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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.62253" ID-GBIF-Dataset="0a292fd0-be2d-427e-b277-c19af79971db" ID-PMC="PMC8222342" ID-Pensoft-Pub="1313-2970-1044-153" ID-Pensoft-UUID="944578388D005036B42D68229F152A89" ID-PubMed="34183876" ID-ZooBank="A2219253DB2B4C5992D677595A6C1438" ModsDocID="1313-2970-1044-153" checkinTime="1623875446139" checkinUser="pensoft" docAuthor="Maddison, David R. &amp; Porch, Nick" docDate="2021" docId="E93DED16DE9F58DDBCEAD5E300B8C612" docLanguage="en" docName="ZooKeys 1044: 153-196" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.62253" docTitle="Tasmanitachoides baehri Maddison &amp; Porch 2021, sp. nov." docType="treatment" docUuid="D4F6DB85-62FE-47DA-A67E-91C2FB54BA2B" docUuidSource="ZooBank" docVersion="5" id="944578388D005036B42D68229F152A89" lastPageNumber="153" masterDocId="944578388D005036B42D68229F152A89" masterDocTitle="A preliminary phylogeny and review of the genus Tasmanitachoides, with descriptions of two new species (Coleoptera, Carabidae, Bembidarenini)" masterLastPageNumber="196" masterPageNumber="153" pageNumber="153" updateTime="1732566981577" updateUser="ExternalLinkService">
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<mods:title id="C02DA172CF55F5D3ED8670247BB7A5DD">A preliminary phylogeny and review of the genus Tasmanitachoides, with descriptions of two new species (Coleoptera, Carabidae, Bembidarenini)</mods:title>
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<mods:name id="803A640EC6F47E143B3C656FC95D2D30" type="personal">
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<mods:namePart id="96EE2D596E841D21F1907EDC655258A4">Maddison, David R.</mods:namePart>
<mods:nameIdentifier id="3FB8477C86067F146A4F524234415C2C" type="ORCID">https://orcid.org/0000-0002-7152-3824</mods:nameIdentifier>
<mods:affiliation id="F5F3929A413224E0BB56B5298A95A7FE">Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA</mods:affiliation>
<mods:nameIdentifier id="6EE40AF7571D0EFB018C900616B98A09" type="email">david.maddison@science.oregonstate.edu</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="364F524FE9353966752485BD0572AF67">Porch, Nick</mods:namePart>
<mods:affiliation id="8988F714D0CEB8F0D730CC202E4859B9">School of Life and Environmental Sciences, Deakin University, Geelong 3216, Australia</mods:affiliation>
</mods:name>
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<mods:date id="F05F7FC6FA54FCAE3EECF62008A6E907">2021</mods:date>
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<mods:number id="66813B852B550AF8281B286D7290DDFD">2021-06-16</mods:number>
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<mods:number id="48249625DC1E4F5D40305A99F50566B9">1044</mods:number>
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</mods:location>
<mods:classification id="9838ACEA9FD090957FA76009A812DBB0">journal article</mods:classification>
<mods:identifier id="DEF6B3F7D4B7873325C69BAC8CD1A1DB" type="DOI">http://dx.doi.org/10.3897/zookeys.1044.62253</mods:identifier>
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<mods:identifier id="EE9A98172A89CF12920975AD8F4C7BA6" type="Pensoft-UUID">944578388D005036B42D68229F152A89</mods:identifier>
</mods:mods>
<treatment id="E93DED16DE9F58DDBCEAD5E300B8C612" ID-GBIF-Taxon="182754123" LSID="urn:lsid:zoobank.org:act:D4F6DB85-62FE-47DA-A67E-91C2FB54BA2B" httpUri="http://treatment.plazi.org/id/E93DED16DE9F58DDBCEAD5E300B8C612" lastPageNumber="153" pageId="0" pageNumber="153" scope_family="Carabidae" scope_order="Coleoptera" scope_tribe="Bembidarenini">
<subSubSection id="6C795D6EB96BBF752E51842E519E06E1" pageId="0" pageNumber="153" type="nomenclature">
<paragraph id="573FCEF54292391B3A0F74828B6FAC13" pageId="0" pageNumber="153">
<taxonomicName id="A90B4FF273D8BAA43A397B9209C7A33A" ID-CoL="8MDB9" LSID="http://zoobank.org/D4F6DB85-62FE-47DA-A67E-91C2FB54BA2B" authority="Maddison &amp; Porch, 2021" authorityName="Maddison &amp; Porch" authorityYear="2021" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides baehri" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="baehri" status="sp. nov.">Tasmanitachoides baehri</taxonomicName>
<taxonomicNameLabel id="E153265F0ED4D91DAC0969DE64387C24" pageId="0" pageNumber="153">sp. nov.</taxonomicNameLabel>
<figureCitation id="2CAC9DC557A5D9BADFFE8001CD02A324" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Adults of the Tasmanitachoides kingi species group as well as T. lutus A T. kingi, voucher V 101468 B T. angullicollis, voucher DNA 5515. C T. hendrichi, holotype. D T. wilsoni, voucher V 101470. E T. baehri, voucher V 101479. F T. lutus, voucher V 101462. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.62253.figure10" httpUri="https://binary.pensoft.net/fig/555132" pageId="0" pageNumber="153">Figures 10E</figureCitation>
<figureCitation id="CEE33D5A97AE31F61A1BAC6B4320F660" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left elytron of Tasmanitachoides A T. kingi, voucher V 101468 B T. angulicollis, voucher DNA 5515 C T. wilsoni, voucher V 101470 D T. baehri, voucher V 101479 E T. lutus, voucher V 101462 F T. katherinei, voucher V 101475 G T. cf. gerdi, voucher DNA 5676 H T. sp. &quot; Angle Crossing 1 &quot;, voucher DNA 5677 J T. obliquiceps, voucher V 101477 K T. mandibularis, voucher V 101473." figureDoi="10.3897/zookeys.1044.62253.figure15" httpUri="https://binary.pensoft.net/fig/555137" pageId="0" pageNumber="153">, 15D</figureCitation>
<figureCitation id="9B755E1FB1B02079EE5716B5242A6CFD" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dorsal view of head of Tasmanitachoides adults A T. wilsoni, voucher V 101470 B T. baehri, voucher V 101479 C T. lutus, voucher V 101462 D T. cf. gerdi, voucher DNA 5676 E T. sp. &quot; Angle Crossing 1 &quot;, voucher DNA 5677 F T. obliquiceps, voucher V 101477 G T. erwini, voucher V 101469 H T. hobarti, voucher V 101463 J T. leai, voucher V 101467 K T. rufescens, voucher V 101478 L T. fitzroyi, voucher V 101471 M T. murrumbidgensis, voucher V 101464." figureDoi="10.3897/zookeys.1044.62253.figure17" httpUri="https://binary.pensoft.net/fig/555139" pageId="0" pageNumber="153">, 17B</figureCitation>
<figureCitation id="42E5EF2604E75F8569B577FF30D34C5B" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Elytral microsculpture of Tasmanitachoides, dorsal view A T. baehri, voucher V 101484 B T. erwini, voucher V 101483. Scale bar: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure18" httpUri="https://binary.pensoft.net/fig/555140" pageId="0" pageNumber="153">, 18A</figureCitation>
<figureCitation id="77F4AD68A89AD5BECF8C25CB1042533F" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Aedeagus of Tasmanitachoides, ventral view A T. baehri, voucher DNA 5569 (holotype) B T. wilsoni, voucher DNA 5514 C T. erwini, voucher V 101481 D T. sp. &quot; Lerderderg R &quot;, voucher DNA 2029. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure19" httpUri="https://binary.pensoft.net/fig/555141" pageId="0" pageNumber="153">, 19A</figureCitation>
<figureCitation id="B25D7A182C0B989D6D37A5AF62BBCC1E" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Aedeagus of Tasmanitachoides, left lateral view A T. baehri, voucher DNA 2032 B T. erwini, voucher DNA 5583 C T. erwini, voucher DNA 5509 (holotype). Scale bar: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure20" httpUri="https://binary.pensoft.net/fig/555142" pageId="0" pageNumber="153">, 20A</figureCitation>
<figureCitation id="9575C58AFFDA35426C9EC7A187F4DBD5" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Known geographic distributions of Tasmanitachoides baehri and T. erwini in southeastern Australia. Circles: T. baehri; stars: T. erwini." figureDoi="10.3897/zookeys.1044.62253.figure21" httpUri="https://binary.pensoft.net/fig/555143" pageId="0" pageNumber="153">, 21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C727EE047E325F96782F81DFD1590156" pageId="0" pageNumber="153" type="materials_examined">
<paragraph id="520143241F467AD6F52D6E9DB519020E" pageId="0" pageNumber="153">Material examined.</paragraph>
<paragraph id="836B179DC7FAE9FA9379D8FFBAD26E20" pageId="0" pageNumber="153">
<materialsCitation id="F7FE4909B846DB8324E40CF7399BA998" accessionNumber="MW291166, MW291260, MW291213, MW291304" collectingDate="2002-09-28" collectorName="N. Porch, David R. Maddison" country="Australia" elevation="440" latitude="-35.245285" location="Male" longLatPrecision="1" longitude="148.95226" specimenCount="1" typeStatus="Holotype">
<emphasis id="953AEC7848C9B9AAF42FBC1847227EA2" bold="true" pageId="0" pageNumber="153">
<emphasis id="58264A2C1337009A32C5F3DF1F7D4F89" italics="true" pageId="0" pageNumber="153">
<typeStatus id="B1E79D5909C18A0722A318747D7E07B2">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<location id="629365FF2E808B91026CEAAF09704F9F" LSID="urn:lsid:plazi:treatment:E93DED16DE9F58DDBCEAD5E300B8C612:629365FF2E808B91026CEAAF09704F9F" country="Australia" latitude="-35.245285" longLatPrecision="1" longitude="148.95226" name="Male">Male</location>
(ANIC), labeled: &quot;
<collectingCountry id="E179CCDA74CBE05B6000B1E8A39EB89B" name="Australia">Australia</collectingCountry>
: ACT: Murrumbidgee River,
<quantity id="1AB9262EE62233EAB1776BD30852CF25" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="km" value="0.15">0.15 km</quantity>
u/s Uriarra Crossing (
<geoCoordinate id="73030D8F6A2B1FBA9595583CBC18F290" degrees="35" direction="south" minutes="14.717" orientation="latitude" precision="1" value="-35.245285">35°14.717'S</geoCoordinate>
,
<geoCoordinate id="EC127D8CC5905F9F4DCF6221CE457068" degrees="148" direction="east" minutes="57.135" orientation="longitude" precision="1" value="148.95226">148°57.135'E</geoCoordinate>
<quantity id="753E0576E2F73BCD5995A390E8B679FE" metricMagnitude="2" metricUnit="m" metricValue="4.4" unit="m" value="440.0">
<elevation id="8F55C44C824DF24BADD646131EA98D3E" metricMagnitude="2" metricUnit="m" metricValue="4.4" unit="m" value="440.0">440 m</elevation>
</quantity>
) Washed fr. gravel/under cobbles at river edge.
<collectorName id="FDB076D4B58D21BCBE6F1259F97E23E2">N. Porch</collectorName>
,
<collectingDate id="33D6080A262538D18BF6EEAD40F9C18B" value="2002-09-28">28 Sep. 2002</collectingDate>
&quot;, &quot;
<collectorName id="B886DA6B22FA92E21EC402D5AEF631CA">David R. Maddison</collectorName>
DNA5569 DNA Voucher&quot; [pale green paper], &quot;
<typeStatus id="FF147DE738007BFE025C407EF92AD8C0">HOLOTYPE</typeStatus>
<taxonomicName id="3FE244A0A36F5D1D8C729094C178C2C2" authorityName="Maddison &amp; Porch" authorityYear="2021" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides baehri" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="baehri">Tasmanitachoides baehri</taxonomicName>
Maddison &amp; Porch&quot; [partly handwritten, on red paper]. Genitalia mounted in Euparal on coverslip pinned with specimen; extracted DNA stored separately. GenBank accession numbers for DNA sequences of the
<typeStatus id="F81487ADCC947A2651302ED89FDBE7B8">holotype</typeStatus>
are
<accessionNumber id="EE4D15C6A21D7A16C0D8ABEFCD77CB8F" isEnumeration="true">MW291166, MW291260, MW291213</accessionNumber>
, and
<accessionNumber id="8BB428277DA8EFBABC35C08229EFC46E" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW291304">MW291304</accessionNumber>
.
</materialsCitation>
</paragraph>
<paragraph id="F9A1DE623B87CF058E9A6CAB4A84A90A" pageId="0" pageNumber="153">
<materialsCitation id="409F3C21C3B5517E79A5B93EE4C17C11" accessionNumber="MW291166, MW291260, MW291213, MW291304" collectingDate="2000-10-14" collectingDateMax="2002-09-28" collectingDateMin="2000-10-14" collectorName="N. Porch, Washed" country="Australia" county="Murrumbidgee River" elevation="440" latitude="-35.245285" location="Same" longLatPrecision="1" longitude="148.95226" municipality="Uriarra Crossing" specimenCount="1" typeStatus="Paratypes">
<emphasis id="6776EC996FB28A3ADE19F7C2A94BA047" bold="true" italics="true" pageId="0" pageNumber="153">
<typeStatus id="273522016A4D4E333F6DC3975BD61CDC">Paratypes</typeStatus>
</emphasis>
(26).
<location id="A022BB24D71DF60BCE29BE87C97B61F0" LSID="urn:lsid:plazi:treatment:E93DED16DE9F58DDBCEAD5E300B8C612:A022BB24D71DF60BCE29BE87C97B61F0" country="Australia" county="Murrumbidgee River" latitude="-35.245285" longLatPrecision="1" longitude="148.95226" municipality="Uriarra Crossing" name="Same">Same</location>
label data as holotype (8; ANIC, OSAC). ACT:
<location id="2A16EFD416E971BA9D01D28C0EA7D6C2" LSID="urn:lsid:plazi:treatment:E93DED16DE9F58DDBCEAD5E300B8C612:2A16EFD416E971BA9D01D28C0EA7D6C2" country="Australia" county="Murrumbidgee River" latitude="-35.245285" longLatPrecision="1" longitude="148.95226" municipality="Uriarra Crossing" name="Murrumbidgee River">Murrumbidgee River</location>
,
<quantity id="0028BB1259C3006156E5FC7674A1AAB2" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="km" value="0.15">0.15 km</quantity>
u/s
<collectingMunicipality id="FD4DEC3AF398EDB83B200BCE22C16D2A">Uriarra Crossing</collectingMunicipality>
(
<geoCoordinate id="B28400BC0C65E2C4792FC2CBC9EA0457" degrees="35" direction="south" minutes="14.717" orientation="latitude" precision="1" value="-35.245285">35°14.717'S</geoCoordinate>
,
<geoCoordinate id="E39AE36DE17DD944C2107675F728B80D" degrees="148" direction="east" minutes="57.135" orientation="longitude" precision="1" value="148.95226">148°57.135'E</geoCoordinate>
<quantity id="EAD9E4C6F7164C155964FF6ECE7492D6" metricMagnitude="2" metricUnit="m" metricValue="4.4" unit="m" value="440.0">
<elevation id="43AB9140CAEFFE6658C9F8F3B60A2299" metricMagnitude="2" metricUnit="m" metricValue="4.4" unit="m" value="440.0">440 m</elevation>
</quantity>
). Washed fr. gravel/under cobbles at edge of river.
<collectorName id="0CB9AFD6000B3E7C554E353B8046A9FB">N. Porch</collectorName>
,
<collectingDate id="F2DAE306C40B48DB024CD7DE3A4AB91B" value="2000-10-14">14 Oct. 2000</collectingDate>
(10; NPC, ANIC, ZSM, NMV, NHMUK, MCZ). ACT:
<collectingCounty id="B45EAE369D38CA9A8D131C799EFDE255">Murrumbidgee River</collectingCounty>
,
<quantity id="BEA047A4BAD63CDA07F5EE804F8B8645" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="km" value="0.15">0.15 km</quantity>
u/s Uriarra Crossing (
<geoCoordinate id="6F62A44DB7957AB883614D9582ECB49F" degrees="35" direction="south" minutes="14.717" orientation="latitude" precision="1" value="-35.245285">35°14.717'S</geoCoordinate>
,
<geoCoordinate id="000EEF92B7A81FB54DD3FA0099593AE1" degrees="148" direction="east" minutes="57.135" orientation="longitude" precision="1" value="148.95226">148°57.135'E</geoCoordinate>
<quantity id="3B8AB0C6FB17D1B54260C20BB8085244" metricMagnitude="2" metricUnit="m" metricValue="4.4" unit="m" value="440.0">
<elevation id="B600683D74BEB11B867C7EDAE786D244" metricMagnitude="2" metricUnit="m" metricValue="4.4" unit="m" value="440.0">440 m</elevation>
</quantity>
).
<collectorName id="3DC7A9562FA79E165EEACD735D457AAA">Washed</collectorName>
fr. gravel/under cobbles at edge of river.
<collectorName id="2B957DA7841438CE4978870E34F55913">N. Porch</collectorName>
,
<collectingDate id="2645E557202022C21175A7C4FDA3B592" value="2002-09-28">28 Sep. 2002</collectingDate>
(8; NPC, ANIC, ZSM, NHMUK, MCZ)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="68F988DABB6D32575434A3B3B4A02520" pageId="0" pageNumber="153" type="materials_examined">
<paragraph id="19AC5FDE5DF4E1C2BDA05534BD283BA0" pageId="0" pageNumber="153">Other material examined.</paragraph>
<paragraph id="F4E4F00A7506A91EA80D8001A765CB2F" pageId="0" pageNumber="153">
We have seen an addition specimen labeled
<normalizedToken id="3B439FD247594E638F17F5B2CC4BA7DE" originalValue="“Paddys">&quot;Paddy's</normalizedToken>
River,
<quantity id="BA78E67B6389F5C1095A27F9DB6B946D" metricMagnitude="3" metricUnit="m" metricValue="1.609344" unit="mi" value="1.0">1 mi.</quantity>
S. of Cotter Dam, ACT,
<date id="7C4CCE1EC7B80DE404EBB86F9937B628" value="1969-04-17">17.iv.1969</date>
. S. Misko&quot; (ANIC; currently in ZSM).
</paragraph>
</subSubSection>
<subSubSection id="BAA92E1A6D83C2FE42B327CD96BA4C60" pageId="0" pageNumber="153" type="type locality">
<paragraph id="D8A286A6C7C523F518B5012209B8FD3D" pageId="0" pageNumber="153">Type locality.</paragraph>
<paragraph id="70C9230CBFBF1B312D705F5482035408" pageId="0" pageNumber="153">
Australia: ACT: Murrumbidgee River, 0.15 km u/s Uriarra Crossing (
<geoCoordinate id="9BD410DDD85920BA4C6006BE95642950" degrees="35" direction="south" minutes="14.717" orientation="latitude" precision="1" value="-35.245285">35°14.717'S</geoCoordinate>
,
<geoCoordinate id="01C4DF246FC25E96CEFD59999217080F" degrees="148" direction="east" minutes="57.135" orientation="longitude" precision="1" value="148.95226">148°57.135'E</geoCoordinate>
440 m).
</paragraph>
</subSubSection>
<subSubSection id="39E301E3C5DE4F8F4B6C5FB35B3A16A6" pageId="0" pageNumber="153" type="derivation of specific epithet">
<paragraph id="EF4EC24E4E68BCB9A51B271487F5144D" pageId="0" pageNumber="153">Derivation of specific epithet.</paragraph>
<paragraph id="3F52273D49D970A0596268305FD71C19" pageId="0" pageNumber="153">
We are honored to name this species after the late Martin Baehr, who discovered and documented many of the carabid species of Australia, and who described 14 of the known species of
<taxonomicName id="53DE302F38051502FBACB03EE7A6375B" authorityName="Erwin" authorityYear="1972" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="D1A744E4BE3D6DD6CD69A8F5006D310F" italics="true" pageId="0" pageNumber="153">Tasmanitachoides</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="004BB51E51FC9E180BA4814ED76CA2EF" pageId="0" pageNumber="153" type="diagnosis">
<paragraph id="91C5F2065B57B208D5568E37A9E29CAB" pageId="0" pageNumber="153">Diagnosis and description.</paragraph>
<paragraph id="0F8DB576A3C34BAC3AE4BAC036036794" pageId="0" pageNumber="153">
Very small, length 1.59-1.63 mm (n = 4). A pale species, body mostly orange, with the front half of the elytra and head a darker reddish orange. Antennae pale testaceous, with antennomeres 5-11 slightly infuscated. Head with moderately long but shallow frontal furrows, reaching approximately the center of the eye, and at least to the anterior supraorbital seta (Fig.
<figureCitation id="26DF0554475F7B0720B32F66211815A2" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dorsal view of head of Tasmanitachoides adults A T. wilsoni, voucher V 101470 B T. baehri, voucher V 101479 C T. lutus, voucher V 101462 D T. cf. gerdi, voucher DNA 5676 E T. sp. &quot; Angle Crossing 1 &quot;, voucher DNA 5677 F T. obliquiceps, voucher V 101477 G T. erwini, voucher V 101469 H T. hobarti, voucher V 101463 J T. leai, voucher V 101467 K T. rufescens, voucher V 101478 L T. fitzroyi, voucher V 101471 M T. murrumbidgensis, voucher V 101464." figureDoi="10.3897/zookeys.1044.62253.figure17" httpUri="https://binary.pensoft.net/fig/555139" pageId="0" pageNumber="153">17B</figureCitation>
); with a groove extending from anterior supraorbital puncture anteriad and mediad to approximately halfway toward the frontal furrow (Fig.
<figureCitation id="CB9B9129D43AC38E4D736CE78D02021C" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dorsal view of head of Tasmanitachoides adults A T. wilsoni, voucher V 101470 B T. baehri, voucher V 101479 C T. lutus, voucher V 101462 D T. cf. gerdi, voucher DNA 5676 E T. sp. &quot; Angle Crossing 1 &quot;, voucher DNA 5677 F T. obliquiceps, voucher V 101477 G T. erwini, voucher V 101469 H T. hobarti, voucher V 101463 J T. leai, voucher V 101467 K T. rufescens, voucher V 101478 L T. fitzroyi, voucher V 101471 M T. murrumbidgensis, voucher V 101464." figureDoi="10.3897/zookeys.1044.62253.figure17" httpUri="https://binary.pensoft.net/fig/555139" pageId="0" pageNumber="153">17B</figureCitation>
). Pronotum convex, narrow, only slightly wider than head (Fig.
<figureCitation id="3EB2B0803C439EDD82A6F651CE6A8486" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Adults of the Tasmanitachoides kingi species group as well as T. lutus A T. kingi, voucher V 101468 B T. angullicollis, voucher DNA 5515. C T. hendrichi, holotype. D T. wilsoni, voucher V 101470. E T. baehri, voucher V 101479. F T. lutus, voucher V 101462. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1044.62253.figure10" httpUri="https://binary.pensoft.net/fig/555132" pageId="0" pageNumber="153">10E</figureCitation>
). Hind angle of pronotum obtuse. Elytra more parallel-sided than
<taxonomicName id="68076A7FAC5F4C91EB2481CE251550BA" lsidName="T. wilsoni" pageId="0" pageNumber="153" rank="species" species="wilsoni">
<emphasis id="5BEAAD0DA9C8F21F6BC9781BFDE77D81" italics="true" pageId="0" pageNumber="153">T. wilsoni</emphasis>
</taxonomicName>
. Striae 2 and 3 shallow, broad, impunctate grooves (Figs
<figureCitation id="D4EFFF008D759F983F77AA3EA168BB6F" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left elytron of Tasmanitachoides A T. kingi, voucher V 101468 B T. angulicollis, voucher DNA 5515 C T. wilsoni, voucher V 101470 D T. baehri, voucher V 101479 E T. lutus, voucher V 101462 F T. katherinei, voucher V 101475 G T. cf. gerdi, voucher DNA 5676 H T. sp. &quot; Angle Crossing 1 &quot;, voucher DNA 5677 J T. obliquiceps, voucher V 101477 K T. mandibularis, voucher V 101473." figureDoi="10.3897/zookeys.1044.62253.figure15" httpUri="https://binary.pensoft.net/fig/555137" pageId="0" pageNumber="153">15D</figureCitation>
,
<figureCitation id="8E38525494276B9705BDE53ED5C7CC02" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Elytral microsculpture of Tasmanitachoides, dorsal view A T. baehri, voucher V 101484 B T. erwini, voucher V 101483. Scale bar: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure18" httpUri="https://binary.pensoft.net/fig/555140" pageId="0" pageNumber="153">18A</figureCitation>
); nearby intervals convex. Stria 5 deeply engraved in anterior half of elytron; stria 5 reaching or nearly reaching the second discal seta (ed5; Fig.
<figureCitation id="1D4A39D70000BE6540E4B6DCA02C8433" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left elytron of Tasmanitachoides A T. kingi, voucher V 101468 B T. angulicollis, voucher DNA 5515 C T. wilsoni, voucher V 101470 D T. baehri, voucher V 101479 E T. lutus, voucher V 101462 F T. katherinei, voucher V 101475 G T. cf. gerdi, voucher DNA 5676 H T. sp. &quot; Angle Crossing 1 &quot;, voucher DNA 5677 J T. obliquiceps, voucher V 101477 K T. mandibularis, voucher V 101473." figureDoi="10.3897/zookeys.1044.62253.figure15" httpUri="https://binary.pensoft.net/fig/555137" pageId="0" pageNumber="153">15D</figureCitation>
). Striae 6 and 7 effaced. Discal setae ed6 apparently in stria 2. Microsculpture without engraved lines; where present on the dorsal surface, the microsculpture is formed as low papillae without defined boundaries (Fig.
<figureCitation id="A07B48CC72FD4DC033BD96441418C64B" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Elytral microsculpture of Tasmanitachoides, dorsal view A T. baehri, voucher V 101484 B T. erwini, voucher V 101483. Scale bar: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure18" httpUri="https://binary.pensoft.net/fig/555140" pageId="0" pageNumber="153">18A</figureCitation>
). Pronotum and head very shiny, virtually without microsculpture. Aedeagus (Figs
<figureCitation id="319F02EB67B6D36BC194929D7B116BDB" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Aedeagus of Tasmanitachoides, ventral view A T. baehri, voucher DNA 5569 (holotype) B T. wilsoni, voucher DNA 5514 C T. erwini, voucher V 101481 D T. sp. &quot; Lerderderg R &quot;, voucher DNA 2029. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure19" httpUri="https://binary.pensoft.net/fig/555141" pageId="0" pageNumber="153">19A</figureCitation>
,
<figureCitation id="7965D9EDA12111704789EC13510E0B56" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Aedeagus of Tasmanitachoides, left lateral view A T. baehri, voucher DNA 2032 B T. erwini, voucher DNA 5583 C T. erwini, voucher DNA 5509 (holotype). Scale bar: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure20" httpUri="https://binary.pensoft.net/fig/555142" pageId="0" pageNumber="153">20A</figureCitation>
) with internal sac sclerites compact, and sinuate, very similar to those of
<taxonomicName id="A08D011656A5A4650571050AF5D06E04" lsidName="T. wilsoni" pageId="0" pageNumber="153" rank="species" species="wilsoni">
<emphasis id="156D41A8EA529B84B9195CFE6C8FAC57" italics="true" pageId="0" pageNumber="153">T. wilsoni</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="4975DDB6F7F6B8F5F843FC1ED2772957" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Aedeagus of Tasmanitachoides, ventral view A T. baehri, voucher DNA 5569 (holotype) B T. wilsoni, voucher DNA 5514 C T. erwini, voucher V 101481 D T. sp. &quot; Lerderderg R &quot;, voucher DNA 2029. Scale bars: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure19" httpUri="https://binary.pensoft.net/fig/555141" pageId="0" pageNumber="153">19B</figureCitation>
).
</paragraph>
<caption id="2C333108F72D9B4B9C2333578FEB3A2E" doi="10.3897/zookeys.1044.62253.figure17" httpUri="https://binary.pensoft.net/fig/555139" pageId="0" pageNumber="153" start="Figure 17" startId="F17">
<paragraph id="AE5B19DB91F9CC30472665F3AA214E2C" pageId="0" pageNumber="153">
<emphasis id="AEB180307B36064255893564FD8D6B6A" bold="true" pageId="0" pageNumber="153">Figure 17.</emphasis>
Dorsal view of head of
<taxonomicName id="415F0ADAE91A77988F930BCC6056D4DE" authorityName="Erwin" authorityYear="1972" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="FD4CF3CFE48EA8FEBA84DEA516223D19" italics="true" pageId="0" pageNumber="153">Tasmanitachoides</emphasis>
</taxonomicName>
adults
<emphasis id="0CB15BC05C3AFA3702A2CAC5DEC28E8F" bold="true" pageId="0" pageNumber="153">A</emphasis>
<taxonomicName id="1AC7491DF6ADA1400A1D08B7A55CD44C" lsidName="T. wilsoni" pageId="0" pageNumber="153" rank="species" species="wilsoni">
<emphasis id="FBCB54AE89030C4CF9D8C48B18EE8A31" italics="true" pageId="0" pageNumber="153">T. wilsoni</emphasis>
</taxonomicName>
, voucher V101470
<emphasis id="2D399D54AAD783E02AC240568BCEFA6A" bold="true" pageId="0" pageNumber="153">B</emphasis>
<taxonomicName id="881721365DF933E6983E057F2721F518" lsidName="T. baehri" pageId="0" pageNumber="153" rank="species" species="baehri">
<emphasis id="6A383992D36D111DE5BC7069B27DF08E" italics="true" pageId="0" pageNumber="153">T. baehri</emphasis>
</taxonomicName>
, voucher V101479
<emphasis id="7B9178A6EEB8BEA6DCDD3F60FD0BBA68" bold="true" pageId="0" pageNumber="153">C</emphasis>
<taxonomicName id="989E93D59D3CEAAF9BC762951B870691" lsidName="T. lutus" pageId="0" pageNumber="153" rank="species" species="lutus">
<emphasis id="A8573BAC7FD1E64B08173321E556D31B" italics="true" pageId="0" pageNumber="153">T. lutus</emphasis>
</taxonomicName>
, voucher V101462
<emphasis id="5CAC0AB7C05395561BFA4DD7A57B6F10" bold="true" pageId="0" pageNumber="153">D</emphasis>
<taxonomicName id="0BB06EB203B79A4B4B38FDB1A0810118" lsidName="T. gerdi" pageId="0" pageNumber="153" rank="species" species="gerdi">T. cf. gerdi</taxonomicName>
, voucher DNA5676
<emphasis id="B9D74864912B35DFF727A9967BECAAF3" bold="true" pageId="0" pageNumber="153">E</emphasis>
<taxonomicName id="162195D44562E5A7BCE19C8C4CBDCDE9" genus="T." lsidName="T." pageId="0" pageNumber="153" rank="genus">
<emphasis id="7C90AEEDCC9DBB362F6D406EC955CDA3" italics="true" pageId="0" pageNumber="153">T.</emphasis>
</taxonomicName>
sp. &quot;Angle Crossing #1&quot;, voucher DNA5677
<emphasis id="BF047D9D042D8DD5241B2BA34E4CFCEC" bold="true" pageId="0" pageNumber="153">F</emphasis>
<taxonomicName id="13E8E380CD98AE045B40CA63B06BF4F5" lsidName="T. obliquiceps" pageId="0" pageNumber="153" rank="species" species="obliquiceps">
<emphasis id="24A5539B5B083E44CDCCEAC5354FCD0F" italics="true" pageId="0" pageNumber="153">T. obliquiceps</emphasis>
</taxonomicName>
, voucher V101477
<emphasis id="75EAB178283E9378F6D5AC2D852A1647" bold="true" pageId="0" pageNumber="153">G</emphasis>
<taxonomicName id="F91767EE0ADC2027AE953D6509E4BFBA" lsidName="T. erwini" pageId="0" pageNumber="153" rank="species" species="erwini">
<emphasis id="2DE2D658D6F22B764EF2ACD58C1C7336" italics="true" pageId="0" pageNumber="153">T. erwini</emphasis>
</taxonomicName>
, voucher V101469
<emphasis id="9233CF5E54B2D7D934A7C4E6BEFD7012" bold="true" pageId="0" pageNumber="153">H</emphasis>
<taxonomicName id="DB64949B53A06CAED82779A7F6186DBC" lsidName="T. hobarti" pageId="0" pageNumber="153" rank="species" species="hobarti">
<emphasis id="5234BAFFB12CA8F4E5AE3CCAF2A2EF6A" italics="true" pageId="0" pageNumber="153">T. hobarti</emphasis>
</taxonomicName>
, voucher V101463
<emphasis id="76EB18D0A835800E5D9578BB209D0341" bold="true" pageId="0" pageNumber="153">J</emphasis>
<taxonomicName id="D952376A5B06694C0713911F6BEB006E" lsidName="T. leai" pageId="0" pageNumber="153" rank="species" species="leai">
<emphasis id="CD3B8AE6678BD35BD73510BD8A39927A" italics="true" pageId="0" pageNumber="153">T. leai</emphasis>
</taxonomicName>
, voucher V101467
<emphasis id="60A6A0D720355E57D917D237F072239D" bold="true" pageId="0" pageNumber="153">K</emphasis>
<taxonomicName id="C6262EC548F172DB147F081C825ED50D" lsidName="T. rufescens" pageId="0" pageNumber="153" rank="species" species="rufescens">
<emphasis id="3A5135189FDCBB3C114999CB1DAE4D64" italics="true" pageId="0" pageNumber="153">T. rufescens</emphasis>
</taxonomicName>
, voucher V101478
<emphasis id="1648030DF465E9E802023821C28F8924" bold="true" pageId="0" pageNumber="153">L</emphasis>
<taxonomicName id="8432429EABEDBC586B8265C7AB1E1D5F" lsidName="T. fitzroyi" pageId="0" pageNumber="153" rank="species" species="fitzroyi">
<emphasis id="284300B2CC3C78E48A03D6E129672235" italics="true" pageId="0" pageNumber="153">T. fitzroyi</emphasis>
</taxonomicName>
, voucher V101471
<emphasis id="258F0370BA3EED10211EEA9DC5113CC5" bold="true" pageId="0" pageNumber="153">M</emphasis>
<taxonomicName id="98C1C8426DA5D65A07D05C7FEC202D54" lsidName="T. murrumbidgensis" pageId="0" pageNumber="153" rank="species" species="murrumbidgensis">
<emphasis id="AB93F762E395ECF21C2DEC1EB0191C74" italics="true" pageId="0" pageNumber="153">T. murrumbidgensis</emphasis>
</taxonomicName>
, voucher V101464.
</paragraph>
</caption>
<caption id="892E25D1CCA0DCE4FDA6E7B2B7081051" doi="10.3897/zookeys.1044.62253.figure18" httpUri="https://binary.pensoft.net/fig/555140" pageId="0" pageNumber="153" start="Figure 18" startId="F18">
<paragraph id="8BF3147373A49E90668BAD8B18934997" pageId="0" pageNumber="153">
<emphasis id="34CA970A0B7520801DEC1F3D8A8AC304" bold="true" pageId="0" pageNumber="153">Figure 18.</emphasis>
Elytral microsculpture of
<taxonomicName id="054A70F457E664B9B1A3CAFBFB40E3F1" authorityName="Erwin" authorityYear="1972" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="8292B9AEB623B31F63F28580E5995BD8" italics="true" pageId="0" pageNumber="153">Tasmanitachoides</emphasis>
</taxonomicName>
, dorsal view
<emphasis id="C9A5A929DA1D75F14791CA72C9262F1B" bold="true" pageId="0" pageNumber="153">A</emphasis>
<taxonomicName id="4CBE06A1CC17795273AAC06D102E8DA2" lsidName="T. baehri" pageId="0" pageNumber="153" rank="species" species="baehri">
<emphasis id="ABAC553ADD1D596361CACE816588E5D6" italics="true" pageId="0" pageNumber="153">T. baehri</emphasis>
</taxonomicName>
, voucher V101484
<emphasis id="633EA1E25F2363E18A40DF9DCC9C96C8" bold="true" pageId="0" pageNumber="153">B</emphasis>
<taxonomicName id="0CAD489B05DB27BD73575E161AC6CF31" lsidName="T. erwini" pageId="0" pageNumber="153" rank="species" species="erwini">
<emphasis id="B7A52DA8A3B1CD076B27446573BBB8C6" italics="true" pageId="0" pageNumber="153">T. erwini</emphasis>
</taxonomicName>
, voucher V101483. Scale bar: 100
<normalizedToken id="FD644F17925658987EE5093C6A42299D" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption id="F9D48FFCF32DB4CF690AECDA7265A318" doi="10.3897/zookeys.1044.62253.figure19" httpUri="https://binary.pensoft.net/fig/555141" pageId="0" pageNumber="153" start="Figure 19" startId="F19">
<paragraph id="2F88CD85F2C394B411186C6ACA826181" pageId="0" pageNumber="153">
<emphasis id="1114CEA311282A41324A2480CFD733F8" bold="true" pageId="0" pageNumber="153">Figure 19.</emphasis>
Aedeagus of
<taxonomicName id="CF2402D887C2D3CFC411967D464F9509" authorityName="Erwin" authorityYear="1972" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="BF1ECCB3530C056FA472FADF624D224C" italics="true" pageId="0" pageNumber="153">Tasmanitachoides</emphasis>
</taxonomicName>
, ventral view
<emphasis id="D9DAB68E4967376C302FC434D98D81F0" bold="true" pageId="0" pageNumber="153">A</emphasis>
<taxonomicName id="286DBA43B65863D42C5E97970C1445B1" lsidName="T. baehri" pageId="0" pageNumber="153" rank="species" species="baehri">
<emphasis id="659F59D8423BEFD20AEFC8380B509AC7" italics="true" pageId="0" pageNumber="153">T. baehri</emphasis>
</taxonomicName>
, voucher DNA5569 (holotype)
<emphasis id="477F4E47CD321A67B2510B8694AA2C98" bold="true" pageId="0" pageNumber="153">B</emphasis>
<taxonomicName id="68F51BEBD913ED9F16599F040AB7AE3E" lsidName="T. wilsoni" pageId="0" pageNumber="153" rank="species" species="wilsoni">
<emphasis id="B38EC043F33AF8AC5926AF207AD5EBF0" italics="true" pageId="0" pageNumber="153">T. wilsoni</emphasis>
</taxonomicName>
, voucher DNA5514
<emphasis id="E6EFC60AC0F5A21C168CBF6AD39F7470" bold="true" pageId="0" pageNumber="153">C</emphasis>
<taxonomicName id="8A5338C78F18B13285903C6325F0B604" lsidName="T. erwini" pageId="0" pageNumber="153" rank="species" species="erwini">
<emphasis id="41070ED926612BC134DBDE0759C77D7D" italics="true" pageId="0" pageNumber="153">T. erwini</emphasis>
</taxonomicName>
, voucher V101481
<emphasis id="DE677C7AAC09504EFD52633F8413F0D4" bold="true" pageId="0" pageNumber="153">D</emphasis>
<taxonomicName id="79333986FCFD50DEE888CBCA5D76EE4C" genus="T." lsidName="T." pageId="0" pageNumber="153" rank="genus">
<emphasis id="8724A9C7B1A745B710B42F2CF073F3E5" italics="true" pageId="0" pageNumber="153">T.</emphasis>
</taxonomicName>
sp. &quot;Lerderderg R&quot;, voucher DNA2029. Scale bars: 100
<normalizedToken id="50E9AF30D2B572EC93E01D08C0A1A48E" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="6BD3BACABBE3BF438B74B7E189631D71" pageId="0" pageNumber="153" type="comparisons">
<paragraph id="D2763C341E9B2CC3F4B54000A769566B" pageId="0" pageNumber="153">Comparison with related species.</paragraph>
<paragraph id="B5230B5CE2B4571F5EB2E53289E10D17" pageId="0" pageNumber="153">
Likely to be confused only with similarly small and compact
<taxonomicName id="4A74B74391B68B54B48CCB1EC6425D1B" lsidName="T. wilsoni" pageId="0" pageNumber="153" rank="species" species="wilsoni">
<emphasis id="6C205A65C1E562F30B6F55B285FD5B9F" italics="true" pageId="0" pageNumber="153">T. wilsoni</emphasis>
</taxonomicName>
, from which it can be distinguished by the narrower pronotum with less rounded lateral margins, and narrower, less rounded elytra. In addition,
<taxonomicName id="8C1D50FD0C486E9AA3D693E2EC2CE77C" lsidName="T. wilsoni" pageId="0" pageNumber="153" rank="species" species="wilsoni">
<emphasis id="FD526B1A1A6F32C60714AB24E81322CD" italics="true" pageId="0" pageNumber="153">T. wilsoni</emphasis>
</taxonomicName>
has much shorter frontal furrows, which do not reach the anterior supraorbital seta (Fig.
<figureCitation id="EA75FFEDE728A72FFF7D875033F6D3AF" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Dorsal view of head of Tasmanitachoides adults A T. wilsoni, voucher V 101470 B T. baehri, voucher V 101479 C T. lutus, voucher V 101462 D T. cf. gerdi, voucher DNA 5676 E T. sp. &quot; Angle Crossing 1 &quot;, voucher DNA 5677 F T. obliquiceps, voucher V 101477 G T. erwini, voucher V 101469 H T. hobarti, voucher V 101463 J T. leai, voucher V 101467 K T. rufescens, voucher V 101478 L T. fitzroyi, voucher V 101471 M T. murrumbidgensis, voucher V 101464." figureDoi="10.3897/zookeys.1044.62253.figure17" httpUri="https://binary.pensoft.net/fig/555139" pageId="0" pageNumber="153">17A</figureCitation>
);
<taxonomicName id="AB2128F45444BBE4899BA1F278C7022E" lsidName="T. wilsoni" pageId="0" pageNumber="153" rank="species" species="wilsoni">
<emphasis id="B68CC7C25772603A4FEB178AFAC4D869" italics="true" pageId="0" pageNumber="153">T. wilsoni</emphasis>
</taxonomicName>
also lacks the notable groove extending forward from the anterior supraorbital seta. The elytral striae in
<taxonomicName id="335E887D0720A1F2CE19D5B9C0E657F2" lsidName="T. wilsoni" pageId="0" pageNumber="153" rank="species" species="wilsoni">
<emphasis id="4FB1C02F073047124917559EBA072330" italics="true" pageId="0" pageNumber="153">T. wilsoni</emphasis>
</taxonomicName>
are less evident than in
<taxonomicName id="2F8F665AD60CA7C1B982DBEC75455BEA" lsidName="T. baehri" pageId="0" pageNumber="153" rank="species" species="baehri">
<emphasis id="B2EB5CC723B1496060C5988F3578F2DC" italics="true" pageId="0" pageNumber="153">T. baehri</emphasis>
</taxonomicName>
:
<taxonomicName id="BD2B842F344DC1ED9DCAB56E9B4BF13B" lsidName="T. baehri" pageId="0" pageNumber="153" rank="species" species="baehri">
<emphasis id="AF4DEC10A66F396CA9DDAB6CB858F786" italics="true" pageId="0" pageNumber="153">T. baehri</emphasis>
</taxonomicName>
has an evident (if shallow and broad) stria 3 between the two anterior discal setae (Figs
<figureCitation id="6EDEABAFE83C9F345224749FCAD52098" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left elytron of Tasmanitachoides A T. kingi, voucher V 101468 B T. angulicollis, voucher DNA 5515 C T. wilsoni, voucher V 101470 D T. baehri, voucher V 101479 E T. lutus, voucher V 101462 F T. katherinei, voucher V 101475 G T. cf. gerdi, voucher DNA 5676 H T. sp. &quot; Angle Crossing 1 &quot;, voucher DNA 5677 J T. obliquiceps, voucher V 101477 K T. mandibularis, voucher V 101473." figureDoi="10.3897/zookeys.1044.62253.figure15" httpUri="https://binary.pensoft.net/fig/555137" pageId="0" pageNumber="153">15D</figureCitation>
,
<figureCitation id="C0D769C88CAC60BB0F0627F313BF5CF6" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Elytral microsculpture of Tasmanitachoides, dorsal view A T. baehri, voucher V 101484 B T. erwini, voucher V 101483. Scale bar: 100 µm." figureDoi="10.3897/zookeys.1044.62253.figure18" httpUri="https://binary.pensoft.net/fig/555140" pageId="0" pageNumber="153">18A</figureCitation>
), whereas in
<taxonomicName id="AFF0DFCD6DD0A67DCD9A0027944B3AC1" lsidName="T. wilsoni" pageId="0" pageNumber="153" rank="species" species="wilsoni">
<emphasis id="AB0ABC5589A1CC8A62231842177F196B" italics="true" pageId="0" pageNumber="153">T. wilsoni</emphasis>
</taxonomicName>
it is either absent or extremely faint and shallow (Fig.
<figureCitation id="F3404AF367C756115892A7E32E34CEF1" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left elytron of Tasmanitachoides A T. kingi, voucher V 101468 B T. angulicollis, voucher DNA 5515 C T. wilsoni, voucher V 101470 D T. baehri, voucher V 101479 E T. lutus, voucher V 101462 F T. katherinei, voucher V 101475 G T. cf. gerdi, voucher DNA 5676 H T. sp. &quot; Angle Crossing 1 &quot;, voucher DNA 5677 J T. obliquiceps, voucher V 101477 K T. mandibularis, voucher V 101473." figureDoi="10.3897/zookeys.1044.62253.figure15" httpUri="https://binary.pensoft.net/fig/555137" pageId="0" pageNumber="153">15C</figureCitation>
); stria 5 in
<taxonomicName id="10FD83AE1477FD7D0502AFD3E630330A" lsidName="T. wilsoni" pageId="0" pageNumber="153" rank="species" species="wilsoni">
<emphasis id="FC7802ECC48C3535F3295715D1255104" italics="true" pageId="0" pageNumber="153">T. wilsoni</emphasis>
</taxonomicName>
is much shorter, only reaching to around half-way in between the two anterior discal setae (Fig.
<figureCitation id="33AD621CA328156535521C4FE3F0CF3D" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left elytron of Tasmanitachoides A T. kingi, voucher V 101468 B T. angulicollis, voucher DNA 5515 C T. wilsoni, voucher V 101470 D T. baehri, voucher V 101479 E T. lutus, voucher V 101462 F T. katherinei, voucher V 101475 G T. cf. gerdi, voucher DNA 5676 H T. sp. &quot; Angle Crossing 1 &quot;, voucher DNA 5677 J T. obliquiceps, voucher V 101477 K T. mandibularis, voucher V 101473." figureDoi="10.3897/zookeys.1044.62253.figure15" httpUri="https://binary.pensoft.net/fig/555137" pageId="0" pageNumber="153">15C</figureCitation>
), as opposed to reaching or nearly reaching the second discal seta (ed5) as it does in
<taxonomicName id="81FFC324B7EB1DA0414FBA593898FAB2" lsidName="T. baehri" pageId="0" pageNumber="153" rank="species" species="baehri">
<emphasis id="2DB5C4D099EA7F7D73F36DF524D2BA80" italics="true" pageId="0" pageNumber="153">T. baehri</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="4F72A104879BE890AC8C73275FFC1223" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Left elytron of Tasmanitachoides A T. kingi, voucher V 101468 B T. angulicollis, voucher DNA 5515 C T. wilsoni, voucher V 101470 D T. baehri, voucher V 101479 E T. lutus, voucher V 101462 F T. katherinei, voucher V 101475 G T. cf. gerdi, voucher DNA 5676 H T. sp. &quot; Angle Crossing 1 &quot;, voucher DNA 5677 J T. obliquiceps, voucher V 101477 K T. mandibularis, voucher V 101473." figureDoi="10.3897/zookeys.1044.62253.figure15" httpUri="https://binary.pensoft.net/fig/555137" pageId="0" pageNumber="153">15D</figureCitation>
)
<taxonomicName id="98ED73CB46E70F8AF106315762A1B6D1" lsidName="T. baehri" pageId="0" pageNumber="153" rank="species" species="baehri">
<emphasis id="ECAD998A184C4EB770B7296F60D40F5A" italics="true" pageId="0" pageNumber="153">T. baehri</emphasis>
</taxonomicName>
and
<taxonomicName id="DCCC9354567F7646C78361438E722B51" lsidName="T. wilsoni" pageId="0" pageNumber="153" rank="species" species="wilsoni">
<emphasis id="BF13699494C93E9726725B7762BBBCA6" italics="true" pageId="0" pageNumber="153">T. wilsoni</emphasis>
</taxonomicName>
look very much like small members of the tribe
<taxonomicName id="5C8E3B75DA0A12A4A79EA28B09BDE396" authorityName="Motschulsky" authorityYear="1862" lsidName="" pageId="0" pageNumber="153" rank="tribe" tribe="Tachyini">Tachyini</taxonomicName>
(e.g.,
<taxonomicName id="4156770E774D1348F041B92637E75BC4" class="Insecta" family="Carabidae" genus="Elaphropus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Elaphropus" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="C1EDB92265E9BEDB769920186BFC81E7" italics="true" pageId="0" pageNumber="153">Elaphropus</emphasis>
</taxonomicName>
,
<taxonomicName id="928843CA5AE9A5239E84AD18198D76E5" authorityName="Motschoulsky" authorityYear="1862" class="Insecta" family="Carabidae" genus="Tachyura" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tachyura" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="4B2FB7B3794FA70F973B4790AB0415BB" italics="true" pageId="0" pageNumber="153">Tachyura</emphasis>
</taxonomicName>
). The two
<taxonomicName id="8FB1FEFBC099BDE8AA1DC4A9B09CCE67" authorityName="Erwin" authorityYear="1972" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="047045165E999C066217537DC5C2A76A" italics="true" pageId="0" pageNumber="153">Tasmanitachoides</emphasis>
</taxonomicName>
can be distinguished by the presence of four setae on the clypeus, as opposed to the two setae present in tachyines.
</paragraph>
</subSubSection>
<subSubSection id="E035AA155304AA7A4CF21EE766CA667A" pageId="0" pageNumber="153" type="distribution">
<paragraph id="7D44DECE28532F10E2E661BC8FEC3CFC" pageId="0" pageNumber="153">Geographic distribution.</paragraph>
<paragraph id="91DE48227AF528A00DA9391A0E181870" pageId="0" pageNumber="153">
Only known from the Australian Capital Territory (Fig.
<figureCitation id="1706F463BDA0E26E4151B03EE3361D6A" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Known geographic distributions of Tasmanitachoides baehri and T. erwini in southeastern Australia. Circles: T. baehri; stars: T. erwini." figureDoi="10.3897/zookeys.1044.62253.figure21" httpUri="https://binary.pensoft.net/fig/555143" pageId="0" pageNumber="153">21</figureCitation>
), but very likely occurring in similar habitats in NSW.
</paragraph>
</subSubSection>
<subSubSection id="B8F5ABEBAFBA50D44127897130570F0F" pageId="0" pageNumber="153" type="habitat">
<paragraph id="04124606CF4A9287006F1CBB3F64279F" pageId="0" pageNumber="153">Habitat.</paragraph>
<paragraph id="3008EC90AD7030EB9E5ADEF1B8A246BF" pageId="0" pageNumber="153">
Collected from pockets of gravelly cobble at the edge of still water of the Murrumbidgee River. The collection locality was amongst riverbank sheoaks (
<taxonomicName id="E313BDAF02B54E7C5615E6F75FA54D20" class="Magnoliopsida" family="Casuarinaceae" genus="Allocasuarina" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Allocasuarina" order="Fagales" pageId="0" pageNumber="153" phylum="Spermatophyta" rank="genus">
<emphasis id="9BBB96DA6AF2EA9ABA178948D0A38930" italics="true" pageId="0" pageNumber="153">Allocasuarina</emphasis>
</taxonomicName>
) and relatively protected. Specimens were recovered by splashing the gravel bank after removal of cobbles. The species was collected with
<taxonomicName id="CD9A1B2E523DF530D993AAC40BD9F174" lsidName="T. murrumbidgensis" pageId="0" pageNumber="153" rank="species" species="murrumbidgensis">
<emphasis id="7DD055D5B01FD1B81C50524D01F03C40" italics="true" pageId="0" pageNumber="153">T. murrumbidgensis</emphasis>
</taxonomicName>
,
<taxonomicName id="DE08AD899A6B2456713B4FB024AF46DF" lsidName="T. rufescens" pageId="0" pageNumber="153" rank="species" species="rufescens">
<emphasis id="6BE1B74090B3DB554D330AA251F968B4" italics="true" pageId="0" pageNumber="153">T. rufescens</emphasis>
</taxonomicName>
, and a single specimen of
<taxonomicName id="58D4F25C13225A8E408ACCE637C1552F" lsidName="T. leai" pageId="0" pageNumber="153" rank="species" species="leai">
<emphasis id="78BA9C812607749DB716F8E9A1A1F273" italics="true" pageId="0" pageNumber="153">T. leai</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="EA69CD3A54B679237458CADF36718D40" pageId="0" pageNumber="153" type="phylogenetic relationships">
<paragraph id="1435033FB98F7423A32E419DAE61E945" pageId="0" pageNumber="153">Phylogenetic relationships.</paragraph>
<paragraph id="D2B6DAAB7BBFB9FBADDEB5B939E16ED3" pageId="0" pageNumber="153">
This species belongs to the
<taxonomicName id="1151059E04AB3458A08453411F467E08" baseAuthorityName="Darlington" baseAuthorityYear="1962" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides kingi" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="kingi">
<emphasis id="A4A88C4B6943DD3A235FE4AC78DA1459" italics="true" pageId="0" pageNumber="153">Tasmanitachoides kingi</emphasis>
</taxonomicName>
species group and appears to be sister to
<taxonomicName id="13EA040C47B5F9B7DE389700F2054BB5" lsidName="T. wilsoni" pageId="0" pageNumber="153" rank="species" species="wilsoni">
<emphasis id="F5DB90DABB1D87DD63B79DC3FD703F9C" italics="true" pageId="0" pageNumber="153">T. wilsoni</emphasis>
</taxonomicName>
among the sampled species (Figs
<figureCitation id="8C43E3864EEDF59F3066F3E794F173AC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Maximum likelihood tree for the concatenated matrix of all gene fragments (main figure) and for the concatenated matrix of all genes except COI (inset; this shows only part of the tree). Branch length is shown proportional to relative divergence, as estimated by IQ-TREE; scale bar indicates 0.01 units. Outgroups (Bembidarenas, Argentinatochoides, and Andinodontis; Table 1) not depicted." figureDoi="10.3897/zookeys.1044.62253.figure5" httpUri="https://binary.pensoft.net/fig/555127" pageId="0" pageNumber="153">5</figureCitation>
-
<figureCitation id="C3866BA5311519F27602FF814A4B9CF4" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Maximum likelihood trees for wg, CAD 4, and CAD 2. Branch length is shown proportional to relative divergence, as estimated by IQ-TREE; scale bars indicate 0.01 units. Outgroups not depicted." figureDoi="10.3897/zookeys.1044.62253.figure9" httpUri="https://binary.pensoft.net/fig/555131" pageId="0" pageNumber="153">9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B06765300F59B4817F2386235E73648B" pageId="0" pageNumber="153" type="notes">
<paragraph id="69497F4A9D5B2B35B7AAFA74B6B9767B" pageId="0" pageNumber="153">Notes.</paragraph>
<paragraph id="A78B7F4C2BAEA4DD24EF6B7B8833EB98" pageId="0" pageNumber="153">
This species was called &quot;
<taxonomicName id="FA0FFBE9DE15B5178E4731D1891FC2FE" authorityName="Baehr" authorityYear="1990" class="Insecta" family="Carabidae" genus="Tasmanitachoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tasmanitachoides rufescens" order="Coleoptera" pageId="0" pageNumber="153" phylum="Arthropoda" rank="species" species="rufescens">Tasmanitachoides cf. rufescens</taxonomicName>
&quot; in
<bibRefCitation id="00CC9D4F2F95E9BBFC1F0AE755A57165" DOI="https://doi.org/10.1016/j.ympev.2018.11.006" author="Maddison, DR" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="153" pagination="151 - 176" refId="B24" refString="Maddison, DR, Kanda, K, Boyd, OF, Faille, A, Porch, N, Erwin, TL, Roig-Junent, S, 2019. Phylogeny of the beetle supertribe Trechitae (Coleoptera: Carabidae): Unexpected clades, isolated lineages, and morphological convergence. Molecular Phylogenetics and Evolution 132: 151 - 176, DOI: https://doi.org/10.1016/j.ympev.2018.11.006" title="Phylogeny of the beetle supertribe Trechitae (Coleoptera: Carabidae): Unexpected clades, isolated lineages, and morphological convergence." url="https://doi.org/10.1016/j.ympev.2018.11.006" volume="132" year="2019">Maddison et al. (2019)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>