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<mods:title id="EE566EC4D7EC31F9A914655723F9F063">New species, new synonymies, and additional records of Leptusa from Turkey and Iran (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<subSubSection id="2AAD1097FFE0FFB7CEABFA9AFD20FA8E" box="[178,713,1286,1308]" pageId="1" pageNumber="1286" type="nomenclature">
<paragraph id="6208431CFFE0FFB7CEABFA9AFD20FA8E" blockId="1.[178,713,1286,1308]" box="[178,713,1286,1308]" pageId="1" pageNumber="1286">
<heading id="3940F470FFE0FFB7CEABFA9AFD20FA8E" box="[178,713,1286,1308]" fontSize="8" level="2" pageId="1" pageNumber="1286" reason="2">
<taxonomicName id="A5B7389FFFE0FFB7CEABFA9AFDB6FA8E" ID-CoL="722DG" authority="PACE 1983" authorityName="PACE" authorityYear="1983" box="[178,607,1286,1308]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="1286" phylum="Arthropoda" rank="species" species="janczyki" subGenus="Neopisalia">
<emphasis id="50C39F0EFFE0FFB7CEABFA9AFDB6FA8E" bold="true" box="[178,607,1286,1308]" pageId="1" pageNumber="1286">
<emphasis id="50C39F0EFFE0FFB7CEABFA9AFEECFA8E" bold="true" box="[178,261,1287,1308]" italics="true" pageId="1" pageNumber="1286">Leptusa</emphasis>
(
<emphasis id="50C39F0EFFE0FFB7CF0AFA9BFE6BFA8E" bold="true" box="[275,386,1286,1308]" italics="true" pageId="1" pageNumber="1286">Neopisalia</emphasis>
)
<emphasis id="50C39F0EFFE0FFB7CF88FA9BFE0EFA8E" bold="true" box="[401,487,1286,1308]" italics="true" pageId="1" pageNumber="1286">janczyki</emphasis>
PACE 1983
</emphasis>
</taxonomicName>
(
<figureCitation id="FA8C5F99FFE0FFB7CC75FA9BFD29FA8E" box="[620,704,1286,1308]" captionStart="Figs 1-9" captionStartId="6.[178,218,1351,1371]" captionTargetBox="[184,1071,219,1311]" captionTargetId="figure-127@6.[178,1085,219,1331]" captionTargetPageId="6" captionText="Figs 1-9: Leptusa janczyki PACE, holotype (1-3) and L. flagellulifera nov.sp. (4-9): (1) pronotum and elytra; (2) male tergite VII; (3, 6-7) median lobe of aedeagus in lateral view; (4) habitus; (5) forebody; (8) apical lobe of paramere; (9) spermatheca. Scale bars: 4: 1.0 mm; 1-2, 5: 0.5 mm; 3, 6- 9: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276619" httpUri="https://zenodo.org/record/5276619/files/figure.png" pageId="1" pageNumber="1286">Figs 1-3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2AAD1097FFE0FFB7CEABFAB0FC43FAE3" pageId="1" pageNumber="1286" type="reference_group">
<paragraph id="6208431CFFE0FFB7CEABFAB0FDBDFAD3" blockId="1.[178,1084,1324,1393]" box="[178,596,1324,1346]" pageId="1" pageNumber="1286">
<taxonomicName id="A5B7389FFFE0FFB7CEABFAB0FDA6FAD3" ID-CoL="722DG" authority="PACE 1983: 74" authorityName="PACE" authorityPageNumber="74" authorityYear="1983" box="[178,591,1324,1346]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="1286" phylum="Arthropoda" rank="species" species="janczyki" subGenus="Neopisalia">
<emphasis id="50C39F0EFFE0FFB7CEABFAB0FF13FAD0" box="[178,250,1325,1346]" italics="true" pageId="1" pageNumber="1286">Leptusa</emphasis>
(
<emphasis id="50C39F0EFFE0FFB7CF1EFAB1FE80FAD3" box="[263,361,1324,1345]" italics="true" pageId="1" pageNumber="1286">Neopisalia</emphasis>
)
<emphasis id="50C39F0EFFE0FFB7CF6FFAB1FE29FAD3" box="[374,448,1324,1345]" italics="true" pageId="1" pageNumber="1286">janczyki</emphasis>
PACE 1983: 74
</taxonomicName>
.
</paragraph>
<paragraph id="6208431CFFE0FFB7CEABFADAFC43FAE3" blockId="1.[178,1084,1324,1393]" pageId="1" pageNumber="1286">
<materialsCitation id="D2DF4941FFE0FFB7CEABFADAFC43FAE3" collectionCode="T, R, NHMW" pageId="1" pageNumber="1286" specimenCount="2" typeStatus="holotype">
<collectionCode id="04A6DBD9FFE0FFB7CEABFADAFF29FAC9" box="[178,192,1351,1371]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="1286">T</collectionCode>
y p e m a t e r i a l e x a m i n e d:
<typeStatus id="BD0CFDBEFFE0FFB7CC49FADAFD74FAC9" box="[592,669,1351,1371]" pageId="1" pageNumber="1286" type="holotype">Holotype</typeStatus>
3: &quot;3 / Lederi Epp., Lagistan, Pont Euxin mer., Dieck / Typus /
<typeStatus id="BD0CFDBEFFE0FFB7CF60FAC0FE31FAE3" box="[377,472,1373,1393]" pageId="1" pageNumber="1286" type="holotype">Holotypus</typeStatus>
<taxonomicName id="A5B7389FFFE0FFB7CFC4FAC0FD91FAE3" ID-CoL="722DG" authorityName="PACE" authorityYear="1983" box="[477,632,1373,1393]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="1286" phylum="Arthropoda" rank="species" species="janczyki">Leptusa janczyki</taxonomicName>
m, det.
<collectionCode id="04A6DBD9FFE0FFB7CCDBFAC0FD3DFAE3" box="[706,724,1373,1393]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="1286">R</collectionCode>
. Pace 1981&quot; (
<collectionCode id="04A6DBD9FFE0FFB7CD49FAC0FC49FAE3" box="[848,928,1373,1393]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="1286" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="2AAD1097FFE0FFB4CEABFA1DFDE8FDA1" lastPageId="2" lastPageNumber="1287" pageId="1" pageNumber="1286" type="description">
<paragraph id="6208431CFFE0FFB7CEABFA1DFDCDF9D7" blockId="1.[178,1086,1408,1605]" pageId="1" pageNumber="1286">
C o m m e n t: The above
<typeStatus id="BD0CFDBEFFE0FFB7CFCFFA1DFDC6FA04" box="[470,559,1408,1430]" pageId="1" pageNumber="1286" type="holotype">holotype</typeStatus>
was examined to clarify the identity of
<taxonomicName id="A5B7389FFFE0FFB7CDD6FA1CFBD7FA07" authorityName="PACE" authorityYear="1983" box="[975,1086,1408,1430]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="1286" phylum="Arthropoda" rank="species" species="janczyki">
<emphasis id="50C39F0EFFE0FFB7CDD6FA1CFBD7FA07" box="[975,1086,1408,1430]" italics="true" pageId="1" pageNumber="1286">L. janczyki</emphasis>
</taxonomicName>
and the status of
<taxonomicName id="A5B7389FFFE0FFB7CF7AFA03FD99FA21" authority="ASSING 2007" authorityName="ASSING" authorityYear="2007" box="[355,624,1437,1459]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="1286" phylum="Arthropoda" rank="species" species="soganlica">
<emphasis id="50C39F0EFFE0FFB7CF7AFA03FE09FA20" box="[355,480,1437,1459]" italics="true" pageId="1" pageNumber="1286">L. soganlica</emphasis>
ASSING 2007
</taxonomicName>
, which was described from the Soganlı Dağı in
<collectingRegion id="A0738DFEFFE0FFB7CEC8FA27FEC0FA42" box="[209,297,1466,1488]" country="Turkey" name="Trabzon" pageId="1" pageNumber="1286">Trabzon</collectingRegion>
. Based on the illustration of the aedeagus of
<taxonomicName id="A5B7389FFFE0FFB7CD3AFA26FC7FFA5D" authorityName="PACE" authorityYear="1983" box="[803,918,1466,1488]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="1286" phylum="Arthropoda" rank="species" species="janczyki">
<emphasis id="50C39F0EFFE0FFB7CD3AFA26FC7FFA5D" box="[803,918,1466,1488]" italics="true" pageId="1" pageNumber="1286">L. janczyki</emphasis>
</taxonomicName>
in the original description, there was a chance that the types of both names were conspecific (
<bibRefCitation id="06263EEDFFE0FFB7CDF6FA4AFF1DF999" author="ASSING V" pageId="1" pageNumber="1286" pagination="427 - 436" refId="ref10861" refString="ASSING V. (2009 a): On the Leptusa species of Turkey. VII. Notes on distribution, a new species, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae). - Linzer biologische Beitrage 41 (1): 427 - 436." type="journal article" year="2009">ASSING 2009a</bibRefCitation>
). The type locality of
<taxonomicName id="A5B7389FFFE0FFB7CFC4FA6BFDA5F998" authorityName="PACE" authorityYear="1983" box="[477,588,1525,1547]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="1286" phylum="Arthropoda" rank="species" species="janczyki">
<emphasis id="50C39F0EFFE0FFB7CFC4FA6BFDA5F998" box="[477,588,1525,1547]" italics="true" pageId="1" pageNumber="1286">L. janczyki</emphasis>
</taxonomicName>
is doubtful; according to ANLAŞ (pers. comm.), Lagistan is situated in northeastern
<collectingCountry id="1AA0038CFFE0FFB7CC04F98FFD9DF9BA" box="[541,628,1554,1576]" name="Turkey" pageId="1" pageNumber="1286">Anatolia</collectingCountry>
and the
<typeStatus id="BD0CFDBEFFE0FFB7CCD4F98FFCCFF9BA" box="[717,806,1554,1576]" pageId="1" pageNumber="1286" type="holotype">holotype</typeStatus>
was probably found somewhere in
<collectingRegion id="A0738DFEFFE0FFB7CF09F9B2FE8DF9D7" box="[272,356,1583,1605]" country="Turkey" name="Trabzon" pageId="1" pageNumber="1286">Trabzon</collectingRegion>
or
<collectingRegion id="A0738DFEFFE0FFB7CF9FF9B2FE5DF9D7" box="[390,436,1583,1605]" country="Turkey" name="Rize" pageId="1" pageNumber="1286">Rize</collectingRegion>
provinces.
</paragraph>
<paragraph id="6208431CFFE3FFB4CEABFF46FDE8FDA1" blockId="2.[178,1085,219,563]" pageId="2" pageNumber="1287">
A comparison of the
<typeStatus id="BD0CFDBEFFE3FFB4CF94FF46FE50FF62" box="[397,441,219,240]" pageId="2" pageNumber="1287">type</typeStatus>
material of
<taxonomicName id="A5B7389FFFE3FFB4CC20FF41FD41FF62" authorityName="PACE" authorityYear="1983" box="[569,680,219,241]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1287" phylum="Arthropoda" rank="species" species="janczyki">
<emphasis id="50C39F0EFFE3FFB4CC20FF41FD41FF62" box="[569,680,219,241]" italics="true" pageId="2" pageNumber="1287">L. janczyki</emphasis>
</taxonomicName>
and
<taxonomicName id="A5B7389FFFE3FFB4CCC5FF41FCB2FF62" authorityName="ASSING" authorityYear="2007" box="[732,859,219,241]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1287" phylum="Arthropoda" rank="species" species="soganlica">
<emphasis id="50C39F0EFFE3FFB4CCC5FF41FCB2FF62" box="[732,859,219,241]" italics="true" pageId="2" pageNumber="1287">L. soganlica</emphasis>
</taxonomicName>
revealed some difference in the male primary and secondary sexual characters. In
<taxonomicName id="A5B7389FFFE3FFB4CD30FF64FC71FE9F" authorityName="PACE" authorityYear="1983" box="[809,920,248,270]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1287" phylum="Arthropoda" rank="species" species="janczyki">
<emphasis id="50C39F0EFFE3FFB4CD30FF64FC71FE9F" box="[809,920,248,270]" italics="true" pageId="2" pageNumber="1287">L. janczyki</emphasis>
</taxonomicName>
, the male elytra (
<figureCitation id="FA8C5F99FFE3FFB4CEA0FE88FF11FEB9" box="[185,248,277,299]" captionStart="Figs 1-9" captionStartId="6.[178,218,1351,1371]" captionTargetBox="[184,1071,219,1311]" captionTargetId="figure-127@6.[178,1085,219,1331]" captionTargetPageId="6" captionText="Figs 1-9: Leptusa janczyki PACE, holotype (1-3) and L. flagellulifera nov.sp. (4-9): (1) pronotum and elytra; (2) male tergite VII; (3, 6-7) median lobe of aedeagus in lateral view; (4) habitus; (5) forebody; (8) apical lobe of paramere; (9) spermatheca. Scale bars: 4: 1.0 mm; 1-2, 5: 0.5 mm; 3, 6- 9: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276619" httpUri="https://zenodo.org/record/5276619/files/figure.png" pageId="2" pageNumber="1287">Fig. 1</figureCitation>
) have elevations on either side of the scutellum (absent in
<taxonomicName id="A5B7389FFFE3FFB4CD9BFE8BFBEAFEB8" authorityName="ASSING" authorityYear="2007" box="[898,1027,277,299]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1287" phylum="Arthropoda" rank="species" species="soganlica">
<emphasis id="50C39F0EFFE3FFB4CD9BFE8BFBEAFEB8" box="[898,1027,277,299]" italics="true" pageId="2" pageNumber="1287">L. soganlica</emphasis>
</taxonomicName>
), the tubercle on the male tergite VII (
<figureCitation id="FA8C5F99FFE3FFB4CC38FEAFFD8AFEDA" box="[545,611,306,328]" captionStart="Figs 1-9" captionStartId="6.[178,218,1351,1371]" captionTargetBox="[184,1071,219,1311]" captionTargetId="figure-127@6.[178,1085,219,1331]" captionTargetPageId="6" captionText="Figs 1-9: Leptusa janczyki PACE, holotype (1-3) and L. flagellulifera nov.sp. (4-9): (1) pronotum and elytra; (2) male tergite VII; (3, 6-7) median lobe of aedeagus in lateral view; (4) habitus; (5) forebody; (8) apical lobe of paramere; (9) spermatheca. Scale bars: 4: 1.0 mm; 1-2, 5: 0.5 mm; 3, 6- 9: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276619" httpUri="https://zenodo.org/record/5276619/files/figure.png" pageId="2" pageNumber="1287">Fig. 2</figureCitation>
) is separated from the posterior margin by approximately its length (situated directly at posterior margin in
<taxonomicName id="A5B7389FFFE3FFB4CD41FECCFC31FEF7" authorityName="ASSING" authorityYear="2007" box="[856,984,336,358]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1287" phylum="Arthropoda" rank="species" species="soganlica">
<emphasis id="50C39F0EFFE3FFB4CD41FECCFC31FEF7" box="[856,984,336,358]" italics="true" pageId="2" pageNumber="1287">L. soganlica</emphasis>
</taxonomicName>
), and the median lobe of the aedeagus (
<figureCitation id="FA8C5F99FFE3FFB4CFFEFEF0FDCCFE11" box="[487,549,365,387]" captionStart="Figs 1-9" captionStartId="6.[178,218,1351,1371]" captionTargetBox="[184,1071,219,1311]" captionTargetId="figure-127@6.[178,1085,219,1331]" captionTargetPageId="6" captionText="Figs 1-9: Leptusa janczyki PACE, holotype (1-3) and L. flagellulifera nov.sp. (4-9): (1) pronotum and elytra; (2) male tergite VII; (3, 6-7) median lobe of aedeagus in lateral view; (4) habitus; (5) forebody; (8) apical lobe of paramere; (9) spermatheca. Scale bars: 4: 1.0 mm; 1-2, 5: 0.5 mm; 3, 6- 9: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276619" httpUri="https://zenodo.org/record/5276619/files/figure.png" pageId="2" pageNumber="1287">Fig. 3</figureCitation>
) has a median process of somewhat different shape (more slender and apically less acute); for comparison see the illustrations of
<taxonomicName id="A5B7389FFFE3FFB4CA31FE16FEFDFE2F" authorityName="ASSING" authorityYear="2007" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1287" phylum="Arthropoda" rank="species" species="soganlica">
<emphasis id="50C39F0EFFE3FFB4CA31FE16FEFDFE2F" italics="true" pageId="2" pageNumber="1287">L. soganlica</emphasis>
</taxonomicName>
in
<bibRefCitation id="06263EEDFFE3FFB4CF2FFE35FE32FE2C" author="ASSING V" box="[310,475,424,446]" pageId="2" pageNumber="1287" pagination="7 - 19" refId="ref10697" refString="ASSING V. (2007 a): Three new species, three new synonyms, and additional records of Leptusa from Turkey (Insecta: Coleoptera: Staphylinidae: Aleocharinae). - Entomological Problems 37: 7 - 19." type="journal article" year="2007">ASSING (2007a)</bibRefCitation>
. Since these differences are not pronounced, the possibility that
<taxonomicName id="A5B7389FFFE3FFB4CEF8FE5BFE88FE48" authorityName="ASSING" authorityYear="2007" box="[225,353,453,475]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1287" phylum="Arthropoda" rank="species" species="soganlica">
<emphasis id="50C39F0EFFE3FFB4CEF8FE5BFE88FE48" box="[225,353,453,475]" italics="true" pageId="2" pageNumber="1287">L. soganlica</emphasis>
</taxonomicName>
and
<taxonomicName id="A5B7389FFFE3FFB4CF81FE5BFDE0FE48" authorityName="PACE" authorityYear="1983" box="[408,521,453,475]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1287" phylum="Arthropoda" rank="species" species="janczyki">
<emphasis id="50C39F0EFFE3FFB4CF81FE5BFDE0FE48" box="[408,521,453,475]" italics="true" pageId="2" pageNumber="1287">L. janczyki</emphasis>
</taxonomicName>
are conspecific can still not be ruled out completely. However, the observed differences are constant in the examined material, so that
<taxonomicName id="A5B7389FFFE3FFB4CA31FE7EFEFDFD87" authorityName="ASSING" authorityYear="2007" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1287" phylum="Arthropoda" rank="species" species="soganlica">
<emphasis id="50C39F0EFFE3FFB4CA31FE7EFEFDFD87" italics="true" pageId="2" pageNumber="1287">L. soganlica</emphasis>
</taxonomicName>
is regarded as a valid species for the time being. More material is needed to confirm or reject this hypothesis.
</paragraph>
</subSubSection>
</treatment>
</document>