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<document id="A43246CA3DD3439A154A33AA2593DC54" ID-DOI="10.35463/j.apr.2024.02.06" ID-ISSN="1842-371x" ID-Zenodo-Dep="14656071" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736960438926" checkinUser="felipe" docAuthor="Iamandei, Stanila, Iamandei, Eugenia, Velitzelos, Dimitrios &amp; Velitzelos, Evangelos" docDate="2024" docId="E93687EBFFBABE08FCC8FD8FFCAFF8C8" docLanguage="en" docName="ActaPalaeontolRom.20.2.61-96.pdf" docOrigin="Acta Palaeontologica Romaniae 20 (2)" docSource="https://doi.org/10.35463/j.apr.2024.02.06" docStyle="DocumentStyle:CA474AAD92E97C533D63E0C5C0F159F1.1:ActaPalaeontolRom.2019-.journal_article" docStyleId="CA474AAD92E97C533D63E0C5C0F159F1" docStyleName="ActaPalaeontolRom.2019-.journal_article" docStyleVersion="1" docTitle="Eucaryoxylon lesbium Iamandei et Iamandei 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="85" masterDocId="150FFF93FFAFBE10FFF0FFDEFFD8FFA8" masterDocTitle="Palaeoxylotomical Studies In The Cenozoic Petrified Forests Of Greece. Part Three - Dicots" masterLastPageNumber="96" masterPageNumber="61" pageNumber="82" updateTime="1736960980745" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A0D6CF116502396A865F072184B92070">Palaeoxylotomical Studies In The Cenozoic Petrified Forests Of Greece. Part Three - Dicots</mods:title>
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<mods:namePart id="14ECD447DADF3CECC872BD58A6BB8CF0">Iamandei, Stanila</mods:namePart>
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<treatment id="E93687EBFFBABE08FCC8FD8FFCAFF8C8" ID-DOI="http://doi.org/10.5281/zenodo.14656085" ID-Zenodo-Dep="14656085" LSID="urn:lsid:plazi:treatment:E93687EBFFBABE08FCC8FD8FFCAFF8C8" httpUri="http://treatment.plazi.org/id/E93687EBFFBABE08FCC8FD8FFCAFF8C8" lastPageId="24" lastPageNumber="85" pageId="21" pageNumber="82">
<subSubSection id="29856576FFBABE05FCC8FD8FFA4CFDC1" box="[824,1428,593,617]" pageId="21" pageNumber="82" type="nomenclature">
<paragraph id="612036FDFFBABE05FCC8FD8FFA4CFDC1" blockId="21.[824,1428,593,617]" box="[824,1428,593,617]" pageId="21" pageNumber="82">
<heading id="3A688191FFBABE05FCC8FD8FFA4CFDC1" box="[824,1428,593,617]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="82" reason="5">
<emphasis id="53EBEAEFFFBABE05FCC8FD8FFA4CFDC1" bold="true" box="[824,1428,593,617]" pageId="21" pageNumber="82">
<taxonomicName id="A69F4D7EFFBABE05FCC8FD8FFAEAFDC1" authority="Iamandei et Iamandei" authorityName="Iamandei et Iamandei" authorityYear="2024" box="[824,1330,593,617]" class="Magnoliopsida" family="Juglandaceae" genus="Eucaryoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="21" pageNumber="82" phylum="Tracheophyta" rank="species" species="lesbium" status="sp. nov.">
<emphasis id="53EBEAEFFFBABE05FCC8FD8FFBF7FDC1" bold="true" box="[824,1071,593,617]" italics="true" pageId="21" pageNumber="82">Eucaryoxylon lesbium</emphasis>
Iamandei et Iamandei
</taxonomicName>
,
<taxonomicNameLabel id="48D85794FFBABE05FACEFD8FFA4CFDC1" box="[1342,1428,593,617]" pageId="21" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="29856576FFBABE05FC6CFDACFAE9FD22" box="[924,1329,626,650]" pageId="21" pageNumber="82" type="description">
<paragraph id="612036FDFFBABE05FC6CFDACFAE9FD22" blockId="21.[924,1329,626,650]" box="[924,1329,626,650]" pageId="21" pageNumber="82">
<figureCitation id="F9A42A78FFBABE05FC6CFDACFC32FD22" box="[924,1002,626,650]" captionStart="Fig" captionStartId="22.[197,236,1946,1970]" captionTargetBox="[198,1411,178,1909]" captionTargetId="figure-42@22.[198,1411,178,1909]" captionTargetPageId="22" captionText="Fig. 10. Eucaryoxylon lesbium Iamandei et Iamandei sp. nov. (Specimen Lsv1). a–c. Cross section – semi-ring porous structure, quite indistinct growth-rings, reticulate axial parenchyma; d–f. Tangential section –, solitary or short vertical chains of crystals (2-2-3), 1-3 seriate rays; g–i. Radial section – simple perforations, solitary or short vertical chains of crystals; heterocellular rays with crystalliferous marginal cells. photos a-i (graphic scale)." figureDoi="http://doi.org/10.5281/zenodo.14656103" httpUri="https://zenodo.org/record/14656103/files/figure.png" pageId="21" pageNumber="82">Fig. 10</figureCitation>
, photos a-i;
<figureCitation id="F9A42A78FFBABE05FB9DFDACFB6AFD22" box="[1133,1202,626,650]" captionStart="Fig" captionStartId="23.[197,236,1918,1942]" captionTargetBox="[203,1406,165,1882]" captionTargetId="figure-23@23.[203,1406,165,1882]" captionTargetPageId="23" captionText="Fig. 11. Eucaryoxylon lesbium Iamandei et Iamandei sp. nov. (Specimen Lsv6). a–c. Cross section – semi-ring porous structure, quite indistinct growth-rings, reticulate axial parenchyma; d–f. Tangential section –, solitary or short vertical chains of 2-3 huge crystals, 1-3 seriate rays; g–i. Radial section – simple perforations, solitary or short vertical chains of 2-3(4) huge crystals; heterocellular rays with marginal cells, sometimes, crystalliferous. photos a-i (graphic scale)." figureDoi="http://doi.org/10.5281/zenodo.14656105" httpUri="https://zenodo.org/record/14656105/files/figure.png" pageId="21" pageNumber="82">Fig.11</figureCitation>
, photos a-i.
</paragraph>
</subSubSection>
<subSubSection id="29856576FFBABE08FCC0FD6BFCAFF8C8" lastPageId="24" lastPageNumber="85" pageId="21" pageNumber="82" type="materials_examined">
<paragraph id="612036FDFFBABE05FCC0FD6BFC7AFC3B" blockId="21.[816,1437,692,2073]" pageId="21" pageNumber="82">
<emphasis id="53EBEAEFFFBABE05FCC0FD6BFC20FD65" bold="true" box="[816,1016,693,717]" pageId="21" pageNumber="82">Studied material.</emphasis>
<materialsCitation id="D1F73CA0FFBABE05FBF0FD6AFC46FC3B" collectionCode="NKUA" collectorName="They &amp; Lsv &amp; Collections &amp; Geoenviron." county="Faculty of Geol." location="Other" municipality="Lesbos Island" pageId="21" pageNumber="82" specimenCount="1">
<location id="64406026FFBABE05FBF0FD6AFBE5FD64" LSID="urn:lsid:plazi:treatment:E93687EBFFBABE08FCC8FD8FFCAFF8C8:64406026FFBABE05FBF0FD6AFBE5FD64" box="[1024,1085,692,716]" county="Faculty of Geol." municipality="Lesbos Island" name="Other" pageId="21" pageNumber="82">Other</location>
two samples of silicified wood, collected from early
<location id="64406026FFBABE05FBB6FD0BFB7BFD45" LSID="urn:lsid:plazi:treatment:E93687EBFFBABE08FCC8FD8FFCAFF8C8:64406026FFBABE05FBB6FD0BFB7BFD45" box="[1094,1187,725,749]" county="Faculty of Geol." municipality="Lesbos Island" name="Miocene" pageId="21" pageNumber="82">Miocene</location>
volcano-sedimentary deposits, of
<collectingMunicipality id="8144AC87FFBABE05FC37FD28FBBDFCA6" box="[967,1125,758,782]" pageId="21" pageNumber="82">Lesbos Island</collectingMunicipality>
showed a similar special juglandaceous xylostructure of
<taxonomicName id="A69F4D7EFFBABE05FB6DFCC6FB07FC87" box="[1181,1247,792,815]" class="Magnoliopsida" family="Juglandaceae" genus="Carya" kingdom="Plantae" order="Fagales" pageId="21" pageNumber="82" phylum="Tracheophyta" rank="genus">
<emphasis id="53EBEAEFFFBABE05FB6DFCC6FB07FC87" box="[1181,1247,792,815]" italics="true" pageId="21" pageNumber="82">Carya</emphasis>
</taxonomicName>
<typeStatus id="BE24885FFFBABE05FB00FCC6FAF9FC98" box="[1264,1313,792,816]" pageId="21" pageNumber="82">type</typeStatus>
.
<collectorName id="CC6A532BFFBABE05FAC5FCC6FAB4FC98" box="[1333,1388,792,816]" pageId="21" pageNumber="82">They</collectorName>
are registered under the field numbers:
<collectorName id="CC6A532BFFBABE05FB46FCE7FB3CFCF9" box="[1206,1252,825,849]" pageId="21" pageNumber="82">Lsv</collectorName>
1,
<collectorName id="CC6A532BFFBABE05FB0AFCE7FAFDFCF9" box="[1274,1317,825,849]" pageId="21" pageNumber="82">Lsv</collectorName>
6, and kept in the
<collectorName id="CC6A532BFFBABE05FC85FC84FC36FCDA" box="[885,1006,858,882]" pageId="21" pageNumber="82">Collections</collectorName>
of the
<collectingCounty id="88414E71FFBABE05FBCDFC84FB36FCDA" box="[1085,1262,858,882]" pageId="21" pageNumber="82">Faculty of Geol.</collectingCounty>
&amp;
<collectorName id="CC6A532BFFBABE05FAE1FC84FA41FCDA" box="[1297,1433,858,882]" pageId="21" pageNumber="82">Geoenviron.</collectorName>
, of
<collectionCode id="078EAE38FFBABE05FCBDFCA5FC46FC3B" box="[845,926,891,915]" pageId="21" pageNumber="82">NKUA</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="612036FDFFBABE05FCC0FC42FB83FB15" blockId="21.[816,1437,692,2073]" pageId="21" pageNumber="82">
<emphasis id="53EBEAEFFFBABE05FCC0FC42FB91FC1C" bold="true" box="[816,1097,924,948]" pageId="21" pageNumber="82">Microscopic description.</emphasis>
The growth rings – are present, with porous to slightly semi-ring-porous structure, showing quite distinct ring boundaries, marked by a few rows of smaller and flattened cells of ground tissue, at the outer border of the growth ring, suddenly followed by normal ground tissue cells, where larger vessels of early-wood appear. Also, in cross-section, the long tangential bands of axial parenchyma, regularly arranged, give the structure a reticulate aspect.
</paragraph>
<paragraph id="612036FDFFBABE05FCC0FB18FAECF81E" blockId="21.[816,1437,692,2073]" pageId="21" pageNumber="82">
The vessels – appear, in cross sections, as solitary pores and radial multiples of 2-3(-5), in a radial arrangement between two successive rays, defining a porous to semi-porous structure, in which the thick-walled large vessels of the early-wood are slightly diminishing in size to the late-wood. The solitary vessels are round to oval-shaped, having in the early-wood a lumina size of 70-200 / 50- 150 µm the radial / tangential diameter. In the late-wood, the vessels are smaller, of 45-60 / 30-45 μm (r/tg.d). When grouped, the vessels are slightly deformed. Their walls are thick, of 8-12 µm the double wall. In longitudinal view, the vessels show exclusively simple perforation plates and the intervessel pits are numerous, polygonal, of bordered
<typeStatus id="BE24885FFFBABE05FBDCF9ABFB84F925" box="[1068,1116,1653,1677]" pageId="21" pageNumber="82">type</typeStatus>
, small (their mean size of 5-7 µm) and have a contiguous alternate arrangement. The vessel-ray pits are quite similar and have much reduced borders, the cross-fields pits are described below. Helical thickenings in the vessel elements were not observed. The mean tangential diameter of vessel lumina is around 120 µm. Vessels' density is 5-10 vessels per square millimeter. The mean vessel element length is between 300-600 µm. Inside the vessels' lumina rare, big, and relatively thin-walled tyloses sometimes appear.
</paragraph>
<paragraph id="612036FDFFBABE05FCC0F861FC23F850" blockId="21.[816,1437,692,2073]" pageId="21" pageNumber="82">Tracheids – or vascular fibres or vasicentric tracheids were not observed.</paragraph>
<paragraph id="612036FDFFBABE08FCC0F7DFFD28FF60" blockId="21.[816,1437,692,2073]" lastBlockId="24.[151,772,176,2020]" lastPageId="24" lastPageNumber="85" pageId="21" pageNumber="82">The libriform fibres – represent the major part of the ground tissue and appear minutely pited and not septate.</paragraph>
<caption id="35E06675FFB9BE06FF35F844FA59F7BD" ID-DOI="http://doi.org/10.5281/zenodo.14656103" ID-Zenodo-Dep="14656103" httpUri="https://zenodo.org/record/14656103/files/figure.png" pageId="22" pageNumber="83" startId="22.[197,236,1946,1970]" targetBox="[198,1411,178,1909]" targetPageId="22" targetType="figure">
<paragraph id="612036FDFFB9BE06FF35F844FA59F7BD" blockId="22.[197,1411,1945,2069]" pageId="22" pageNumber="83">
<emphasis id="53EBEAEFFFB9BE06FF35F844FEC3F81A" bold="true" box="[197,283,1946,1970]" pageId="22" pageNumber="83">Fig. 10.</emphasis>
<taxonomicName id="A69F4D7EFFB9BE06FED4F847FCDFF819" authority="Iamandei et Iamandei" authorityName="Iamandei et Iamandei" authorityYear="2024" box="[292,775,1945,1969]" class="Magnoliopsida" family="Juglandaceae" genus="Eucaryoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="22" pageNumber="83" phylum="Tracheophyta" rank="species" species="lesbium" status="sp. nov.">
<emphasis id="53EBEAEFFFB9BE06FED4F847FDCCF819" box="[292,532,1945,1969]" italics="true" pageId="22" pageNumber="83">Eucaryoxylon lesbium</emphasis>
Iamandei et Iamandei
</taxonomicName>
<taxonomicNameLabel id="48D85794FFB9BE06FCE0F844FCBFF819" box="[784,871,1946,1969]" pageId="22" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
(Specimen Lsv1).
<emphasis id="53EBEAEFFFB9BE06FBC7F844FBBDF81A" bold="true" box="[1079,1125,1946,1970]" pageId="22" pageNumber="83">a–c.</emphasis>
Cross section – semi-ring porous structure, quite indistinct growth-rings, reticulate axial parenchyma;
<emphasis id="53EBEAEFFFB9BE06FC0AF865FBFEF87B" bold="true" box="[1018,1062,1979,2003]" pageId="22" pageNumber="83">d–f.</emphasis>
Tangential section –, solitary or short vertical chains of crystals (2-2-3), 1-3 seriate rays;
<emphasis id="53EBEAEFFFB9BE06FCB1F802FCB2F85C" bold="true" box="[833,874,2012,2036]" pageId="22" pageNumber="83">g–i.</emphasis>
Radial section – simple perforations, solitary or short vertical chains of crystals; heterocellular rays with crystalliferous marginal cells.
<emphasis id="53EBEAEFFFB9BE06FB94F823FB0DF7BD" bold="true" box="[1124,1237,2045,2069]" pageId="22" pageNumber="83">photos a-i</emphasis>
(graphic scale).
</paragraph>
</caption>
<caption id="35E06675FFB8BE07FF35F8A0FE3AF7B2" ID-DOI="http://doi.org/10.5281/zenodo.14656105" ID-Zenodo-Dep="14656105" httpUri="https://zenodo.org/record/14656105/files/figure.png" pageId="23" pageNumber="84" startId="23.[197,236,1918,1942]" targetBox="[203,1406,165,1882]" targetPageId="23" targetType="figure">
<paragraph id="612036FDFFB8BE07FF35F8A0FE3AF7B2" blockId="23.[197,1412,1917,2074]" pageId="23" pageNumber="84">
<emphasis id="53EBEAEFFFB8BE07FF35F8A0FEC3F83E" bold="true" box="[197,283,1918,1942]" pageId="23" pageNumber="84">Fig. 11.</emphasis>
<taxonomicName id="A69F4D7EFFB8BE07FED4F8A3FCDFF83D" authority="Iamandei et Iamandei" authorityName="Iamandei et Iamandei" authorityYear="2024" box="[292,775,1917,1941]" class="Magnoliopsida" family="Juglandaceae" genus="Eucaryoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="23" pageNumber="84" phylum="Tracheophyta" rank="species" species="lesbium" status="sp. nov.">
<emphasis id="53EBEAEFFFB8BE07FED4F8A3FDCCF83D" box="[292,532,1917,1941]" italics="true" pageId="23" pageNumber="84">Eucaryoxylon lesbium</emphasis>
Iamandei et Iamandei
</taxonomicName>
<taxonomicNameLabel id="48D85794FFB8BE07FCE0F8A0FCBFF83D" box="[784,871,1918,1941]" pageId="23" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
(Specimen Lsv6).
<emphasis id="53EBEAEFFFB8BE07FBC7F8A0FBBDF83E" bold="true" box="[1079,1125,1918,1942]" pageId="23" pageNumber="84">a–c.</emphasis>
Cross section – semi-ring porous structure, quite indistinct growth-rings, reticulate axial parenchyma;
<emphasis id="53EBEAEFFFB8BE07FC0AF841FBFEF81F" bold="true" box="[1018,1062,1951,1975]" pageId="23" pageNumber="84">d–f.</emphasis>
Tangential section –, solitary or short vertical chains of 2-3 huge crystals, 1-3 seriate rays;
<emphasis id="53EBEAEFFFB8BE07FCBAF81EFCABF870" bold="true" box="[842,883,1984,2008]" pageId="23" pageNumber="84">g–i.</emphasis>
Radial section – simple perforations, solitary or short vertical chains of 2-3(4) huge crystals; heterocellular rays with marginal cells, sometimes, crystalliferous.
<emphasis id="53EBEAEFFFB8BE07FF35F7DCFEEEF7B2" bold="true" box="[197,310,2050,2074]" pageId="23" pageNumber="84">photos a-i</emphasis>
(graphic scale).
</paragraph>
</caption>
<paragraph id="612036FDFFB7BE08FF67FF0FFD00FE5A" blockId="24.[151,772,176,2020]" pageId="24" pageNumber="85">
The axial parenchyma – is present in cross-section, paratracheal fewer, apotracheal more, as long tangential 1-3-seriate bands, quite regularly arranged giving a reticulate aspect. In longitudinal section the parenchyma can be recognized as vertical rows of rectangular cells. Sometimes some cells are chambered, hypertrophied and crystalliferous, appearing as huge solitary and/or in short chains of 2-3 enormous barel-like cells (rarely more:
<quantity id="A6679B18FFB7BE08FD29FE67FCDBFE79" box="[729,771,441,465]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="24" pageNumber="85" unit="in" value="4.0">4 in</quantity>
Lsv6), bearing a singular big polygonal crystal inside.
</paragraph>
<paragraph id="612036FDFFB7BE08FF67FE25FEFBFCF6" blockId="24.[151,772,176,2020]" pageId="24" pageNumber="85">
The rays – appear as fine rays, usually 1-3(-4) seriate. The ray height is usually low to high, sometimes of more than 30 cells. The ray density is between 8-12 rays/mm tangential. As cellular composition, in radial view, the rays are of heterocellular
<typeStatus id="BE24885FFFB7BE08FE42FD5EFE3CFD30" box="[434,484,640,664]" pageId="24" pageNumber="85">type</typeStatus>
: the body ray cells are all procumbent, having 1-2 row of upright and/or square marginal cells which have a white or dark content and, sometimes, crystalliferous. In cross-fields with vessels, small polygonal rounded pits, in 1-2 horizontal rows, often not visible due to presence of dark content, or poor preservation.
</paragraph>
<paragraph id="612036FDFFB7BE08FF67FCB9FE85FBEE" blockId="24.[151,772,176,2020]" pageId="24" pageNumber="85">Storied structures are absent. Secretory elements seem to be absent. Mineral inclusions – appear as large prismatic crystals, solitary or in short vertical chains of 2-3(-4), as a single big crystal inside an enlarged cell (or chamber of axial parenchyma cell). Also, as noted above, crystal sand appears in rays, especially in the upright and/or square marginal ray cells.</paragraph>
<paragraph id="612036FDFFB7BE08FF67FB91FDE4FAD8" blockId="24.[151,772,176,2020]" pageId="24" pageNumber="85">
<emphasis id="53EBEAEFFFB7BE08FF67FB91FE1AFBCF" bold="true" box="[151,450,1103,1127]" pageId="24" pageNumber="85">Affinities and discussions.</emphasis>
From the numerous petrified wood remains with Juglandaceous affinities, studied here,
<specimenCount id="7799FD74FFB7BE08FF67FB4CFEE1FB01" box="[151,313,1169,1193]" count="2" pageId="24" pageNumber="85" type="generic">two specimens</specimenCount>
showed a special xylostructure, showing thick-walled vessels, with exclusively simple perforations and numerous alternate bordered pits; with banded and crystalliferous parenchyma in a reticulate arrangement, vasicentric less; vertically, the axial parenchyma appear as solitary cells or short chains of 2-3 enlarged chambers, each bearing a single big crystal inside.
</paragraph>
<paragraph id="612036FDFFB7BE08FF67FAA6FEBDF931" blockId="24.[151,772,176,2020]" pageId="24" pageNumber="85">
Thus, the xylotomy of the studied specimens is comparable with that of the current
<taxonomicName id="A69F4D7EFFB7BE08FDD6FA44FD71FA19" authority="Nutt." authorityName="Nutt." box="[550,681,1434,1457]" class="Magnoliopsida" family="Juglandaceae" genus="Carya" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="53EBEAEFFFB7BE08FDD6FA44FDB1FA19" box="[550,617,1434,1457]" italics="true" pageId="24" pageNumber="85">Carya</emphasis>
Nutt.
</taxonomicName>
(known as hickory, or pecan) which has also, besides the typical juglandaceous xylotomy with thick-walled vessels, the apotracheal parenchyma as long continuous bands in cross section and, the crystalliferous parenchyma appears with big solitary crystals in some enormous „
<emphasis id="53EBEAEFFFB7BE08FD75F9E1FD2FF9FF" box="[645,759,1599,1623]" italics="true" pageId="24" pageNumber="85">barrel-like</emphasis>
“ solitary cells which, in vertical view, appear solitary or in short chains of 2-3.
</paragraph>
<paragraph id="612036FDFFB7BE08FF67F97CFCB5FF41" blockId="24.[151,772,176,2020]" lastBlockId="24.[816,1437,176,1888]" pageId="24" pageNumber="85">
The fossil correspondent of the current
<taxonomicName id="A69F4D7EFFB7BE08FD90F97DFD3EF912" authority="Nutt." authorityName="Nutt." box="[608,742,1699,1722]" class="Magnoliopsida" family="Juglandaceae" genus="Carya" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="53EBEAEFFFB7BE08FD90F97DFD7AF912" box="[608,674,1699,1722]" italics="true" pageId="24" pageNumber="85">Carya</emphasis>
Nutt.
</taxonomicName>
is
<taxonomicName id="A69F4D7EFFB7BE08FF67F91DFD05F974" authority="(Muller-Stoll et Madel) Duperon, 1988" authorityName="Duperon" authorityYear="1988" baseAuthorityName="Muller-Stoll et Madel" box="[151,733,1731,1756]" class="Magnoliopsida" family="Juglandaceae" genus="Eucaryoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="53EBEAEFFFB7BE08FF67F91DFEF3F973" box="[151,299,1731,1755]" italics="true" pageId="24" pageNumber="85">Eucaryoxylon</emphasis>
(Müller-Stoll et Mädel) Dupéron, 1988
</taxonomicName>
. Its emended diagnosis shows that „the wood- structure is porous, with solitary vessels or short multiples, with thick to very thick walls, has simple perforations, alternate intervascular pitting rather big, paratracheal and apotracheal banded parenchyma, 1-2(4)-seriate, long and rather regular; has crystalliferous parenchyma as large idioblasts, with solitary crystals in such barrel-like idioblasts, which appear isolated or in short vertical chains of 2-3 cells; has rays 1-3(5)-seriate, and septate pith“.
</paragraph>
<paragraph id="612036FDFFB7BE08FCC0FF2CFCAEFE79" blockId="24.[816,1437,176,1888]" pageId="24" pageNumber="85">
This discussion and the critical overview of the features of our studied specimens strongly suggest that we are facing to a
<taxonomicName id="A69F4D7EFFB7BE08FC31FEEAFB6CFEE4" box="[961,1204,308,332]" pageId="24" pageNumber="85">
<emphasis id="53EBEAEFFFB7BE08FC31FEEAFB8DFEE4" box="[961,1109,308,332]" italics="true" pageId="24" pageNumber="85">Eucaryoxylon</emphasis>
species
</taxonomicName>
, since the observed details are perfectly consistent with the generic diagnosis, especially by the presence of the enormous crystalliferous cells,
<emphasis id="53EBEAEFFFB7BE08FC86FE46FC30FE18" box="[886,1000,408,432]" italics="true" pageId="24" pageNumber="85">barrel-like</emphasis>
(see Dupéron, 1988, citing
<bibRefCitation id="050E4B0CFFB7BE08FAC5FE46FCB1FE79" author="Manning, W. E." pageId="24" pageNumber="85" pagination="1058 - 1087" refId="ref24468" refString="Manning, W. E., 1978. The classification within the Juglandaceae. Annals of the Missouri Botanical Garden, 65 (4): 1058 - 1087. Published by: Missouri Botanical Garden Press." type="journal article" year="1978">Manning, 1978</bibRefCitation>
).
</paragraph>
<paragraph id="612036FDFFB7BE08FCC0FE04FCB1FBEE" blockId="24.[816,1437,176,1888]" pageId="24" pageNumber="85">
Comparing the xylotomy of our studied specimens with some valid fossil species described till now, as:
<taxonomicName id="A69F4D7EFFB7BE08FCC0FDC2FAD8FD9C" authority="Duperon, 1977" authorityName="Duperon" authorityYear="1977" box="[816,1280,540,564]" class="Magnoliopsida" family="Juglandaceae" genus="Eucaryoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="85" phylum="Tracheophyta" rank="species" species="boureaui">
<emphasis id="53EBEAEFFFB7BE08FCC0FDC2FBE1FD9C" box="[816,1081,540,564]" italics="true" pageId="24" pageNumber="85">Eucaryoxylon boureaui</emphasis>
Dupéron, 1977
</taxonomicName>
;
<taxonomicName id="A69F4D7EFFB7BE08FAE9FDC3FC05FDFD" authority="Greguss, 1969" authorityName="Greguss" authorityYear="1969" class="Magnoliopsida" family="Juglandaceae" genus="Eucaryoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="85" phylum="Tracheophyta" rank="species" species="budense">
<emphasis id="53EBEAEFFFB7BE08FAE9FDC3FA44FD9C" box="[1305,1436,540,564]" italics="true" pageId="24" pageNumber="85">E. budense</emphasis>
Greguss, 1969
</taxonomicName>
;
<taxonomicName id="A69F4D7EFFB7BE08FC09FDE0FC66FDDE" authority="Muller-Stoll et Madel, 1960" authorityName="Muller-Stoll et Madel" authorityYear="1960" class="Magnoliopsida" family="Juglandaceae" genus="Eucaryoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="85" phylum="Tracheophyta" rank="species" species="crystallophorum">
<emphasis id="53EBEAEFFFB7BE08FC09FDE0FB0FFDFD" box="[1017,1239,573,597]" italics="true" pageId="24" pageNumber="85">E. crystallophorum</emphasis>
<bibRefCitation id="050E4B0CFFB7BE08FB1EFDE3FC66FDDE" author="Muller-Stoll, W. R. &amp; Madel, E." pageId="24" pageNumber="85" pagination="255 - 295" refId="ref25146" refString="Muller-Stoll, W. R. &amp; Madel, E., 1960. Juglandaceen- Holzer aus dem Tertiar des pannonischen Beckens. Senckenbergiana lethaea, 41: 255 - 295, 13 Abb., 1 Tab., 7 Tab; Frankfurt am Maine." type="book chapter" year="1960">Müller-Stoll et Mädel, 1960</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="A69F4D7EFFB7BE08FC3CFD81FA40FDDE" authority="Muller-Stoll et Madel, 1983" authorityName="Muller-Stoll et Madel" authorityYear="1983" box="[972,1432,606,630]" class="Magnoliopsida" family="Juglandaceae" genus="Eucaryoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="85" phylum="Tracheophyta" rank="species" species="guembelii">
<emphasis id="53EBEAEFFFB7BE08FC3CFD81FB8EFDDE" box="[972,1110,606,630]" italics="true" pageId="24" pageNumber="85">E. guembelii</emphasis>
Müller-Stoll et Mädel, 1983
</taxonomicName>
;
<taxonomicName id="A69F4D7EFFB7BE08FCC0FD5EFABCFD3F" authority="Muller-Stoll et Madel, 1983" authorityName="Muller-Stoll et Madel" authorityYear="1983" box="[816,1380,639,663]" class="Magnoliopsida" family="Juglandaceae" genus="Eucaryoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="85" phylum="Tracheophyta" rank="species" species="moenanum">
<emphasis id="53EBEAEFFFB7BE08FCC0FD5EFC01FD3F" box="[816,985,640,663]" italics="true" pageId="24" pageNumber="85">E. moenanum</emphasis>
Müller-Stoll et Mädel, 1983
</taxonomicName>
;
<taxonomicName id="A69F4D7EFFB7BE08FA75FD5EFAB6FD10" authority="(Watari) Muller-Stoll et Madel, 1960" authorityName="Muller-Stoll et Madel" authorityYear="1960" baseAuthorityName="Watari" class="Magnoliopsida" family="Juglandaceae" genus="Eucaryoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="85" phylum="Tracheophyta" rank="species" species="protojaponicum">
<emphasis id="53EBEAEFFFB7BE08FA75FD5EFC02FD10" italics="true" pageId="24" pageNumber="85">E. protojaponicum</emphasis>
(Watari)
<bibRefCitation id="050E4B0CFFB7BE08FBB3FD7EFAB6FD10" author="Muller-Stoll, W. R. &amp; Madel, E." box="[1091,1390,672,696]" pageId="24" pageNumber="85" pagination="255 - 295" refId="ref25146" refString="Muller-Stoll, W. R. &amp; Madel, E., 1960. Juglandaceen- Holzer aus dem Tertiar des pannonischen Beckens. Senckenbergiana lethaea, 41: 255 - 295, 13 Abb., 1 Tab., 7 Tab; Frankfurt am Maine." type="book chapter" year="1960">Müller-Stoll et Mädel, 1960</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="A69F4D7EFFB7BE08FCC0FD1DFADFFD72" authority="Iamandei et Iamandei, 2002" authorityName="Iamandei et Iamandei" authorityYear="2002" box="[816,1287,706,730]" class="Magnoliopsida" family="Juglandaceae" genus="Eucaryoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="85" phylum="Tracheophyta" rank="species" species="zarandense">
<emphasis id="53EBEAEFFFB7BE08FCC0FD1DFC13FD72" box="[816,971,706,730]" italics="true" pageId="24" pageNumber="85">E. zarandense</emphasis>
Iamandei et Iamandei, 2002
</taxonomicName>
, we observed many similar features, but not identical. Thus, the here studied specimens show some specific features as follows: semi-ring-porous structure with thick-walled vessels, solitary or in short radial multiples of 2-3(-5), with perforations exclusively simple and intervessel pitting alternate; long banded parenchyma with a reticulate arrangement, crystalliferous, as solitary or short vertical chains of 2-3(-5) enlarged chambers with big crystals; rays 1-3(-4) seriate, heterocellular, with 1-2 rows of square or upright marginal cells with crystalsand.
</paragraph>
<paragraph id="612036FDFFB7BE08FCC0FB91FC31FA38" blockId="24.[816,1437,176,1888]" pageId="24" pageNumber="85">
All these xylotomical features, described in the studied specimens, compared with previous studied species, allow us to define a new species that we name
<taxonomicName id="A69F4D7EFFB7BE08FCC0FB6CFAF2FB62" authority="Iamandei et Iamandei" authorityName="Iamandei et Iamandei" authorityYear="2024" box="[816,1322,1202,1226]" class="Magnoliopsida" family="Juglandaceae" genus="Eucaryoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="85" phylum="Tracheophyta" rank="species" species="lesbium" status="sp. nov.">
<emphasis id="53EBEAEFFFB7BE08FCC0FB6CFBFDFB62" box="[816,1061,1202,1226]" italics="true" pageId="24" pageNumber="85">Eucaryoxylon lesbium</emphasis>
Iamandei et Iamandei
</taxonomicName>
,
<taxonomicNameLabel id="48D85794FFB7BE08FACBFB6DFA40FB62" box="[1339,1432,1203,1226]" pageId="24" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
, after the name of the provenance place (Lesbos Island). It could be a possible ancestor of some disappeared
<typeStatus id="BE24885FFFB7BE08FAB7FB2BFAA7FAA5" box="[1351,1407,1269,1293]" pageId="24" pageNumber="85">types</typeStatus>
of
<taxonomicName id="A69F4D7EFFB7BE08FCC0FAC8FCABFA85" box="[816,883,1302,1325]" class="Magnoliopsida" family="Juglandaceae" genus="Carya" kingdom="Plantae" order="Fagales" pageId="24" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="53EBEAEFFFB7BE08FCC0FAC8FCABFA85" box="[816,883,1302,1325]" italics="true" pageId="24" pageNumber="85">Carya</emphasis>
</taxonomicName>
, that lived, during the Cenozoic time in Europe, as shown by the identifications of Dupéron (1977), Greguss (1969),
<bibRefCitation id="050E4B0CFFB7BE08FC77FA86FB18FAD8" author="Muller-Stoll, W. R. &amp; Madel, E." box="[903,1216,1368,1392]" pageId="24" pageNumber="85" pagination="255 - 295" refId="ref25146" refString="Muller-Stoll, W. R. &amp; Madel, E., 1960. Juglandaceen- Holzer aus dem Tertiar des pannonischen Beckens. Senckenbergiana lethaea, 41: 255 - 295, 13 Abb., 1 Tab., 7 Tab; Frankfurt am Maine." type="book chapter" year="1960">Müller-Stoll &amp; Mädel (1960</bibRefCitation>
, 1983), Iamandei &amp; Iamandei (2002).
</paragraph>
<paragraph id="612036FDFFB7BE08FCC0FA44FCAFF8C8" blockId="24.[816,1437,176,1888]" pageId="24" pageNumber="85">
So, we designate the specimen Lsv1 as
<typeStatus id="BE24885FFFB7BE08FB11FA44FAE6FA1A" box="[1249,1342,1434,1458]" pageId="24" pageNumber="85" type="holotype">holotype</typeStatus>
and, the specimen Lsv6 as
<typeStatus id="BE24885FFFB7BE08FBF0FA62FB86FA7C" box="[1024,1118,1468,1492]" pageId="24" pageNumber="85" type="paratype">paratype</typeStatus>
, and this is the diagnosis of the new species: “Growth rings present, with porous to semi-ring-porous structure and with long tangential bands of axial parenchyma giving a reticulate aspect; vessels are solitary or in short radial multiples of 2-3(-5), thick-walled, circular to oval, 50-150 µm tg.d., 5-10(-20) vessels on sq. mm; exclusively simple perforations plates, alternate intervascular pitting; parenchyma banded reticulate, with solitary or vertical chains of 2-3 enormous crystals; rays 1-3(-4) seriate, heterocellular, with 1-2 rows of square or upright marginal cells sometimes crystalliferous; cross field small, few simple pits, in 1-2 rows”.
</paragraph>
</subSubSection>
</treatment>
</document>
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