageNumber="1294">
T y p e m a t e r i a l e x a m i n e d
<typeStatus id="BD0CFDBEFFE8FFBFCC46FB05FD45FB3E" box="[607,684,1176,1196]" pageId="9" pageNumber="1294" type="holotype">Holotype</typeStatus>
3: &quot;
<collectingCountry id="1AA0038CFFE8FFBFCCF4FB05FCFCFB3E" box="[749,789,1176,1196]" name="Iran" pageId="9" pageNumber="1294">Iran</collectingCountry>
,
<collectingRegion id="A0738DFEFFE8FFBFCD3FFB05FC8FFB3E" box="[806,870,1176,1196]" country="Iran" name="Gilan" pageId="9" pageNumber="1294">Guilân</collectingRegion>
, sur Asâlem,
<quantity id="A54FEEF9FFE8FFBFCDEFFB05FBD0FB3E" box="[1014,1081,1176,1196]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="1294" unit="m" value="450.0">450 m</quantity>
,
<geoCoordinate id="078325DBFFE8FFBFCEABFB32FF17FB51" box="[178,254,1199,1219]" degrees="37" direction="north" minutes="42" orientation="latitude" pageId="9" pageNumber="1294" precision="925" value="37.7">37°42'N</geoCoordinate>
,
<geoCoordinate id="078325DBFFE8FFBFCF15FB32FEBEFB51" box="[268,343,1199,1219]" degrees="48" direction="east" minutes="53" orientation="longitude" pageId="9" pageNumber="1294" precision="925" value="48.883335">48°53'E</geoCoordinate>
, A. Senglet, 10.6.75 /
<typeStatus id="BD0CFDBEFFE8FFBFCC21FB32FD7EFB51" box="[568,663,1199,1219]" pageId="9" pageNumber="1294" type="holotype">Holotypus</typeStatus>
<taxonomicName id="A5B7389FFFE8FFBFCCB8FB32FCD3FB51" ID-CoL="722F8" authorityName="PACE" authorityYear="1984" box="[673,826,1199,1219]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1294" phylum="Arthropoda" rank="species" species="sengleti">Leptusa sengleti</taxonomicName>
mihi, det. R. Pace 1983 /
<taxonomicName id="A5B7389FFFE8FFBFCEABFB5BFEAFFB48" ID-CoL="722F8" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" box="[178,326,1222,1242]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1294" phylum="Arthropoda" rank="species" species="sengleti" status="sp. nov.">Leptusa sengleti</taxonomicName>
<taxonomicNameLabel id="4BF02275FFE8FFBFCF55FB5BFE96FB48" box="[332,383,1222,1242]" pageId="9" pageNumber="1294" rank="species">n. sp.</taxonomicNameLabel>
, det. R. Pace 1983 (MHNG).
</paragraph>
</subSubSection>
<subSubSection id="2AAD1097FFE8FFBFCEABFB75FEB4FAE1" pageId="9" pageNumber="1294" type="description">
<paragraph id="6208431CFFE8FFBFCEABFB75FEB4FAE1" blockId="9.[178,1085,1256,1395]" pageId="9" pageNumber="1294">
C o m m e n t: The original description of
<taxonomicName id="A5B7389FFFE8FFBFCC98FB74FD05FB6F" authorityName="Assing" authorityYear="2009" box="[641,748,1256,1278]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1294" phylum="Arthropoda" rank="species" species="sengleti">
<emphasis id="50C39F0EFFE8FFBFCC98FB74FD05FB6F" box="[641,748,1256,1278]" italics="true" pageId="9" pageNumber="1294">L. sengleti</emphasis>
</taxonomicName>
is based on a single male (
<bibRefCitation id="06263EEDFFE8FFBFCA1CFB75FF00FA89" author="PACE R" pageId="9" pageNumber="1294" pagination="211 - 219" refId="ref11133" refString="PACE R. (1984): Nuove Aleocharinae microftalme mediterranee e dell'Iran, del Museum d'Histoire naturelle di Ginevra (Coleoptera, Staphylinidae). - Archives Scientifiques Geneve 37: 211 - 219." type="journal article" year="1984">PACE 1984</bibRefCitation>
). The aedeagus (
<figureCitation id="FA8C5F99FFE8FFBFCF85FA98FE00FA89" box="[412,489,1285,1307]" captionStart="Figs 49-54" captionStartId="16.[178,218,1491,1511]" captionTargetBox="[186,1079,879,1436]" captionTargetId="figure-319@16.[178,1084,873,1476]" captionTargetPageId="16" captionText="Figs 49-54: Leptusa ionopolitana PACE (49-51), L. amisensis PACE (52), and L. sengleti PACE (53- 54): (49, 53) median lobe of aedeagus in lateral view; (50-52) spermatheca; (54) male sternite VII. Scale bars: 54: 0.2 mm; 49-53: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276633" httpUri="https://zenodo.org/record/5276633/files/figure.png" pageId="9" pageNumber="1294">Fig. 53</figureCitation>
) and the male sternite VII (
<figureCitation id="FA8C5F99FFE8FFBFCD0FFA98FC8DFA89" box="[790,868,1285,1307]" captionStart="Figs 49-54" captionStartId="16.[178,218,1491,1511]" captionTargetBox="[186,1079,879,1436]" captionTargetId="figure-319@16.[178,1084,873,1476]" captionTargetPageId="16" captionText="Figs 49-54: Leptusa ionopolitana PACE (49-51), L. amisensis PACE (52), and L. sengleti PACE (53- 54): (49, 53) median lobe of aedeagus in lateral view; (50-52) spermatheca; (54) male sternite VII. Scale bars: 54: 0.2 mm; 49-53: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276633" httpUri="https://zenodo.org/record/5276633/files/figure.png" pageId="9" pageNumber="1294">Fig. 54</figureCitation>
) are most similar to those of
<taxonomicName id="A5B7389FFFE8FFBFCF15FABEFE66FAAA" authorityName="Assing" authorityYear="2009" box="[268,399,1315,1336]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1294" phylum="Arthropoda" rank="species" species="piciformis">
<emphasis id="50C39F0EFFE8FFBFCF15FABEFE66FAAA" box="[268,399,1315,1336]" italics="true" pageId="9" pageNumber="1294">L. piciformis</emphasis>
</taxonomicName>
and
<taxonomicName id="A5B7389FFFE8FFBFCFDBFABEFDD6FAAA" authorityName="Assing" authorityYear="2009" box="[450,575,1315,1336]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1294" phylum="Arthropoda" rank="species" species="flagrifera">
<emphasis id="50C39F0EFFE8FFBFCFDBFABEFDD6FAAA" box="[450,575,1315,1336]" italics="true" pageId="9" pageNumber="1294">L. flagrifera</emphasis>
</taxonomicName>
, respectively. This species was the first representative of the genus to become known from
<collectingCountry id="1AA0038CFFE8FFBFCC73FADDFD7FFAC7" box="[618,662,1344,1365]" name="Iran" pageId="9" pageNumber="1294">Iran</collectingCountry>
. It is currently known only from the
<typeStatus id="BD0CFDBEFFE8FFBFCA08FADDFBD4FAC7" box="[1041,1085,1344,1365]" pageId="9" pageNumber="1294">type</typeStatus>
locality (
<figureCitation id="FA8C5F99FFE8FFBFCF12FAC0FEB9FAE1" box="[267,336,1373,1395]" captionStart="Map 4" captionStartId="14.[178,223,731,751]" captionTargetBox="[178,1084,212,707]" captionTargetId="figure-1@14.[178,1084,212,707]" captionTargetPageId="14" captionText="Map 4: Distributions of the Leptusa species of the subgenus Roubaliusa SCHEERPELTZ in northern Iran: L. sengleti PACE (filled square), L. piciformis nov.sp. (open circle), L. flagrifera nov.sp. (open circle), L. delphiniformis nov.sp. (filled circle), and L. soriciformis nov.sp. (open square). Localities where only females were found are omitted." figureDoi="http://doi.org/10.5281/zenodo.5276631" httpUri="https://zenodo.org/record/5276631/files/figure.png" pageId="9" pageNumber="1294">Map 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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