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<document id="C7B11D911F5B9426AB716627C004E4D8" ID-CLB-Dataset="300292" ID-DOI="10.3897/zookeys.1208.122126" ID-GBIF-Dataset="92f37e90-7f3a-42f4-987d-aeae1889d378" ID-ZooBank="56CC7AA9-4B0A-4EA0-86A5-B066E6A57FC3" ID-publisher-id="122126" URI-arpha="994AC1D6-2908-5674-A0CD-B8A92786C6CB" URI-zoobank="http://zoobank.org/56CC7AA9-4B0A-4EA0-86A5-B066E6A57FC3" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1721924801512" checkinUser="pensoft" docAuthor="Doménech, Carles" docDate="2024" docId="EA156D06F21B585F9BDF75B11DEB39C1" docLanguage="en" docName="ZooKeys 1208: 49-80" docOrigin="ZooKeys 1208" docSource="https://zookeys.pensoft.net/article/122126/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Scolopendra chlorotes L. Koch 1856" docType="treatment" docVersion="4" dtd-version="3.0" lastPageNumber="80" masterDocId="994AC1D629085674A0CDB8A92786C6CB" masterDocTitle="Taxonomic and nomenclatural reassessment of the Iberian Peninsulas nomina obscura, Scolopendra viridipes Dufour, 1820 and S. chlorotes L. Koch in Rosenhauer, 1856 (Chilopoda, Scolopendromorpha, Scolopendridae)" masterLastPageNumber="80" masterPageNumber="49" pageNumber="49" updateTime="1721938564851" updateUser="ExternalLinkService">
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<mods:title id="AF76264472919E6DB6F3489E4D44595A">Taxonomic and nomenclatural reassessment of the Iberian Peninsulas nomina obscura, Scolopendra viridipes Dufour, 1820 and S. chlorotes L. Koch in Rosenhauer, 1856 (Chilopoda, Scolopendromorpha, Scolopendridae)</mods:title>
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<mods:namePart id="BB7F901F96A02C9915040BB279763A02">Doménech, Carles</mods:namePart>
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<mods:affiliation id="CA4830743D4925BBCD667955BA51765B">Departament de Ciències Ambientals i Recursos Naturals, Universitat dAlacant, Carretera de Sant Vicent del Raspeig s / n C. P. 03690, San Vicent del Raspeig, Alacant, Spain</mods:affiliation>
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<treatment id="EA156D06F21B585F9BDF75B11DEB39C1" ID-DOI="http://doi.org/10.5281/zenodo.12827514" ID-Zenodo-Dep="12827514" ID-arpha="EA156D06-F21B-585F-9BDF-75B11DEB39C1" LSID="urn:lsid:plazi:treatment:EA156D06F21B585F9BDF75B11DEB39C1" httpUri="http://treatment.plazi.org/id/EA156D06F21B585F9BDF75B11DEB39C1">
<subSubSection id="E9655B554A3CBE700DC782327A7E89E4" type="nomenclature">
<paragraph id="56D51985C07537218C7D1DBF8852B8E6">
<taxonomicName id="22ABDDB314476065F9DAD073D390D724" ID-CoL="4VS6H" authority="L. Koch in Rosenhauer, 1856" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="13139E0A4937662477FB93D0227101F2" italics="true">Scolopendra chlorotes</emphasis>
L. Koch in
<bibRefCitation id="5AB2528DD6920713944AA6DF5D8BB596" DOI="10.5962/bhl.title.66016" author="Rosenhauer WG" journalOrPublisher="Verlag von Theodor Blaesing, Erlangen" refId="B65" refString="Rosenhauer WG (1856) Die Thiere Andalusiens nach dem resulte einer Reise. Ed. Verlag von Theodor Blaesing, Erlangen, 429 pp. https: // doi. org / 10.5962 / bhl. title. 66016" title="Die Thiere Andalusiens nach dem resulte einer Reise. Ed." year="1856">Rosenhauer, 1856</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="2DD8B6D44751B47A8FCDD64E9914F7E4" type="description">
<paragraph id="5ED1FB499FCFE050588717A1A71EB09E">
<figureCitation id="81F86F10FB5050A704A815B5855E2B0E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A topotype collection localities in the Iberian Peninsula (southwestern Europe): S. viridipes topotypes (= S. oraniensis); V 1 = Moixent and V 2 = Xàtiva (previously San Felipe), Valencia province, Valencian Country / Community, Spain and S. chlorotes topotypes (= S. oraniensis); C 1 = Málaga municipality, C 2 = Alahurín de la Torre, C 3 = Casabermeja, C 4 = Totalán and C 5 = Estepona, Málaga province, Andalucía, Spain. The “ O ” points S. oraniensis type localities, O 1 = Santons mountains [corrected from “ Sauton ” (sic.); Lucas 1846] and O 2 = “ Between Oran and Mers-el-Kebir ”, Oran province (Wilayah), Algeria B “ S. viridipes ” topotype 2, female, (CEUAMr 22) from C Moixent, Alacant D “ S. chlorotes ” topotype 6, female, (CEUAMr 30) from E Alahurín de la Torre, Málaga." figureDoi="10.3897/zookeys.1208.122126.figure2" httpUri="https://binary.pensoft.net/fig/1101812">Figs 2 D, E</figureCitation>
,
<figureCitation id="8056250481F2A79A74E78C3A80030A06" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A cephalic plate, antennae, and TT 13 B forcipular segment and sternites 1, 2 C tooth plates D tergites 1014 E sternites 812. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure5" httpUri="https://binary.pensoft.net/fig/1101815">5</figureCitation>
,
<figureCitation id="E7DDAF90210FE1B6311DFDF4E66FBCE2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A ULBS s and UL s dorsal view B ULBS s and UL s ventral view C ULBS s, coxopleuron, and UL s ventral lateral view. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure6" httpUri="https://binary.pensoft.net/fig/1101816">6</figureCitation>
,
<figureCitation id="2368B3B7B9C005C9B5214D3F1F1C4B16" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Scolopendra oraniensis genitalia A male (CEUAMr 24); and B female (CEUAMr 31), and detail of the setae (arrows) on tarsus 1 (C, D) on the same specimens C male and D female. Observe that on the contrary that Würmli (1980) indicated, the numbers of setae are actually similar in both genders. Legend: AV, anal valve; G. gonopod (folded back); LA, lamina adanalis; LS, lamina subanalis; P, penis; SGS I, sternite of genital segment 1; SGS II, sternite of genital segment 2. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1208.122126.figure9" httpUri="https://binary.pensoft.net/fig/1101819">9 B, D</figureCitation>
,
<tableCitation id="04AB4C7417A8D3F7A029EC491D99985D" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes and S. chlorotes topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">Table 3</tableCitation>
<xref id="6559DB6B6359899B703F7269A1737F12" ref-type="supplementary-material" rid="S1">, Suppl. material 1: file 3</xref>
</paragraph>
</subSubSection>
<subSubSection id="8772E2451218B14F6C9113E0E4A8A954" type="reference_group">
<paragraph id="E5A9B867DF30D130EC2A8AB43A44BABE">
<treatmentCitationGroup id="70DC274E3B0983BD119D679C81A79BAA">
<treatmentCitation id="A316AD3F93C7510FF9F470327EDEF7F8" author="Rosenhauer WG" page="417" year="1856">
<taxonomicName id="3A65F9EF83EB9F5FD9E2EFB7029D304C" ID-CoL="4VS6H" authority="L. Koch in Rosenhauer, 1856: 417" authorityName="L. Koch" authorityPageNumber="417" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="A12EDCDECD507B5F1D37346EBEEC9AF8" italics="true">Scolopendra chlorotes</emphasis>
<comment id="B9C475C069C24A6F383B7EC41A1E2251">
L. Koch in
<bibRefCitation id="269366380312195AA723E83B38F9B558" DOI="10.5962/bhl.title.66016" author="Rosenhauer WG" journalOrPublisher="Verlag von Theodor Blaesing, Erlangen" refId="B65" refString="Rosenhauer WG (1856) Die Thiere Andalusiens nach dem resulte einer Reise. Ed. Verlag von Theodor Blaesing, Erlangen, 429 pp. https: // doi. org / 10.5962 / bhl. title. 66016" title="Die Thiere Andalusiens nach dem resulte einer Reise. Ed." year="1856">Rosenhauer, 1856: 417</bibRefCitation>
.
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="0B494BE800FBA1D02265A79F6396CBB2">
<treatmentCitationGroup id="3DDAC3F867380CDA9517E7EEF9D70A7A">
<comment id="A813476C3F510641946E942DAD2ED6CE">
?
<treatmentCitation id="1933841F3ABC4E751371FF0ED6497E4B">
<taxonomicName id="73B9FB5F380D72C331761330346ECEF3" ID-CoL="79XG2" authority="Lucas, 1846" authorityName="Lucas" authorityYear="1846" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="F7D694F43E8CAD189615DC8A1BB4EE1F" italics="true">S. oraniensis</emphasis>
<bibRefCitation id="9372442AC5AD00D8FA5FDC7FE8F44E64" author="Lucas PH" journalOrPublisher="Revue Zoologique" pagination="283-289" refId="B52" refString="Lucas PH (1846) Note sur quelques nouvelles especes d'insectes qui habitent les possessions francaises du Nord de l'Afrique. Revue Zoologique 9: 283 - 289." title="Note sur quelques nouvelles espèces dinsectes qui habitent les possessions francaises du Nord de lAfrique." volume="9" year="1846">Lucas, 1846</bibRefCitation>
</taxonomicName>
</treatmentCitation>
:
<treatmentCitation id="691A13F4FF10CEBD37FAE590517CC771" author="Kraepelin K" page="246" year="1903">
<bibRefCitation id="80B4B0505A2FE09EA5A2C4E27FCCFA65" author="Kraepelin K" firstAuthor="Kraepelin" issue="2" journalOrPublisher="Mitteilungen aus dem Naturhistorischen Museum in Hamburg" page="246" pagination="1-276" refId="B40" refString="Kraepelin K (1903) Revision der Scolopendriden. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 20 (2): 1276." title="Revision der Scolopendriden." volume="20" year="1903">Kraepelin 1903: 246</bibRefCitation>
</treatmentCitation>
.
</comment>
</treatmentCitationGroup>
</paragraph>
<paragraph id="61D2E34E9BA76F6080691B0EEAD6318B">
<treatmentCitationGroup id="D6FAD5E900D0ED95778A19962F1C6CCE">
<taxonomicName id="A684B9C7AB66D5CEE4676E1C0B0039C0" ID-CoL="4VS6H" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="2FB8E921FC51A81F57F214D4821E883F" italics="true">S. chlorotes</emphasis>
</taxonomicName>
<comment id="D44CAC5F09D59A98016D22003B47B1C3">
:
<treatmentCitation id="77EDC8DF932AF8ADB0D0212BE87F8A05" author="Attems C" page="49" year="1930">
<bibRefCitation id="5DC5FAD9F54F37704189C882E8B660A9" DOI="10.1515/9783112373002" author="Attems C" firstAuthor="Attems" journalOrPublisher="De Gruyter, Berlin and Leipzig 54" page="49" refId="B1" refString="Attems C (1930) Myriopoda. 2. Scolopendromorpha. Das Tierreich. W. De Gruyter, Berlin and Leipzig 54, 308 pp. https://doi.org/10.1515/9783112373002" title="Myriopoda. 2. Scolopendromorpha. Das Tierreich. W." year="1930">Attems 1930: 49</bibRefCitation>
</treatmentCitation>
, as unrecognisable taxon.
</comment>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EF3CI" type="etymology">
<paragraph id="AC5CEF2B1BF2F6B64C703F742197572D">
<heading id="4AC7823E45773BA83DD3B7B57CF81725" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="F698ADFBAE169E86890A74BDE2331EA8">
From the Greek word
<emphasis id="675BB1C7D01B30FA1D17FA659BEA2F4F" italics="true">χλοερός</emphasis>
(
<emphasis id="6033F871FD0B8FD00D4CE21462B8BEB3" italics="true">khloerós</emphasis>
, verdant) and
<emphasis id="81396AC77C0C5FF75125DB0F0F7B4FB9" italics="true">χλόη</emphasis>
(
<emphasis id="63D066ED81ABD53F492778A3F5D45F61" italics="true">khlóē</emphasis>
, “ the green of new growth ”) meaning greenish yellow, pale green, pale, pallid, or verdant, referencing their pale greenish and yellowish leg colouration.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ET3CI" type="Type series and type depository">
<paragraph id="F1AB8028906C7DB86AB8D99251DEA969">
<heading id="042CB36E1E9CE857D1ED61C93F760BF9" reason="title">Type series and type depository.</heading>
</paragraph>
<paragraph id="C3C4DD49CFA260F573DDB5CB2A51A865">Types currently lost.</paragraph>
</subSubSection>
<subSubSection id="SECID0EY3CI" type="Collector and collection date">
<paragraph id="EED7302ADB4214371259F01CC49DAE0C">
<heading id="AD34777163568D59EC76DC259E825419" reason="title">Collector and collection date.</heading>
</paragraph>
<paragraph id="A9095E7324DA7D058D12D7033C2E4E3E">
W. G. Rosenhauer, in 1849 (
<bibRefCitation id="F7EFD71013DD0C777849A399CBBE5603" DOI="10.5962/bhl.title.66016" author="Rosenhauer" firstAuthor="Rosenhauer" journalOrPublisher="Verlag von Theodor Blaesing, Erlangen" refId="B65" refString="Rosenhauer WG (1856) Die Thiere Andalusiens nach dem resulte einer Reise. Ed. Verlag von Theodor Blaesing, Erlangen, 429 pp. https://doi.org/10.5962/bhl.title.66016" title="Die Thiere Andalusiens nach dem resulte einer Reise. Ed." year="1856">Rosenhauer 1856</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EC4CI" type="type locality">
<paragraph id="7C2E82DBA50D8680275E0FBFA0C0CF45">
<heading id="044CA4F92C768487DC52E2C4C596E4F7" reason="title">Type locality.</heading>
</paragraph>
<paragraph id="A292C3A81307ACC3A96341F359347902">
“ Near Málaga ”,
<collectingRegion id="72FC9CF6BF774C989E719057D536B3C2" country="Spain" name="Andalucia">Andalusia</collectingRegion>
,
<collectingCountry id="2ECFAF39FFE811153C51CFAAA844E82D" name="Spain">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EH4CI" type="distribution">
<paragraph id="CA819E1C175D32E65684D4D49DC69E5F">
<heading id="F3EC5BE6DB61DC1D11154509F5D97724" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="EF128213B18A426916BDCE9481270D86">As for type locality.</paragraph>
</subSubSection>
<subSubSection id="SECID0EM4CI" type="Neotype designation">
<paragraph id="C9C84ADA18E42C8A0930A78CBE2B7FA4">
<heading id="82486B48C718A65BD0CD7D3D1FBF0F87" reason="title">
<typeStatus id="23084EEF9775639D55DB27CC35FDB3F0" type="neotype">Neotype</typeStatus>
designation.
</heading>
</paragraph>
<paragraph id="5BE8163A49B1F4C66C54FF54E2278C5E">
With the express purpose of clarifying the taxonomic status and the type locality of the nominal taxon
<taxonomicName id="4423440C0090086DFFB27C275B496782" authority="L. Koch" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="CE05ED65FA2F0A9D01E91D2176EFCE35" italics="true">S. chlorotes</emphasis>
L. Koch
</taxonomicName>
in
<bibRefCitation id="BE2D3CB1A6D43F7042D88991B39FB748" DOI="10.5962/bhl.title.66016" author="Rosenhauer WG" journalOrPublisher="Verlag von Theodor Blaesing, Erlangen" refId="B65" refString="Rosenhauer WG (1856) Die Thiere Andalusiens nach dem resulte einer Reise. Ed. Verlag von Theodor Blaesing, Erlangen, 429 pp. https: // doi. org / 10.5962 / bhl. title. 66016" title="Die Thiere Andalusiens nach dem resulte einer Reise. Ed." year="1856">Rosenhauer, 1856</bibRefCitation>
, the following
<typeStatus id="4E6E455242CB7EDEF56F2675CC99C016" type="neotype">neotype</typeStatus>
is designated (
<bibRefCitation id="2654224C7991292AD68E1B6F135E788A" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN 1999</bibRefCitation>
: Art. 75 and 76.3):
</paragraph>
<paragraph id="3CED623B2C345076007874C4D186EF5A">
<materialsCitation id="E05C7134C6212EDB47C96EF310DFD22D" collectingDate="2021-11-18" collectionCode="CEUA" collectorName="C. Domenech" country="Spain" elevation="104" latitude="36.660278" location="Carretera de Coin" longitude="-4.516778" municipality="Alahurin de la Torre" specimenCount="1" specimenCount-male="1" stateProvince="Malaga province" typeStatus="neotype">
<specimenCount id="7CB87BCB30B3768B6855BAD536FC1D8B" count="1" type="male">Male</specimenCount>
.
<location id="79C96D2DFD1D21EC510B41C80F33DB9C" LSID="urn:lsid:plazi:treatment:EA156D06F21B585F9BDF75B11DEB39C1:79C96D2DFD1D21EC510B41C80F33DB9C" country="Spain" municipality="Alahurin de la Torre" name="Carretera de Coin" stateProvince="Malaga province">Carretera de Coín</location>
,
<collectingMunicipality id="F7EE3D6BFAF2728E6B4AFC665A7D9E69">Alahurín de la Torre</collectingMunicipality>
,
<collectingRegion id="5193588FA3F3242AEDD503C7D9B2FEBA">Málaga province</collectingRegion>
(
<collectingCountry id="B6E26ACBA1852737C4725C50D77BF511" name="Spain">Spain</collectingCountry>
) (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EG5CI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-4.516778,36.660278]}">
<geoCoordinate id="25BA0F19FEC80E935716DBBDF0038783" degrees="36" direction="north" minutes="39" orientation="latitude" seconds="37" value="36.660278">36 ° 39 ' 37 ' N</geoCoordinate>
,
<geoCoordinate id="EF24E92C5D6C576001ED58DADC7B8AFF" degrees="4" direction="west" minutes="31" orientation="longitude" seconds="0.4" value="-4.516778">4 ° 31 ' 0.4 ' W</geoCoordinate>
</named-content>
<quantity id="46E7A08B7056E8AA6E8FB22D04041AB2" metricMagnitude="2" metricUnit="m" metricValue="1.04" unit="m" value="104.0">
<elevation id="1D0475FBD9FA7AE84D2F8F0521E8EA71" metricMagnitude="2" metricUnit="m" metricValue="1.04" unit="m" value="104.0">104 m</elevation>
</quantity>
a. s. l.) (Figs
<figureCitation id="62FAC77F6F3AB48D109FE206C06F7FA3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A cephalic plate, antennae, and TT 13 B forcipular segment and sternites 1, 2 C tooth plates D tergites 1014 E sternites 812. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure5" httpUri="https://binary.pensoft.net/fig/1101815">5</figureCitation>
,
<figureCitation id="75C5CAC83F213C733BB33B80BEAD8EAB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A ULBS s and UL s dorsal view B ULBS s and UL s ventral view C ULBS s, coxopleuron, and UL s ventral lateral view. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure6" httpUri="https://binary.pensoft.net/fig/1101816">6</figureCitation>
; Table
<tableCitation id="568D876FFF1A1AC893014205166BDCA0" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes and S. chlorotes topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">3</tableCitation>
; Suppl. material
<xref id="9F161E0F31CAE50EE843C80B6EF3C2AB" ref-type="supplementary-material" rid="S1">1</xref>
: file 1). Coll.
<date id="C5FDC76DDCF0C17373FD82601263B0F7" value="2021-11-18">
<collectingDate id="3D4D90867254BDDDB962A8D7191881EF" value="2021-11-18">18 Nov. 2021</collectingDate>
</date>
.
<collectorName id="ED57680780CDEB13C8EF33EA4E8469F5">C. Doménech</collectorName>
Leg. Repository
<named-content id="C7B873C59BF00C130B390226811796BC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidad-de-alicante" xlink_title="Colección Entomológica de la Universidad de Alicante San Vicent del Raspeig">
<collectionCode id="9A8ED2F3847E00ED162B028A65F1026F">CEUA</collectionCode>
</named-content>
with the collection number CEUAMr 29 (In this text also referred as “
<taxonomicName id="6D88672534FF9F32DBF0D4A8F8F14E18" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="E243797F4CA38CF717AAC46BEBDBC544" italics="true">S. chlorotes</emphasis>
</taxonomicName>
topotype 5 &quot; before its
<typeStatus id="0FD92501053080B0E6C4F082CD326E65" type="neotype">neotype</typeStatus>
designation)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EK6CI" type="Proposed new nomenclatural status">
<paragraph id="C2CBBBA0FF94318A38949A2509A5E7BF">
<heading id="C59EF7B459089AA0B90AE57E62E738FB" reason="title">Proposed new nomenclatural status.</heading>
</paragraph>
<paragraph id="0638F79590F2693EC2CC2BA8B52C35C4">
<taxonomicName id="AAFB010DB7462F1BE17C52150D94EAB7" authority="L. Koch" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="B94354CABEBEA41A0D89E4A8ECEEC212" italics="true">Scolopendra chlorotes</emphasis>
L. Koch
</taxonomicName>
in
<bibRefCitation id="E96CB761A18896E50278C2A80EF6575C" DOI="10.5962/bhl.title.66016" author="Rosenhauer WG" journalOrPublisher="Verlag von Theodor Blaesing, Erlangen" refId="B65" refString="Rosenhauer WG (1856) Die Thiere Andalusiens nach dem resulte einer Reise. Ed. Verlag von Theodor Blaesing, Erlangen, 429 pp. https: // doi. org / 10.5962 / bhl. title. 66016" title="Die Thiere Andalusiens nach dem resulte einer Reise. Ed." year="1856">Rosenhauer, 1856</bibRefCitation>
is an invalid name, here subjectively proposed as a junior synonym of
<taxonomicName id="F00A06B2A330F912B945009792ED6A58" authority="Lucas, 1846" authorityName="Lucas" authorityYear="1846" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="39D74F02D6705412D9D8F2DF1D764B56" italics="true">Scolopendra oraniensis</emphasis>
<bibRefCitation id="22B7914956696C503083E58A72B1BF43" author="Lucas PH" journalOrPublisher="Revue Zoologique" pagination="283-289" refId="B52" refString="Lucas PH (1846) Note sur quelques nouvelles especes d'insectes qui habitent les possessions francaises du Nord de l'Afrique. Revue Zoologique 9: 283 - 289." title="Note sur quelques nouvelles espèces dinsectes qui habitent les possessions francaises du Nord de lAfrique." volume="9" year="1846">Lucas, 1846</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGADI" type="description">
<paragraph id="2EE14F666ED883DC487F81C4E0159172">
<heading id="7399796883069AFF3A6910733183DB27" reason="title">Translation of the original descriptions from German</heading>
</paragraph>
<paragraph id="BD65C11F9494A2C95E66B812AF6FBCB8">
[annotations in brackets]. (Original description available from:
<ext-link id="467EC26F79CE0529F081AC5EAF6E53DE" ext-link-type="uri" xlink_href="https://www.biodiversitylibrary.org/page/42185817#page/425/mode/1up" xlink_type="simple">https://www.biodiversitylibrary.org/page/42185817#page/425/mode/1up</ext-link>
)
</paragraph>
<paragraph id="00F16A1DD073F2EE470FA04761CDB74A">
<taxonomicName id="6CF73617686FA93C89BBACB103377407" authority="Koch." authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="652DC6BBDA70330E9C2F8D8AAA45D2CC" italics="true">Scolopendra chlorotes</emphasis>
Koch.
</taxonomicName>
</paragraph>
<paragraph id="87C3580336C15888574618E6C28454D7">
Brownish green, feeding pliers [forcipules], end-shield [
<abbrev id="ABBRID0EABDI" xlink_title="ultimate leg-bearing segment">
<collectionCode id="63B938E3E187D6CF4AD109AB69719559">ULBS</collectionCode>
</abbrev>
tergite] and last pair of legs [terminal legs] reddish brown, on the last seven tergites, a furrow at the lateral edges [margination in tergites], on the first podomere of the last pair of legs [
<abbrev id="ABBRID0EEBDI" xlink_title="ultimate legs">UL</abbrev>
prefemur] 19 small teeth on the underside [ventral position], seven small teeth directed inward on the upper-side [medial and dorsomedial positions].
</paragraph>
<paragraph id="E46DCBC9146889B9F0BA7B2D5F0B5EB4">
Length 20 lignes [1 German Nuremberg lignes ≈
<quantity id="D55C6205FCB22A1D686AE94F8717AE40" metricMagnitude="-3" metricUnit="m" metricValue="2.11" unit="mm" value="2.11">2.11 mm</quantity>
;
<quantity id="10EA42502862CC174231B746FEFC4514" metricMagnitude="-2" metricUnit="m" metricValue="4.2219999999999995" unit="mm" value="42.22">42.22 mm</quantity>
].
</paragraph>
<paragraph id="4290EC828715EA962856F9F4B9888A5C">
Shiny; head [cephalic plate] longish, rather narrow, the head area dentate [probably referring to tooth plates] in the middle, tergite sides straight, those of the 3
<superScript id="DCB35A245560331895BBBC055E177BE5">rd</superScript>
and the penultimate tergite rounded anteriorly, the seven last tergites with a furrow at the lateral edges [tergite margination]. Tergites with the two normal stripes [paramedian sutures], except the first and the last tergites; the end-shield [
<abbrev id="ABBRID0ENBDI" xlink_title="ultimate leg-bearing segment">
<collectionCode id="41032DCEF3038C8160C15CAD969DC639">ULBS</collectionCode>
</abbrev>
tergite] shows a distinct median longitudinal furrow [suture]. The sternites have two longitudinal furrows [paramedian sutures], except the last sternite. The last pair of legs short, dorsally flat, the tooth-like process at the inner angle of the first podomere short [
<abbrev id="ABBRID0ERBDI" xlink_title="ultimate legs">UL</abbrev>
prefemoral process] with two blunt teeth [spines] at the tip; at the inner side [medial position] of this podomere [
<abbrev id="ABBRID0EVBDI" xlink_title="ultimate legs">UL</abbrev>
prefemur] seven small teeth [spines] at the upper side [medial position], 19 small teeth [spines] arranged in four rows at the underside [ventral position]. Head [cephalic plate], antennae, and tergites, except the last one, brownish green, the first podomeres [prefemur, femur, and even tibia] of the legs yellowish, the last [tibia and tarsi 1 and 2] green; maxillipeds, lower lip [probably trochanterprefemoral parts of the forcipula, tooth plates or / and coxosternite], end-shield [
<abbrev id="ABBRID0EZBDI" xlink_title="ultimate leg-bearing segment">
<collectionCode id="E3ED180F4AD2810C04DBE2C2E275ECFE">ULBS</collectionCode>
</abbrev>
tergite] and last pair of legs [
<abbrev id="ABBRID0E4BDI" xlink_title="ultimate legs">UL</abbrev>
] reddish brown, the capture-claw [forcipules] brownish black from the middle [tarsungula].
</paragraph>
<paragraph id="AB9C163C7486B37154CE73C7555F2B1F">Near Malaga, sporadic.</paragraph>
</subSubSection>
<subSubSection id="SECID0ECCDI" type="remarks">
<paragraph id="D56BF1B69266963463909B95FC062E08">
<heading id="16C5638F312B69C1B037947E8ED5F463" reason="title">Remarks involving the type series and type series depository.</heading>
</paragraph>
<paragraph id="363856CB322834F419322310106CB248">
All attempts to locate the type series of
<taxonomicName id="F6D6C40E94167851F17A35AC1D27F14F" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="06B7D36585210288BFFF0633AEEF2BE4" italics="true">S. chlorotes</emphasis>
</taxonomicName>
were unsuccessful; the curators in their four probable depositories (
<abbrev id="ABBRID0ETCDI" xlink_title="Naturhistorisches Museum Nürnberg">
<collectionCode id="291066727B6549DCBAE7E7C6C4CE2684" country="United Kingdom" httpUri="http://grbio.org/cool/m8e8-jkbv" name="Nottingham Natural History Museum (Wollaton Hall)" type="">NHMN</collectionCode>
</abbrev>
,
<named-content id="316C602543004763CFE8AA3ECE2FFC9C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="C8E31E0590156B9E83F45308B3CB5619" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
,
<named-content id="BAE4B09E124D2F848B88AEECB5344CDC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museum-f%C3%BCr-naturkunde-berlin-zoological-collections" xlink_title="Museum für Naturkunde, Berlin">
<collectionCode id="6FC9956721A269C4A91BADF1F00ED6FC" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" type="Museum">ZMB</collectionCode>
</named-content>
, and
<abbrev id="ABBRID0E6CDI" xlink_title="Friedrich-Alexander Universität">
<collectionCode id="F052514D581E6FDCB5D11974CB5A552E" country="USA" lsid="urn:lsid:biocol.org:col:15720" name="Florida Atlantic University" type="Herbarium">FAU</collectionCode>
</abbrev>
) found no samples in their collections (pers. comm.
<date id="602854FD74AF7AD9B5C537F3986BE486" value="2020-08">Aug. 2020</date>
to
<date id="4EAFD5C0250958AF0BA09732544B048D" value="2021-11">Nov. 2021</date>
).
</paragraph>
<paragraph id="B7B20276A003E0C9251ADDEA81543091">
According to
<bibRefCitation id="FAE329113E0910D3CDB042C65A584462" author="Rühm J" firstAuthor="Ruhm" refId="B66" refString="Rühm J (1925) Der Neurenberger Naturforscher Dr. Ludwig Koch. Ein Gedenktblatt zum seinem 100. Geburstage am 8 Nov. 1925. Naturhistorischen Gesellschaft zu Nürnberg, 13 pp." year="1925">Rühm (1925)</bibRefCitation>
and
<bibRefCitation id="2269B227B98D9FE6B490EBD4510ACBFA" author="Hessel B" firstAuthor="Hessel" refId="B31" refString="Hessel B (2000) Dr. Karl Ludwig Christian Koch (8. 11. 1825 - 1. 11. 1908) Eine kurze Übersicht über das Leben und Wirken eines großen Nürnberger Naturforschers. Naturhistorische Gesellschaft Nürnberg e. V. 3946." year="2000">Hessel (2000)</bibRefCitation>
the types of this species most probable depository should be the
<named-content id="7E2E409ECFE4B6A8685B6FFEB88637FD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="Natural History Museum, London">
<collectionCode id="B007E08B0373CA09DD02BD1E6C02B842" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
, on the basis that in 1925 the Naturhistorische Gesellschaft (
<collectionCode id="4AE2FBDAE4FBA5E156B44229C6A7D682" country="Germany" lsid="urn:lsid:biocol.org:col:15855" name="Naturhistorische Gesellschaft e. V." type="Herbarium">NHG</collectionCode>
), lacking financial resources, decided to move a large number of their specimens preserved in ethanol to the British Museum of Natural History. Nevertheless, the types of
<taxonomicName id="79D202859CCA7DC989D5C258BA69D9A5" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="8DC209E435102454D962FD7F6A80E45D" italics="true">S. chlorotes</emphasis>
</taxonomicName>
are not registered nor found there.
</paragraph>
<paragraph id="280933207F7889EDDBE0BCAA1C6BC17E">
The other smaller part of Kochs wet collection, probably containing this species type series, did remain at the
<abbrev id="ABBRID0E5DDI" xlink_title="Naturhistorisches Museum Nürnberg">
<collectionCode id="F3DAEE9EEED7F0B3666267F446E79C7D" country="United Kingdom" httpUri="http://grbio.org/cool/m8e8-jkbv" name="Nottingham Natural History Museum (Wollaton Hall)" type="">NHMN</collectionCode>
</abbrev>
. However, the NHG rooms, their catalogues, and a large part of the collections were damaged during World War II in 1945 and presumably also the
<taxonomicName id="8478120558BDCE0BB68F0767D4F973E3" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="B8F2E5EEEA2FC34E33C3537EA5653B46" italics="true">S. chlorotes</emphasis>
</taxonomicName>
types (EM Neupert (
<collectionCode id="DC8881754D0537443A5BC470ECEB0C85" country="Germany" lsid="urn:lsid:biocol.org:col:15855" name="Naturhistorische Gesellschaft e. V." type="Herbarium">NHG</collectionCode>
) pers. comm.
<date id="D5CCAC1FB589A0B8F1CF360FF17EB9AF" value="2020-09">Sep. 2020</date>
). Therefore, the type series of
<taxonomicName id="42C7B26B324B32ADD80C3F3BB231D9F2" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="7A0D9B350D6CF928D3BCCF37CF7F9641" italics="true">S. chlorotes</emphasis>
</taxonomicName>
is considered to be lost.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYEDI" type="description">
<paragraph id="C6147801A66A1DED8C19A4F81B1C9C0C">
<heading id="FB02D63603041B89F25966EDB7248A4E" reason="title">Original description comparison.</heading>
</paragraph>
<paragraph id="828A4AEB02FB8A511F8CCB4FFD377DE8">
Because the type material is unavailable, the features in the original description are compared to those of other
<taxonomicName id="8B01EFF8AC8A137762AFBD168423861E" authorityName="Pocock" authorityYear="1895" class="Chilopoda" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="order">Scolopendromorpha</taxonomicName>
found in Peninsular Spain to determine to which taxon L. Koch was referring when he erected
<taxonomicName id="463B252DAAC9BEFA4B03EFD8A2D6A53F" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="F6D9BA381151F18062E4BE7E6C3E9C91" italics="true">S. chlorotes</emphasis>
</taxonomicName>
as a new species (Table
<tableCitation id="0A00E27F82429FB8FF742D8F72E6AD13" captionStart="Table 2" captionStartId="T2" captionText="Table 2. The original descriptive morphology of Scolopendra chlorotes in comparison to other Scolopendromorpha species described in Peninsular Spain. Bold letters indicate compatible features in other species. * indicates common to the entire Scolopendromorpha, interpretable, or poorly differentiated feature. Nr pp = Not reported, presence possible. NA = Information not available." httpUri="http://table.plazi.org/id/83E668CADC992295344F956E110DB611" tableUuid="83E668CADC992295344F956E110DB611">2</tableCitation>
). In this case, the original description of
<taxonomicName id="61B49E33AED94E739F6ADAB19D674504" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="77B5C668A775088E2CA216AE3E5C560F" italics="true">S. chlorotes</emphasis>
</taxonomicName>
allows exclusion of all
<taxonomicName id="5B53A4FC81CB18B1960912C1B8591F74">
<emphasis id="F96BBB44E0D859D15908443633A7E219" italics="true">Cryptops</emphasis>
species
</taxonomicName>
because the sternite sutures are cruciform or trigonal (rather than only the two paramedial ones) and the
<abbrev id="ABBRID0EEGDI" xlink_title="ultimate legs">UL</abbrev>
prefemoral spines and (most likely referred to as) coxosternal tooth plates are absent in this genus (
<bibRefCitation id="1C5645A726C8B9B0AE66E25E39E0CDEC" DOI="10.11646/zootaxa.4825.1.1" author="Schileyko" etAl="et al." firstAuthor="Schileyko" issue="1" journalOrPublisher="Zootaxa" pagination="1-64" refId="B67" refString="Schileyko AA, Vathera V, Edgecombe GD (2020) An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): A new identification key and updated diagnoses. Zootaxa 4825 (1): 164. https://doi.org/10.11646/zootaxa.4825.1.1" title="An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): A new identification key and updated diagnoses." volume="4825" year="2020">Schileyko et al. 2020</bibRefCitation>
). Neither of the two Mediterranean representatives of the family
<taxonomicName id="C463358821888422D8D53EE83F403B67" authorityName="Bollman" authorityYear="1893" class="Chilopoda" family="Plutoniumidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="family">Plutoniumidae</taxonomicName>
match with the morphological description of
<taxonomicName id="73BD6C177F5673624AC34C8A43260375" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="BF3F8516A00366BA218D191C9547E74C" italics="true">S. chlorotes</emphasis>
</taxonomicName>
since both have sternites with a distinctive single medial longitudinal suture and none, or just one, spine on
<abbrev id="ABBRID0E3GDI" xlink_title="ultimate legs">UL</abbrev>
prefemur (
<bibRefCitation id="6CEB4ACCD0F66FC7E46A7DDE04BB5A0D" DOI="10.1093/zoolinnean/zlw026" author="Bonato L &amp; Orlando M &amp; Zapparoli M &amp; Fusco G &amp; Bartolin F" issue="4" journalOrPublisher="Zoological Journal of the Linnean Society" pagination="887-909" refId="B6" refString="Bonato L, Orlando M, Zapparoli M, Fusco G, Bartolin F (2017 b) New insights into Plutonium, one of the largest and least known European centipedes (Chilopoda): Distribution, evolution and morphology. Zoological Journal of the Linnean Society 180 (4): 887909. https://doi.org/10.1093/zoolinnean/zlw026" title="New insights into Plutonium, one of the largest and least known European centipedes (Chilopoda): Distribution, evolution and morphology." volume="180">Bonato et al. 2017 b</bibRefCitation>
). The compatibility of
<taxonomicName id="5E71642143AEEC80EAF72FA4DDED0D9F" authorityName="Latreille" authorityYear="1829" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="cingulata">
<emphasis id="E8778F8F5508C9D1390E1C4B8CBD00B8" italics="true">S. cingulata</emphasis>
</taxonomicName>
is also rejected due to the absence of the longitudinal suture on the T 21 with the incompatibilities in the prefemoral spinulation of the
<abbrev id="ABBRID0EPHDI" xlink_title="ultimate legs">UL</abbrev>
(Table
<tableCitation id="AADFECD0B9DC0DFC839DE40A4524B2EA" captionStart="Table 2" captionStartId="T2" captionText="Table 2. The original descriptive morphology of Scolopendra chlorotes in comparison to other Scolopendromorpha species described in Peninsular Spain. Bold letters indicate compatible features in other species. * indicates common to the entire Scolopendromorpha, interpretable, or poorly differentiated feature. Nr pp = Not reported, presence possible. NA = Information not available." httpUri="http://table.plazi.org/id/83E668CADC992295344F956E110DB611" tableUuid="83E668CADC992295344F956E110DB611">2</tableCitation>
). Furthermore, all of these taxa can be also ruled out because their colours do not match with those of
<taxonomicName id="FAA640FA5CE218B9B0DA406E162D93D1" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="EA718795D657ED1137B1DC8C067B3D1B" italics="true">S. chlorotes</emphasis>
</taxonomicName>
(Table
<tableCitation id="DC3AE1E1014DF3D099A8B4035B2E93C5" captionStart="Table 2" captionStartId="T2" captionText="Table 2. The original descriptive morphology of Scolopendra chlorotes in comparison to other Scolopendromorpha species described in Peninsular Spain. Bold letters indicate compatible features in other species. * indicates common to the entire Scolopendromorpha, interpretable, or poorly differentiated feature. Nr pp = Not reported, presence possible. NA = Information not available." httpUri="http://table.plazi.org/id/83E668CADC992295344F956E110DB611" tableUuid="83E668CADC992295344F956E110DB611">2</tableCitation>
).
</paragraph>
<paragraph id="35EC5DDAE3BBE8809836527314DCEF75">
Hence, its unambiguous morphology and colouration, compatible location, and the exclusion of all other Iberian
<taxonomicName id="2DD93176FD6D8987CE4B140B27A9A44C" authorityName="Pocock" authorityYear="1895" class="Chilopoda" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="order">Scolopendromorpha</taxonomicName>
are facts that, combined, strongly suggest that if it is not the same taxon, the closest relative to
<taxonomicName id="0BD645C9EE1C4BC3EA07DB38E96B14FF" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="597F3CFA8D80108DBF2B01EFF76D0F21" italics="true">S. chlorotes</emphasis>
</taxonomicName>
is
<taxonomicName id="4F7DFD6165678180151FEDC37B3D676F" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="A0101C8CF0AB38E489F9DD4530AA460C" italics="true">S. oraniensis</emphasis>
</taxonomicName>
(Table
<tableCitation id="F083606777A8CB4D34A6E3208E780ED8" captionStart="Table 2" captionStartId="T2" captionText="Table 2. The original descriptive morphology of Scolopendra chlorotes in comparison to other Scolopendromorpha species described in Peninsular Spain. Bold letters indicate compatible features in other species. * indicates common to the entire Scolopendromorpha, interpretable, or poorly differentiated feature. Nr pp = Not reported, presence possible. NA = Information not available." httpUri="http://table.plazi.org/id/83E668CADC992295344F956E110DB611" tableUuid="83E668CADC992295344F956E110DB611">2</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHJDI" type="Neotype and &quot;topotypes&quot; morphological comparisons">
<paragraph id="476112517ACED52F612CD34FBFD7D406">
<heading id="32B3BB40E39F115CB003096D0FD7F04E" reason="title">
<typeStatus id="026C6D8C37DA7C4EF36BD4D19F23FD51" type="neotype">Neotype</typeStatus>
and “
<typeStatus id="7DD032C8DAFD285B3DE09AF9C2F8E753" type="topotype">topotypes</typeStatus>
” morphological comparisons.
</heading>
</paragraph>
<paragraph id="C9A16422378806E8FEB89F8E616968E5">
A total of fifteen
<taxonomicName id="878CF608D4949598C3EEC3289AC800AA" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="365A734B0D4661E6BA4C288F7E8E6749" italics="true">S. chlorotes</emphasis>
</taxonomicName>
topotypical specimens were collected in five municipalities “ near Málaga ” (L.
<bibRefCitation id="4B3F03C792A94EDCEA96F19B83B9022A" author="Koch" firstAuthor="Koch" refId="B39" refString="Koch L (1856) . Myriapoda. In: Die Thiere Andalusiens. Rosenhauer WG (Ed.) Erlangen, 416 pp." year="1856">Koch 1856</bibRefCitation>
; Figs
<figureCitation id="A30F04A4B06BFAB3F9C39996D2E72B4A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A topotype collection localities in the Iberian Peninsula (southwestern Europe): S. viridipes topotypes (= S. oraniensis); V 1 = Moixent and V 2 = Xàtiva (previously San Felipe), Valencia province, Valencian Country / Community, Spain and S. chlorotes topotypes (= S. oraniensis); C 1 = Málaga municipality, C 2 = Alahurín de la Torre, C 3 = Casabermeja, C 4 = Totalán and C 5 = Estepona, Málaga province, Andalucía, Spain. The “ O ” points S. oraniensis type localities, O 1 = Santons mountains [corrected from “ Sauton ” (sic.); Lucas 1846] and O 2 = “ Between Oran and Mers-el-Kebir ”, Oran province (Wilayah), Algeria B “ S. viridipes ” topotype 2, female, (CEUAMr 22) from C Moixent, Alacant D “ S. chlorotes ” topotype 6, female, (CEUAMr 30) from E Alahurín de la Torre, Málaga." figureDoi="10.3897/zookeys.1208.122126.figure2" httpUri="https://binary.pensoft.net/fig/1101812">2 A, D, E</figureCitation>
,
<figureCitation id="383BB8A965566BBADD8F8A4BC676C945" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A cephalic plate, antennae, and TT 13 B forcipular segment and sternites 1, 2 C tooth plates D tergites 1014 E sternites 812. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure5" httpUri="https://binary.pensoft.net/fig/1101815">5</figureCitation>
,
<figureCitation id="D1D44D653C6831966B1BC1E0888B5FAC" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A ULBS s and UL s dorsal view B ULBS s and UL s ventral view C ULBS s, coxopleuron, and UL s ventral lateral view. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure6" httpUri="https://binary.pensoft.net/fig/1101816">6</figureCitation>
; Suppl. material
<xref id="0AFED488C535EB3F9719EC4BC7D6EAC7" ref-type="supplementary-material" rid="S1">1</xref>
: file 3). All these specimens were examined and confirmed to be conspecific (Suppl. material
<xref id="A67EAEECCFD808B719AC0593C16B881C" ref-type="supplementary-material" rid="S1">1</xref>
: file 3). Among those, a
<typeStatus id="EFBC9A4303635A161560B31B6E0ADC8E" type="neotype">neotype</typeStatus>
was selected (
<bibRefCitation id="CECE486854942648BC4ACB043C19F56D" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN 1999</bibRefCitation>
: Art. 75; Figs
<figureCitation id="6A9D8B2C5A5D0216B552FB3C73014AAC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A cephalic plate, antennae, and TT 13 B forcipular segment and sternites 1, 2 C tooth plates D tergites 1014 E sternites 812. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure5" httpUri="https://binary.pensoft.net/fig/1101815">5</figureCitation>
,
<figureCitation id="CE3D6DC4B9235272740766D39435B627" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A ULBS s and UL s dorsal view B ULBS s and UL s ventral view C ULBS s, coxopleuron, and UL s ventral lateral view. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure6" httpUri="https://binary.pensoft.net/fig/1101816">6</figureCitation>
, Table
<tableCitation id="7DF044375FA38C6515A146EE91B06CA5" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes and S. chlorotes topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">3</tableCitation>
, Suppl. material
<xref id="1558DBF1295C99B934C166E7E82B3FBC" ref-type="supplementary-material" rid="S1">1</xref>
: file 3). The comparison of the
<typeStatus id="D1EE11B44A402A3FB984EC1A9FFB3F1C" type="neotype">neotype</typeStatus>
and the other “
<typeStatus id="D38246EEDEFD298111A3FB66628036CB" type="topotype">topotypes</typeStatus>
” with the
<specimenCount id="0CA55C0CA619E0C9D0F1E831583A7A17" count="7" type="generic" typeStatus="syntype">seven syntypes</specimenCount>
of the closest taxon
<taxonomicName id="FD820FAF0D35AC8E42A6F5BDD7B9E1A3" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="9881A7BA124B154BEEF5256F8BF6C7C4" italics="true">S. oraniensis</emphasis>
</taxonomicName>
demonstrated that all these specimens possess identical diagnostic morphological features (see diagnosis and redescription above; Table
<tableCitation id="94B9FB20FB49CA9D4433804FADB8F99B" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes and S. chlorotes topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">3</tableCitation>
; compare Figs
<figureCitation id="1F12DD4FAB3A34DC44370064A4491EF0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A cephalic plate, antennae, and TT 13 B forcipular segment and sternites 1, 2 C tooth plates D tergites 1014 E sternites 812. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure5" httpUri="https://binary.pensoft.net/fig/1101815">5</figureCitation>
,
<figureCitation id="57BAF5B84FBA7CA9D6A58383220C97D4" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A ULBS s and UL s dorsal view B ULBS s and UL s ventral view C ULBS s, coxopleuron, and UL s ventral lateral view. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure6" httpUri="https://binary.pensoft.net/fig/1101816">6</figureCitation>
with Figs
<figureCitation id="32C775A1B5EF5BA3512A349E0BAC2586" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A cephalic plate, antennae, and T 1 B forcipular segment and sternites 1, 2 C tooth plates D tergites 36 E sternites 68. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure7" httpUri="https://binary.pensoft.net/fig/1101817">7</figureCitation>
,
<figureCitation id="770CC604350CD533C6E3594F866F6D05" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelins 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8</figureCitation>
) and therefore, both taxa should be considered to be the same species.
</paragraph>
<paragraph id="BCDA9C1E993790DF573D99B4404382AD">
Additionally, the
<typeStatus id="DC2D201A9B6850C1F29C83E5E56E5F12" type="neotype">neotype</typeStatus>
of
<taxonomicName id="1ECE95F84EF90EE7CB8A3BF02DE2588C" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="C9242A10406B63BCD4BE890D3E7C5983" italics="true">S. chlorotes</emphasis>
</taxonomicName>
and the other “
<typeStatus id="1E86B93D2F758F8D4FBA2EBC93C99486" type="topotype">topotypes</typeStatus>
” were compared with those of
<taxonomicName id="AF5807AED2BE4DD4D42C139769B74859" authorityName="Dufour" authorityYear="1820" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="viridipes">
<emphasis id="7D9A0A71CE7032C7C5B9FDF85C13FCC4" italics="true">S. viridipes</emphasis>
</taxonomicName>
, and conspecificity was also confirmed (compare Figs
<figureCitation id="A13044E8AD10FC37F00E57BFB69808B9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra viridipes neotype (= S. oraniensis); male; (“ topotype 1 &quot;, CEUAMr 21) A cephalic plate, antennae, and TT 15 B forcipular segment and sternites 13 C tooth plates D tergites 1318 E sternites 1015. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure3" httpUri="https://binary.pensoft.net/fig/1101813">3</figureCitation>
,
<figureCitation id="A6BC03FAB6CF0C8325C606A882DC07E0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopendra viridipes neotype (= S. oraniensis); male; (“ topotype 1 &quot;, CEUAMr 21) A ULBS and UL s dorsal view B ULBS s and UL s ventral view C ULBS s, coxopleuron, and UL s ventral lateral view. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure4" httpUri="https://binary.pensoft.net/fig/1101814">4</figureCitation>
with Figs
<figureCitation id="2479676D883D10E109EA8F6ABEB62B31" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A cephalic plate, antennae, and TT 13 B forcipular segment and sternites 1, 2 C tooth plates D tergites 1014 E sternites 812. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure5" httpUri="https://binary.pensoft.net/fig/1101815">5</figureCitation>
,
<figureCitation id="A7CCBAB11D0F03EBB9DBBF91CA6F7729" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopendra chlorotes neotype (= S. oraniensis); male; (“ topotype 5 &quot;, CEUAMr 29) A ULBS s and UL s dorsal view B ULBS s and UL s ventral view C ULBS s, coxopleuron, and UL s ventral lateral view. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure6" httpUri="https://binary.pensoft.net/fig/1101816">6</figureCitation>
; Table
<tableCitation id="0980B9905379872C98CC661235C83311" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes and S. chlorotes topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">3</tableCitation>
). Consequently,
<taxonomicName id="248904E1AB8B46BA401B9BA69BC2C70A" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="9D38B2F14D6ACEDD165C633A381DDCEE" italics="true">S. chlorotes</emphasis>
</taxonomicName>
,
<taxonomicName id="FA97336EE9A9FD6163AAD6806EDEC55F" authorityName="Dufour" authorityYear="1820" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="viridipes">
<emphasis id="A012450D8526B4DDE75A2D7B4917067E" italics="true">S. viridipes</emphasis>
</taxonomicName>
, and
<taxonomicName id="2A4228A7FBD4F2C06911A76C8E1D0B42" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="DE7CC49BC4F1BCCC51F29320065020D0" italics="true">S. oraniensis</emphasis>
</taxonomicName>
are considered a single taxon with three names (Figs
<figureCitation id="4795248197098747345C2071F14B90AC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopendra viridipes neotype (= S. oraniensis); male; (“ topotype 1 &quot;, CEUAMr 21) A cephalic plate, antennae, and TT 15 B forcipular segment and sternites 13 C tooth plates D tergites 1318 E sternites 1015. Scale bars: 0.2 mm (C); 1 mm (A, B); 2 mm (D, E)." figureDoi="10.3897/zookeys.1208.122126.figure3" httpUri="https://binary.pensoft.net/fig/1101813">3</figureCitation>
<figureCitation id="2D62B578479DDCB0EED5D4BD1D41D469" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Scolopendra oraniensis syntype “ 1 &quot;; unsexed; (MNHN) A dorsal view of ULBS and UL B ventral view of ULBS and UL C lateral view of ULBS and UL D Original manuscript labels ordered chronologically. Top: old label [Scolopendra canidens Newp. oraniensis Luc. H. Lucas. [Leg.] Oran [type locality]; Middle: Kraepelins 1903 labels [“ Type de H. Lucas. Scol. oraniensis Luc. (non canidens Newp.) ”]. Lower: MNHN staff “ modern ” summary label. Scale bars: 1 mm (A, B); 2 mm (C)." figureDoi="10.3897/zookeys.1208.122126.figure8" httpUri="https://binary.pensoft.net/fig/1101818">8</figureCitation>
; Table
<tableCitation id="6EC62E832CD7E0E9E3F97753DFB29D10" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Scolopendra oraniensis type series morphological comparison with S. viridipes and S. chlorotes topotypes (= S. oraniensis; composite data from Supplementary tables 1 and 2). AP, apical spines; SAP, subapical spines; DS, dorsal spines; LS, lateral spines; VL, ventro-lateral spines; V, ventral spines; VM, ventro-median spines; M, median spines; DM, dorso-median spiness; SP, prefemoral process spines; UL; ultimate legs, ULBS, ultimate leg-bearing segment; T, tergite; TT, tergites, S, sternite; SS, sternites, RG, Retracted genitalia; * No visible (cephalic plate flexed over firsts SS), interpretable or damaged." httpUri="http://table.plazi.org/id/3DA5B8200171122C71CB5BEFCC3C311D" tableUuid="3DA5B8200171122C71CB5BEFCC3C311D">3</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3ODI" type="Taxonomic and nomenclatural status">
<paragraph id="76ADCEE42C369BE5430C5A3B56C63940">
<heading id="89F1F09D2089B4E4D16A02AFBF97405F" reason="title">Taxonomic and nomenclatural status.</heading>
</paragraph>
<paragraph id="BE8058F74F9C6AE35BC055A3AE129F30">
As long as the taxon
<taxonomicName id="6D8FFC79676569510C8C687B05F62E78" authority="L. Koch" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="5DBA2EC9FA8460AB6BEA5D8195F83A8F" italics="true">S. chlorotes</emphasis>
L. Koch
</taxonomicName>
in
<bibRefCitation id="4C1F616CF35D0B0DF37E87DFEBAA155B" DOI="10.5962/bhl.title.66016" author="Rosenhauer WG" journalOrPublisher="Verlag von Theodor Blaesing, Erlangen" refId="B65" refString="Rosenhauer WG (1856) Die Thiere Andalusiens nach dem resulte einer Reise. Ed. Verlag von Theodor Blaesing, Erlangen, 429 pp. https: // doi. org / 10.5962 / bhl. title. 66016" title="Die Thiere Andalusiens nach dem resulte einer Reise. Ed." year="1856">Rosenhauer, 1856</bibRefCitation>
is recognised as conspecific with
<taxonomicName id="089B1C49A986E6C5D5D5C082425724EA" authority="Lucas, 1846" authorityName="Lucas" authorityYear="1846" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="EFCD60264C2D8FF82759DD8CCA751068" italics="true">S. oraniensis</emphasis>
<bibRefCitation id="D28A504FBBBFE77D47A4A4C26DAAFEE5" author="Lucas PH" journalOrPublisher="Revue Zoologique" pagination="283-289" refId="B52" refString="Lucas PH (1846) Note sur quelques nouvelles especes d'insectes qui habitent les possessions francaises du Nord de l'Afrique. Revue Zoologique 9: 283 - 289." title="Note sur quelques nouvelles espèces dinsectes qui habitent les possessions francaises du Nord de lAfrique." volume="9" year="1846">Lucas, 1846</bibRefCitation>
</taxonomicName>
, the principle of priority indicates that the valid name of
<taxonomicName id="FEBD3C88B457CF83F7BF27078DDA2C38" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="C48511B3714955B4B88482441DB9B038" italics="true">S. chlorotes</emphasis>
</taxonomicName>
is
<taxonomicName id="33782A1CDA1A096A4BFF159576A75A3F" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="09544AEC7ECCC59D07D7931382B27ABB" italics="true">S. oraniensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="C4ABCDAD0EFC0E97273ABDF23974608E" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN 1999</bibRefCitation>
: Art. 23.3). Hence,
<taxonomicName id="F8246A77279CBA767E7C10EA855E688A" authorityName="L. Koch" authorityYear="1856" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="chlorotes">
<emphasis id="BAD3551D5317C7F8E0822FC261274B41" italics="true">S. chlorotes</emphasis>
</taxonomicName>
is subjectively designated as an invalid name since it is here considered a junior synonym of
<taxonomicName id="983C32D4759B896C79095FAB77C0A32C" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="oraniensis">
<emphasis id="AD682A1F2109702693C289721E3527DE" italics="true">S. oraniensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="FDE45811D738E46545945F540069D705" refId="B32" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK. https://www.iczn.org/the-code/the-code-online/" url="https://www.iczn.org/the-code/the-code-online/" year="1999">ICZN 1999</bibRefCitation>
: Art. 23.3).
</paragraph>
</subSubSection>
</treatment>
</document>