174 lines
19 KiB
XML
174 lines
19 KiB
XML
<document id="6D715CBB7DC67234E25B6487763D792F" ID-DOI="10.5281/zenodo.200830" ID-GBIF-Dataset="722f38b4-abd7-47e6-b64d-9a7b122122dd" ID-ISSN="1175-5326" ID-Zenodo-Dep="200830" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460277456165" checkinUser="plazi" docAuthor="Rifkind, Jacques & Barr, William F." docDate="2011" docId="E93D878080288E4A75A7FB27FA97FD1C" docLanguage="en" docName="zt02821p061.pdf" docOrigin="Zootaxa 2821" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Colyphus lamed Rifkind, n. sp." docType="treatment" docVersion="8" lastPageNumber="60" masterDocId="1504FFF8802B8E4F7530FC28FFE7FFF3" masterDocTitle="New species of Colyphus Spinola (Coleoptera: Cleridae: Clerinae) from Central America, with taxonomic and distributional notes on some others" masterLastPageNumber="61" masterPageNumber="55" pageNumber="58" updateTime="1698248780870" updateUser="plazi">
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<mods:title id="E59C5D2681B70F07290622716F8A4AFE">New species of Colyphus Spinola (Coleoptera: Cleridae: Clerinae) from Central America, with taxonomic and distributional notes on some others</mods:title>
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<mods:name id="511FF06110FA359B25B1AC16BB208E78" type="personal">
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<mods:namePart id="94F70BE47410A72C86EBA5491D7050F0">Rifkind, Jacques</mods:namePart>
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<mods:namePart id="665445439328A2D00729BC03701E839F">Barr, William F.</mods:namePart>
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<mods:typeOfResource id="43B4DBE701D5D88B55DE6FAD6FC21DF7">text</mods:typeOfResource>
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<mods:title id="389F3F70BB9C5D81D87E82A8A832ED43">Zootaxa</mods:title>
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<mods:date id="5606266D9202DD31D3C259C51EE2CA34">2011</mods:date>
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<mods:detail id="6CC3EB386340E571AA8EC404A9716CBB" type="volume">
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<mods:identifier id="83A1AA87D29387EC7586AE90F1856496" type="DOI">10.5281/zenodo.200830</mods:identifier>
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<mods:identifier id="53CB7292D21E35A33B92311F979B3B67" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="022ACC23A78E7DD96B45CC5634A8EC1D" type="Zenodo-Dep">200830</mods:identifier>
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<treatment id="E93D878080288E4A75A7FB27FA97FD1C" ID-DOI="http://doi.org/10.5281/zenodo.6188306" ID-GBIF-Taxon="119396262" ID-Zenodo-Dep="6188306" LSID="urn:lsid:plazi:treatment:E93D878080288E4A75A7FB27FA97FD1C" httpUri="http://treatment.plazi.org/id/E93D878080288E4A75A7FB27FA97FD1C" lastPageId="5" lastPageNumber="60" pageId="3" pageNumber="58">
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<subSubSection id="298E651D80288E4C75A7FB27FF0CF8BF" pageId="3" pageNumber="58" type="nomenclature">
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<paragraph id="612B369680288E4C75A7FB27FDE7F8DB" blockId="3.[151,512,1807,1868]" box="[151,512,1807,1832]" pageId="3" pageNumber="58">
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<heading id="3A6381FA80288E4C75A7FB27FDE7F8DB" bold="true" box="[151,512,1807,1832]" fontSize="10" level="2" pageId="3" pageNumber="58" reason="2">
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<emphasis id="53E0EA8480288E4C75A7FB27FDE7F8DB" bold="true" box="[151,512,1807,1832]" pageId="3" pageNumber="58">
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<taxonomicName id="A6944D1580288E4C75A7FB27FE51F8DB" authority="Rifkind" authorityName="Rifkind" box="[151,438,1807,1832]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="lamed" status="sp. nov.">
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<emphasis id="53E0EA8480288E4C75A7FB27FEB6F8DB" bold="true" box="[151,337,1807,1832]" italics="true" pageId="3" pageNumber="58">Colyphus lamed</emphasis>
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Rifkind
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</taxonomicName>
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,
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<taxonomicNameLabel id="48D357FF80288E4C74F2FB38FDE7F8DB" box="[450,512,1808,1832]" pageId="3" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="612B369680288E4C75A7FB1CFF0CF8BF" blockId="3.[151,512,1807,1868]" box="[151,235,1844,1868]" pageId="3" pageNumber="58">
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(
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<figureCitation id="F9AF2A1380288E4C75AFFB1CFF05F8BF" box="[159,226,1844,1868]" captionStart="FIGURE 3" captionStartId="4.[151,250,1258,1281]" captionText="FIGURE 3. Habitus of Colyphus lamed Rifkind, n. sp. (holotype female)." pageId="3" pageNumber="58">Fig. 3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="298E651D80288E4C75A7FB54FB6DF84B" pageId="3" pageNumber="58" type="materials_examined">
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<paragraph id="612B369680288E4C75A7FB54FB6DF84B" blockId="3.[151,1436,1915,1977]" pageId="3" pageNumber="58">
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<emphasis id="53E0EA8480288E4C75A7FB54FEBEF867" bold="true" box="[151,345,1915,1940]" pageId="3" pageNumber="58">
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<typeStatus id="BE2F883480288E4C75A7FB54FF35F867" box="[151,210,1916,1940]" pageId="3" pageNumber="58">Type</typeStatus>
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specimens.
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</emphasis>
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<typeStatus id="BE2F883480288E4C7450FB54FE2FF867" box="[352,456,1916,1940]" pageId="3" pageNumber="58" type="holotype">Holotype</typeStatus>
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female:
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<collectingCountry id="1983760680288E4C771BFB54FD7CF867" box="[555,667,1916,1940]" name="Honduras" pageId="3" pageNumber="58">Honduras</collectingCountry>
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, Octopeque [sic] [should read "Ocotepeque"], El Portillo, Guisayote Bio Preserve,
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<date id="152A105680288E4C7405FB88FE33F84B" box="[309,468,1952,1976]" pageId="3" pageNumber="58" value="1995-05-21" valueMax="1995-05-22" valueMin="1995-05-21">21-22-V-1995</date>
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, R. Morris. (
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<collectionCode id="0785AE5380288E4C7753FB88FD55F84B" LSID="urn:lsid:biocol.org:col:33156" box="[611,690,1952,1976]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="3" pageNumber="58">CSCA</collectionCode>
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).
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<typeStatus id="BE2F883480288E4C77F5FB89FCC0F84A" box="[709,807,1953,1977]" pageId="3" pageNumber="58" type="paratype">Paratype</typeStatus>
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(male): same data as
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<typeStatus id="BE2F883480288E4C712AFB88FB61F84B" box="[1050,1158,1952,1976]" pageId="3" pageNumber="58" type="holotype">Holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="298E651D802F8E4B75F7FCBFFEEFFECC" pageId="4" pageNumber="59" type="diagnosis">
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<paragraph id="612B3696802F8E4B75F7FCBFFEEFFECC" blockId="4.[151,1437,151,320]" pageId="4" pageNumber="59">
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<emphasis id="53E0EA84802F8E4B75F7FCBFFEA6FF43" bold="true" box="[199,321,151,176]" pageId="4" pageNumber="59">Diagnosis.</emphasis>
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Distinguishable from its congeners on the basis of a unique combination of body shape, integumental sculpturing, setation and elytral markings.
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<taxonomicName id="A6944D15802F8E4B77A9FC94FCA8FF27" box="[665,847,188,212]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="lamed">
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<emphasis id="53E0EA84802F8E4B77A9FC94FCA8FF27" box="[665,847,188,212]" italics="true" pageId="4" pageNumber="59">Colyphus lamed</emphasis>
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</taxonomicName>
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is superficially similar to
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<taxonomicName id="A6944D15802F8E4B7145FC95FA70FF27" authority="Gorham, 1882" authorityName="Gorham" authorityYear="1882" box="[1141,1431,188,212]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="pictus">
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<emphasis id="53E0EA84802F8E4B7145FC95FB3EFF27" box="[1141,1241,189,212]" italics="true" pageId="4" pageNumber="59">C. pictus</emphasis>
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(
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<bibRefCitation id="05054B67802F8E4B71D8FC94FA68FF27" author="Gorham" box="[1256,1423,188,212]" pageId="4" pageNumber="59" refString="Gorham, H. S. (1882) Family Cleridae. In Goodman and Salvin (editors). Biologia Centrali-Americana. Insecta, Coleoptera 3 (2), 129 - 168. Taylor & Francis, London." type="journal article" year="1882">Gorham, 1882</bibRefCitation>
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)
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</taxonomicName>
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, but has different elytral markings and lacks that species' distinct band of cinereous vestiture at the elytral apices. The new species does not fit neatly within any of the species groups proposed by
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<bibRefCitation id="05054B67802F8E4B717AFD2CFB28FEEF" author="Ekis" box="[1098,1231,260,284]" pageId="4" pageNumber="59" refString="Ekis, G. (1977) Classification and evolution of the Central American beetle genus Colyphus (Cleridae). Systematic Entomology, 2, 199 - 224." type="journal article" year="1977">Ekis (1977)</bibRefCitation>
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in his revision of
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<taxonomicName id="A6944D15802F8E4B75A7FD00FEE6FEB3" box="[151,257,296,320]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="genus">
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<emphasis id="53E0EA84802F8E4B75A7FD00FEE6FEB3" box="[151,257,296,320]" italics="true" pageId="4" pageNumber="59">Colyphus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<caption id="35EB661E802F8E4B75A7F8C2FC6FFAF3" box="[151,904,1258,1281]" pageId="4" pageNumber="59">
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<paragraph id="612B3696802F8E4B75A7F8C2FC6FFAF3" blockId="4.[151,904,1258,1281]" box="[151,904,1258,1281]" pageId="4" pageNumber="59">
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<emphasis id="53E0EA84802F8E4B75A7F8C2FEF5FAF3" bold="true" box="[151,274,1258,1281]" pageId="4" pageNumber="59">FIGURE 3.</emphasis>
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Habitus of
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<taxonomicName id="A6944D15802F8E4B74B9F8C2FD63FAF3" authority="Rifkind" authorityName="Rifkind" box="[393,644,1258,1280]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="lamed" status="sp. nov.">
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<emphasis id="53E0EA84802F8E4B74B9F8C2FDCAFAF3" box="[393,557,1258,1280]" italics="true" pageId="4" pageNumber="59">Colyphus lamed</emphasis>
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Rifkind
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</taxonomicName>
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,
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<taxonomicNameLabel id="48D357FF802F8E4B77BDF8C3FD20FAF3" box="[653,711,1259,1280]" pageId="4" pageNumber="59" rank="species">
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<emphasis id="53E0EA84802F8E4B77BDF8C3FD20FAF3" bold="true" box="[653,711,1259,1280]" pageId="4" pageNumber="59">n. sp.</emphasis>
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</taxonomicNameLabel>
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(holotype female).
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</paragraph>
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</caption>
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<subSubSection id="298E651D802F8E4A75F7F905FC29FE3C" lastPageId="5" lastPageNumber="60" pageId="4" pageNumber="59" type="description">
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<paragraph id="612B3696802F8E4A75F7F905FF25FE97" blockId="4.[151,1437,1325,2033]" lastBlockId="5.[151,1436,151,607]" lastPageId="5" lastPageNumber="60" pageId="4" pageNumber="59">
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<emphasis id="53E0EA84802F8E4B75F7F905FE05FAB5" bold="true" box="[199,482,1325,1350]" pageId="4" pageNumber="59">
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Description (
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<typeStatus id="BE2F8834802F8E4B7453F905FE32FAB5" box="[355,469,1325,1350]" pageId="4" pageNumber="59" type="holotype">Holotype</typeStatus>
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).
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</emphasis>
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Length 8.6mm. Form: medium sized, elongate, dorsoventrally compressed. Color: integument black to reddish black (elytra slightly iridescent in some lights), except antennomeres 1–3 testaceous ventrally, maxillary palpi and labial palpomeres 1–2 testaceous, legs reddish brown, epipleuron dark testaceous, and each elytron with a stramineous checkmark-shaped marking as in
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<figureCitation id="F9AF2A13802F8E4B7685F9B1FC1EFA42" box="[949,1017,1433,1457]" captionStart="FIGURE 3" captionStartId="4.[151,250,1258,1281]" captionText="FIGURE 3. Habitus of Colyphus lamed Rifkind, n. sp. (holotype female)." pageId="4" pageNumber="59">Fig. 3</figureCitation>
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. Head: surface shining, finely, rather sparsely punctate; vestiture sparse, composed of mostly fine, suberect, silvery setae, with a thin admixture of longer, erect black setae; front distinctly bi-impressed; eyes moderately convex, emargination shorter than length of antennal scape; antennae just attaining base of pronotum, antennomeres 8–11 forming a rather loose, gradually expanded club, 11 elongate, inner margin subsinuate, with apex subacute. Gular process rectangulate, distinctly concave at its base. Pronotum: weakly convex, slightly broader than long, wider than head measured across eyes, narrower than base of elytra; surface shining, shallowly, indistinctly and rather minutely punctate, obscurely and transversely rugulose; vestiture consisting of a rather sparse covering of suberect, fine, silvery setae with a thin commixture of longer, erect dark setae; anterior transverse impression deeply incised across disk, broadly Ushaped at middle; lateral foveae small, anterior margin transverse, sides constricted at anterior transverse impression, convexly expanded to maximum width at middle, then arcuately narrowed to subparallel sides of rather short basal collar; posterior pronotal slope acute but shallow. Scutellum rounded apically, surface set with pale setae. Elytra: quite shallow, more than
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<date id="152A1056802F8E4B773EFB61FDCEF892" box="[526,553,1865,1889]" pageId="4" pageNumber="59">2x</date>
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as long as wide, widest just posterior to middle; surface shining, indistinctly rugulose (more obscurely so on posterior 1/2), densely but rather shallowly punctate, punctures smaller on posterior 1/2; vestiture rather inconspicuous, consisting of very fine, subrecumbent, rather short, pale setae, sparsely arranged on anterior 1/2, more densely arrayed on posterior 1/2, intermixed with fewer fine, somewhat longer, suberect black setae; disk flattened above, subbasal tumescences absent; umbones oblique; sides distinctly precipitous anteriorly, margins broadly subsinuate posterior to base, broadly, arcuately rounded from middle, converging to notably dehiscent, subacute apices; apical slope very gradual. Legs: moderately long. Posterior mesosternal process and anterior metasternal process not elevated. Abdomen: sternites shining, rather smooth, very sparsely set with scattered fine, pale setae; visible sternite 5 with sides slightly convergent posteriorly, hind angles rounded and hind margin shallowly, arcuately emarginate; visible sternite 6 with sides converging posteriorly to rounded hind margin.
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</paragraph>
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<paragraph id="612B3696802E8E4A75F7FD47FC29FE3C" blockId="5.[151,1436,151,607]" pageId="5" pageNumber="60">
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<emphasis id="53E0EA84802E8E4A75F7FD47FEDCFE7B" bold="true" box="[199,315,367,392]" pageId="5" pageNumber="60">Variation.</emphasis>
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The male
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<typeStatus id="BE2F8834802E8E4A7489FD58FDFEFE7B" box="[441,537,368,392]" pageId="5" pageNumber="60" type="paratype">paratype</typeStatus>
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is 7.8mm in length. It has the pale elytral markings reduced and broadly interrupted at the middle; its elytral surface is more uniformly and coarsely punctate. Visible abdominal sternite 6 is feebly, arcuately emarginate at the posterior margin. The tegmen is sagittate.
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</paragraph>
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</subSubSection>
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<subSubSection id="298E651D802E8E4A75F7FDF4FD74FDE4" pageId="5" pageNumber="60" type="etymology">
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<paragraph id="612B3696802E8E4A75F7FDF4FD74FDE4" blockId="5.[151,1436,151,607]" pageId="5" pageNumber="60">
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<emphasis id="53E0EA84802E8E4A75F7FDF4FEA9FE07" bold="true" box="[199,334,476,500]" pageId="5" pageNumber="60">Etymology.</emphasis>
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The specific epithet is the Hebrew letter "
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<taxonomicName id="A6944D15802E8E4A7605FDF4FC66FE07" box="[821,897,476,500]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="lamed">lamed</taxonomicName>
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," archaic cursive forms of which are similar in shape to the elytral markings of the
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<typeStatus id="BE2F8834802E8E4A7719FDD7FD68FDE4" box="[553,655,511,535]" pageId="5" pageNumber="60" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="298E651D802E8E4A75F7FE0BFA97FD1C" pageId="5" pageNumber="60" type="biology_ecology">
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<paragraph id="612B3696802E8E4A75F7FE0BFEC3FDAC" blockId="5.[151,1436,151,607]" pageId="5" pageNumber="60">
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||
<emphasis id="53E0EA84802E8E4A75F7FE0BFEC0FDCF" bold="true" box="[199,295,547,572]" pageId="5" pageNumber="60">Biology.</emphasis>
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Habitat at the Biological Reserve Guisayote
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<typeStatus id="BE2F8834802E8E4A7603FE0DFC84FDCE" box="[819,867,549,573]" pageId="5" pageNumber="60">type</typeStatus>
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locality is composed of tropical moist forest and cloud forest.
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</paragraph>
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<paragraph id="612B3696802E8E4A75A7FEA7FA97FD1C" blockId="5.[151,1436,655,751]" pageId="5" pageNumber="60">
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<emphasis id="53E0EA84802E8E4A75A7FEA7FDA5FD5B" bold="true" box="[151,578,655,680]" pageId="5" pageNumber="60">Systematic and distributional notes.</emphasis>
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We have examined the
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<typeStatus id="BE2F8834802E8E4A767BFEB8FC61FD5B" box="[843,902,656,680]" pageId="5" pageNumber="60">types</typeStatus>
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and other specimens of three species of Central American
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<taxonomicName id="A6944D15802E8E4A743DFE9CFE28FD3F" authority="Gahan" authorityName="Gahan" box="[269,463,692,716]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="genus">
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<emphasis id="53E0EA84802E8E4A743DFE9CFE98FD3F" box="[269,383,692,716]" italics="true" pageId="5" pageNumber="60">Enoclerus</emphasis>
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Gahan
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</taxonomicName>
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and determined that they more properly fit within the generic concept of
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<taxonomicName id="A6944D15802E8E4A7023FE9CFA9AFD3F" box="[1299,1405,692,716]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="53E0EA84802E8E4A7023FE9CFA9AFD3F" box="[1299,1405,692,716]" italics="true" pageId="5" pageNumber="60">Colyphus</emphasis>
|
||
</taxonomicName>
|
||
as established by
|
||
<bibRefCitation id="05054B67802E8E4A740DFEFFFE23FD1C" author="Ekis" box="[317,452,727,751]" pageId="5" pageNumber="60" refString="Ekis, G. (1977) Classification and evolution of the Central American beetle genus Colyphus (Cleridae). Systematic Entomology, 2, 199 - 224." type="journal article" year="1977">Ekis (1977)</bibRefCitation>
|
||
. Because all are rare in collections, we also give new locality data for these species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |