treatments-xml/data/E9/2E/87/E92E878E522DFFE8D8D16540FD79FDD5.xml

591 lines
69 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="7C27D2369C92D6470D4745BF576FBEBF" ID-DOI="10.11646/zootaxa.5536.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14239180" ID-ZooBank="A63FF362-AFF3-434C-843E-46A9F1EF55DB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732799396256" checkinUser="plazi" docAuthor="Ito, Tomiko, Kuhara, Naotoshi &amp; Malicky, Hans" docDate="2024" docId="E92E878E522DFFE8D8D16540FD79FDD5" docLanguage="en" docName="zootaxa.5536.4.3.pdf" docOrigin="Zootaxa 5536 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacophila misago Ito &amp; Kuhara &amp; Malicky 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="565" masterDocId="1517FFF65227FFE6D8466056FFE6FFCC" masterDocTitle="Five new species of the genus Rhyacophila Stephens (Trichoptera, Rhyacophilidae) from southwestern Hokkaido, northern Japan" masterLastPageNumber="568" masterPageNumber="551" pageNumber="561" updateTime="1732839613807" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2C89CDDAA4D4F3FCAB519ED3D87EC1A8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="07279109727BC584D04C2DBCD8302256">
<mods:title id="F79E12BF4C3AB7DCB75C1B9E5DDA3373">Five new species of the genus Rhyacophila Stephens (Trichoptera, Rhyacophilidae) from southwestern Hokkaido, northern Japan</mods:title>
</mods:titleInfo>
<mods:name id="3A924329C5697D5C05AB5AE5DBE7EA39" type="personal">
<mods:role id="540835D2AF0C107D0B0F1CC8E3106398">
<mods:roleTerm id="B6CBAB22C5F90BBFEF7B1688F846769B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FE7462BC09E2043513D2CE813341013B">Ito, Tomiko</mods:namePart>
<mods:affiliation id="999B87849F50EDAA6CED448165C84A31">Hokkaido Aquatic Biology, Hakuyo-chô, 3 - 3 - 5, Eniwa, Hokkaido, 061 - 1434 Japan</mods:affiliation>
</mods:name>
<mods:name id="064F15509CB7D05FE7DE8C0DAE1EE761" type="personal">
<mods:role id="4EE44F9A293D41CCF6CF5A7C813415F4">
<mods:roleTerm id="499E3FB3E1F323003E021E1B6B7E5652">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="39DFB846906D206FF8CCEEAE01906B6B">Kuhara, Naotoshi</mods:namePart>
<mods:affiliation id="D5D68B37DC1D504A16A18B1ABBE2921A">Kasuga-chô 4 - 2 - 15 - 107, Chitose, Hokkaido 066 - 0065, Japan</mods:affiliation>
</mods:name>
<mods:name id="5FD4761BC5CF15C94071A2FFC36AE291" type="personal">
<mods:role id="76C1C8EC6B2A20E8EF02DE8006D5530F">
<mods:roleTerm id="7D61F1A5C618FB8F7AFC71A84A98C04F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC5DE6E7DA73162297D88BFD4833B579">Malicky, Hans</mods:namePart>
<mods:affiliation id="2F3CE2E80A67AEE9FAE7A04A5B456A84">Sonnengasse 13, A- 3293 Lunz am See, Austria</mods:affiliation>
</mods:name>
<mods:typeOfResource id="00C5FF36185040DC081553AFAD559545">text</mods:typeOfResource>
<mods:relatedItem id="177C0915B3195FB33E3ADE744B7736A8" type="host">
<mods:titleInfo id="371F558E4857085AC4FCCC40132282CC">
<mods:title id="286B736A3DB07612EEA6549CEDED5FFC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8C8758ADA13B7DE9B87BF7FA65FEBDB8">
<mods:date id="7DD31055D6EF4B60545AA89037633258">2024</mods:date>
<mods:detail id="A407FC6DC7561A508EC889D324A4E2F0" type="pubDate">
<mods:number id="6DD677BA9A0D336F74A83C5AFF3DB680">2024-11-06</mods:number>
</mods:detail>
<mods:detail id="768A018B14CF0217F442BF6C86DF3A9E" type="volume">
<mods:number id="BA45257A4E119314555F2C967C936D18">5536</mods:number>
</mods:detail>
<mods:detail id="1FDD0024806C2DA84C92F4FB092CD34C" type="issue">
<mods:number id="0B839928725701141DFAD33E60A1E527">4</mods:number>
</mods:detail>
<mods:extent id="21AA49E91400123EBC19583A5E3AD710" unit="page">
<mods:start id="1DDF0FF3AB2DAE5B0F87D3EAAFA03B2E">551</mods:start>
<mods:end id="A57DF8FC2385A8A89C995F370752C7CE">568</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="4A80CDC7DFA292908C52F681B8FACDD6">
<mods:url id="86CEC08FB5D3EEE4B624E82814A58F8D">http://dx.doi.org/10.11646/zootaxa.5536.4.3</mods:url>
</mods:location>
<mods:classification id="54CB2A9B996D19F0A85F5564CFEBB52A">journal article</mods:classification>
<mods:identifier id="2E07542ED002418CD556272BCE7591E2" type="DOI">10.11646/zootaxa.5536.4.3</mods:identifier>
<mods:identifier id="A552D8FA77FE518BFB9AA405A57FC974" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="859A7ABD94652A1597CAC6F8482F2B6D" type="Zenodo-Dep">14239180</mods:identifier>
<mods:identifier id="5B461ABCC7B6CECEAED792F964F9469C" type="ZooBank">A63FF362-AFF3-434C-843E-46A9F1EF55DB</mods:identifier>
</mods:mods>
<treatment id="E92E878E522DFFE8D8D16540FD79FDD5" ID-DOI="http://doi.org/10.5281/zenodo.14239689" ID-Zenodo-Dep="14239689" LSID="urn:lsid:plazi:treatment:E92E878E522DFFE8D8D16540FD79FDD5" httpUri="http://treatment.plazi.org/id/E92E878E522DFFE8D8D16540FD79FDD5" lastPageId="14" lastPageNumber="565" pageId="10" pageNumber="561">
<subSubSection id="299D6513522DFFECD8D16540FD41FAFD" box="[151,679,1302,1329]" pageId="10" pageNumber="561" type="nomenclature">
<paragraph id="61383698522DFFECD8D16540FD41FAFD" blockId="10.[151,679,1302,1365]" box="[151,679,1302,1329]" pageId="10" pageNumber="561">
<heading id="3A7081F4522DFFECD8D16540FD41FAFD" bold="true" box="[151,679,1302,1329]" fontSize="11" level="1" pageId="10" pageNumber="561" reason="1">
<emphasis id="53F3EA8A522DFFECD8D16540FD41FAFD" bold="true" box="[151,679,1302,1329]" pageId="10" pageNumber="561">
<taxonomicName id="A6874D1B522DFFECD8D16540FDA4FAFD" authority="Ito &amp; Kuhara" authorityName="Ito &amp; Kuhara &amp; Malicky" authorityYear="2024" box="[151,578,1302,1329]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="561" phylum="Arthropoda" rank="species" species="misago" status="sp. nov.">
<emphasis id="53F3EA8A522DFFECD8D16540FE68FAFD" bold="true" box="[151,398,1302,1329]" italics="true" pageId="10" pageNumber="561">Rhyacophila misago</emphasis>
Ito &amp; Kuhara
</taxonomicName>
<taxonomicNameLabel id="48C057F1522DFFECDA0C6541FD41FAFD" box="[586,679,1303,1329]" pageId="10" pageNumber="561" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="299D6513522DFFECD8D1656DFC70FA5B" pageId="10" pageNumber="561" type="description">
<paragraph id="61383698522DFFECD8D1656DFE5FFA99" blockId="10.[151,679,1302,1365]" box="[151,441,1338,1365]" pageId="10" pageNumber="561">
(
<figureCitation id="F9BC2A1D522DFFECD8D9656DFF1DFA99" box="[159,251,1339,1365]" captionStart="FIGURE 1" captionStartId="2.[151,250,1655,1679]" captionTargetBox="[165,1422,186,1625]" captionTargetId="figure-29@2.[151,1436,179,1630]" captionTargetPageId="2" captionText="FIGURE 1. Recorded collection sites of Rhyacophila species. 1A, location of Hokkaido and recorded sites of known species discussed in present study. 1B1E, recorded and collected sites in Hokkaido: 1B, 1C, R. eniwa sp. nov. and R. arefini Lukyanchenko; 1D, R. bifue sp. nov., R. imakane sp. nov., R. misago sp. nov., and R. hamata Hattori &amp; Ohkawa; 1E, R. shimamaki sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14239182" httpUri="https://zenodo.org/record/14239182/files/figure.png" pageId="10" pageNumber="561">Figs 1D</figureCitation>
,
<figureCitation id="F9BC2A1D522DFFECD94E656CFEB4FA99" box="[264,338,1338,1365]" captionStart="FIGURE 5" captionStartId="11.[151,250,1787,1811]" captionTargetBox="[254,1333,184,1758]" captionTargetId="figure-20@11.[241,1346,179,1761]" captionTargetPageId="11" captionText="FIGURE 5. Adult of Rhyacophila misago sp. nov. 5A5D, male: 5A, genitalia, left lateral, omitting left inferior appendage; 5B, same, dorsal; 5C, right inferior appendage, dorsal; 5D, right inferior appendage and phallic apparatus, ventral. 5E5I, female: 5E, abdominal segments VIIIXI, left lateral; 5F, segment VIII, dorsal; 5G, same, ventral; 5H, vaginal apparatus, left lateral; 5I, same, ventral.Abbreviations:VIIIXI = abdominal segments VIIIXI; a inf app = apical segment of inferior appendage (paired); ae = aedeagus; an sc = anal sclerite (paired); ap ba = apical band (paired); b inf app = basal segment of inferior appendage (paired); do me pr = dorsomesal process (paired); par = paramere (paired); pp = posterior process; ps = processus spermathecae; ru po IX = rugged posterior part of segment IX; sl cu pr = slender curved process (paired); te st = tergal strap (paired)." figureDoi="http://doi.org/10.5281/zenodo.14239190" httpUri="https://zenodo.org/record/14239190/files/figure.png" pageId="10" pageNumber="561">5A5I</figureCitation>
,
<figureCitation id="F9BC2A1D522DFFECD918656DFE54FA99" box="[350,434,1339,1365]" captionStart="FIGURE 6" captionStartId="12.[151,250,1710,1734]" captionTargetBox="[158,1426,191,1678]" captionTargetId="figure-29@12.[151,1436,181,1686]" captionTargetPageId="12" captionText="FIGURE 6. Larva of Rhyacophila misago sp. nov. 6A, habitus, left lateral; 6B, head, dorsal; 6C, mandibles, ventral; 6D, pronotum, dorsal; 6E, pleuron and right foreleg, right lateral; 6F, pleuron and right hind leg, right lateral; 6G, left anal proleg, left lateral." figureDoi="http://doi.org/10.5281/zenodo.14239192" httpUri="https://zenodo.org/record/14239192/files/figure.png" pageId="10" pageNumber="561">6A6G</figureCitation>
)
</paragraph>
<paragraph id="61383698522DFFECD8D16529FC70FA5B" blockId="10.[151,918,1407,1431]" box="[151,918,1407,1431]" pageId="10" pageNumber="561">
<taxonomicName id="A6874D1B522DFFECD8D16529FEB6FA5B" box="[151,336,1407,1431]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="561" phylum="Arthropoda" rank="species" species="undefined-3">
<emphasis id="53F3EA8A522DFFECD8D16529FEF1FA5B" box="[151,279,1407,1431]" italics="true" pageId="10" pageNumber="561">Rhyacophila</emphasis>
sp. 3
</taxonomicName>
:
<bibRefCitation id="05164B69522DFFECD91D65D6FE0DFA5B" author="Ito, T. &amp; Kuhara, N. &amp; Hattori, T. &amp; Ohkawa, A." box="[347,491,1407,1431]" pageId="10" pageNumber="561" pagination="51 - 85" refId="ref12429" refString="Ito, T., Kuhara, N., Hattori, T. &amp; Ohkawa, A. (2010) Caddisfly (Trichoptera) fauna of Oshima Peninsula, Hokkaido, northern Japan. Biology of Inland Waters, 25, 51 - 85. [in Japanese with English abstract]" type="journal article" year="2010">
Ito
<emphasis id="53F3EA8A522DFFECD93B65D6FE4CFA5B" box="[381,426,1407,1431]" italics="true" pageId="10" pageNumber="561">et al</emphasis>
. 2010
</bibRefCitation>
, pp. 56, 81,
<collectingRegion id="A343F87A522DFFECDA2E6529FD2AFA5B" box="[616,716,1407,1431]" country="Japan" name="Hokkaido" pageId="10" pageNumber="561">Hokkaido</collectingRegion>
(Hiyama, Oshima).
</paragraph>
</subSubSection>
<subSubSection id="299D6513522DFFEBD8D16593FD6AFE61" lastPageId="13" lastPageNumber="564" pageId="10" pageNumber="561" type="diagnosis">
<paragraph id="61383698522DFFECD8D16593FDBBF88B" blockId="10.[151,1437,1477,1972]" pageId="10" pageNumber="561">
<emphasis id="53F3EA8A522DFFECD8D16593FE00FA13" bold="true" box="[151,486,1477,1503]" pageId="10" pageNumber="561">Species group and diagnosis</emphasis>
. This new species belongs to the
<taxonomicName id="A6874D1B522DFFECDB266593FBB1FA13" authorityName="Levanidova &amp; Schmid" authorityYear="1977" box="[864,1111,1477,1503]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="561" phylum="Arthropoda" rank="species" species="kaltatica">
<emphasis id="53F3EA8A522DFFECDB266593FBB1FA13" box="[864,1111,1477,1503]" italics="true" pageId="10" pageNumber="561">Rhyacophila kaltatica</emphasis>
</taxonomicName>
Species Group in having the dorsal posterior margin of segment IX protruded caudad with several spiny processes, a long depressed segment X, large anal sclerite and thick inferior appendages with long basal and short apical segments. The male of this new species resembles that of
<taxonomicName id="A6874D1B522DFFECD9FF6664FCFBF987" authority="Kobayashi 1973" authorityName="Kobayashi" authorityYear="1973" box="[441,797,1585,1611]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="561" phylum="Arthropoda" rank="species" species="azumaensis">
<emphasis id="53F3EA8A522DFFECD9FF6664FDBCF987" box="[441,602,1586,1611]" italics="true" pageId="10" pageNumber="561">R. azumaensis</emphasis>
<bibRefCitation id="05164B69522DFFECDA246667FCFBF987" author="Kobayashi, M." box="[610,797,1585,1611]" pageId="10" pageNumber="561" pagination="21 - 44" refId="ref12691" refString="Kobayashi, M. (1973) Caddisfly fauna of the vicinity of Yamagata Prefecture, with descriptions of thirteen new species. Bulletin of the Kanagawa Prefectural Museum (Natural Science), 6, 21 - 44, 10 pls." type="journal article" year="1973">Kobayashi 1973</bibRefCitation>
</taxonomicName>
, found in
<collectingCountry id="19907608522DFFECDBD66664FC36F980" box="[912,976,1586,1612]" name="Japan" pageId="10" pageNumber="561">Japan</collectingCountry>
(northern Honshu;
<figureCitation id="F9BC2A1D522DFFECDCE96667FAE1F987" box="[1199,1287,1585,1611]" captionStart="FIGURE 1" captionStartId="2.[151,250,1655,1679]" captionTargetBox="[165,1422,186,1625]" captionTargetId="figure-29@2.[151,1436,179,1630]" captionTargetPageId="2" captionText="FIGURE 1. Recorded collection sites of Rhyacophila species. 1A, location of Hokkaido and recorded sites of known species discussed in present study. 1B1E, recorded and collected sites in Hokkaido: 1B, 1C, R. eniwa sp. nov. and R. arefini Lukyanchenko; 1D, R. bifue sp. nov., R. imakane sp. nov., R. misago sp. nov., and R. hamata Hattori &amp; Ohkawa; 1E, R. shimamaki sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14239182" httpUri="https://zenodo.org/record/14239182/files/figure.png" pageId="10" pageNumber="561">Fig. 1A</figureCitation>
, black stars), in having pair of slender lateral processes of the rugged posterior area of segment IX and a pair of parameres. However, the male of
<taxonomicName id="A6874D1B522DFFECD9D6662CFDE7F95F" authorityName="Ito &amp; Kuhara &amp; Malicky" authorityYear="2024" box="[400,513,1658,1683]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="561" phylum="Arthropoda" rank="species" species="misago" status="sp. nov.">
<emphasis id="53F3EA8A522DFFECD9D6662CFDE7F95F" box="[400,513,1658,1683]" italics="true" pageId="10" pageNumber="561">R. misago</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48C057F1522DFFECDA4E662FFD85F958" box="[520,611,1657,1684]" pageId="10" pageNumber="561" rank="species">
<emphasis id="53F3EA8A522DFFECDA4E662FFDC4F95F" bold="true" box="[520,546,1657,1683]" pageId="10" pageNumber="561">sp</emphasis>
.
<emphasis id="53F3EA8A522DFFECDA76662FFDBAF95F" bold="true" box="[560,604,1657,1683]" pageId="10" pageNumber="561">nov</emphasis>
.
</taxonomicNameLabel>
is clearly distinguishable from that of
<taxonomicName id="A6874D1B522DFFECDC53662CFB50F95F" authorityName="Kobayashi" authorityYear="1973" box="[1045,1206,1658,1683]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="561" phylum="Arthropoda" rank="species" species="azumaensis">
<emphasis id="53F3EA8A522DFFECDC53662CFB50F95F" box="[1045,1206,1658,1683]" italics="true" pageId="10" pageNumber="561">R. azumaensis</emphasis>
</taxonomicName>
as follows: The pair of long processes of the rugged posterior part of segment IX make a circle in dorsal view in
<taxonomicName id="A6874D1B522DFFECDC8A66C8FAD9F97B" authorityName="Ito &amp; Kuhara &amp; Malicky" authorityYear="2024" box="[1228,1343,1694,1719]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="561" phylum="Arthropoda" rank="species" species="misago">
<emphasis id="53F3EA8A522DFFECDC8A66C8FAD9F97B" box="[1228,1343,1694,1719]" italics="true" pageId="10" pageNumber="561">R. misago</emphasis>
</taxonomicName>
and the dorsomesal process of the basal segment of each inferior appendage is arising from the subapicodorsal edge, short, and strongly curved apicodorsad; but the pair of slender processes is much shorter and gently curved caudad in dorsal view in
<taxonomicName id="A6874D1B522DFFECD979675CFE06F8EF" authorityName="Kobayashi" authorityYear="1973" box="[319,480,1802,1827]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="561" phylum="Arthropoda" rank="species" species="azumaensis">
<emphasis id="53F3EA8A522DFFECD979675CFE06F8EF" box="[319,480,1802,1827]" italics="true" pageId="10" pageNumber="561">R. azumaensis</emphasis>
</taxonomicName>
and the dorsomesal process of the basal segment of each inferior appendage is long, arising at midlength, and nearly straight.
</paragraph>
<paragraph id="61383698522DFFECD8816707FDB4F878" blockId="10.[151,1437,1477,1972]" pageId="10" pageNumber="561">
The female of this new species is also similar to that of
<taxonomicName id="A6874D1B522DFFECDB6F6704FC2EF8A7" authorityName="Kobayashi" authorityYear="1973" box="[809,968,1874,1899]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="561" phylum="Arthropoda" rank="species" species="azumaensis">
<emphasis id="53F3EA8A522DFFECDB6F6704FC2EF8A7" box="[809,968,1874,1899]" italics="true" pageId="10" pageNumber="561">R. azumaensis</emphasis>
</taxonomicName>
, but can be readily discriminated from the latter as follows: In lateral view, segment VIII has a pair of small curved ridges ventroposteriorly in
<taxonomicName id="A6874D1B522DFFECDCB36720FA73F843" authorityName="Kobayashi" authorityYear="1973" box="[1269,1429,1910,1935]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="561" phylum="Arthropoda" rank="species" species="azumaensis">
<emphasis id="53F3EA8A522DFFECDCB36720FA73F843" box="[1269,1429,1910,1935]" italics="true" pageId="10" pageNumber="561">R. azumaensis</emphasis>
</taxonomicName>
, but is without such ridges in
<taxonomicName id="A6874D1B522DFFECD99D67CCFDADF87F" authorityName="Ito &amp; Kuhara &amp; Malicky" authorityYear="2024" box="[475,587,1946,1971]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="561" phylum="Arthropoda" rank="species" species="misago">
<emphasis id="53F3EA8A522DFFECD99D67CCFDADF87F" box="[475,587,1946,1971]" italics="true" pageId="10" pageNumber="561">R. misago</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="35F86610522CFFEDD8D166ADFAA2F827" ID-DOI="http://doi.org/10.5281/zenodo.14239190" ID-Zenodo-Dep="14239190" httpUri="https://zenodo.org/record/14239190/files/figure.png" pageId="11" pageNumber="562" startId="11.[151,250,1787,1811]" targetBox="[254,1333,184,1758]" targetPageId="11" targetType="figure">
<paragraph id="61383698522CFFEDD8D166ADFAA2F827" blockId="11.[151,1437,1787,2028]" pageId="11" pageNumber="562">
<emphasis id="53F3EA8A522CFFEDD8D166ADFEEAF8DF" bold="true" box="[151,268,1787,1811]" pageId="11" pageNumber="562">FIGURE 5</emphasis>
. Adult of
<taxonomicName id="A6874D1B522CFFEDD92966ADFDDBF8DF" authorityName="Ito &amp; Kuhara &amp; Malicky" authorityYear="2024" box="[367,573,1787,1811]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="562" phylum="Arthropoda" rank="species" species="misago" status="sp. nov.">
<emphasis id="53F3EA8A522CFFEDD92966ADFDDBF8DF" box="[367,573,1787,1811]" italics="true" pageId="11" pageNumber="562">Rhyacophila misago</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48C057F1522CFFEDDA0566ADFD72F8DF" box="[579,660,1787,1811]" pageId="11" pageNumber="562" rank="species">
<emphasis id="53F3EA8A522CFFEDDA0566ADFD6BF8DF" bold="true" box="[579,653,1787,1811]" pageId="11" pageNumber="562">sp. nov</emphasis>
.
</taxonomicNameLabel>
5A5D, male: 5A, genitalia, left lateral, omitting left inferior appendage; 5B, same, dorsal; 5C, right inferior appendage, dorsal; 5D, right inferior appendage and phallic apparatus, ventral. 5E5I, female: 5E, abdominal segments VIIIXI, left lateral; 5F, segment VIII, dorsal; 5G, same, ventral; 5H, vaginal apparatus, left lateral; 5I, same, ventral. Abbreviations: VIIIXI = abdominal segments VIIIXI; a inf app = apical segment of inferior appendage (paired); ae = aedeagus; an sc = anal sclerite (paired); ap ba = apical band (paired); b inf app = basal segment of inferior appendage (paired); do me pr = dorsomesal process (paired); par = paramere (paired); pp = posterior process; ps = processus spermathecae; ru po IX = rugged posterior part of segment IX; sl cu pr = slender curved process (paired); te st = tergal strap (paired).
</paragraph>
</caption>
<caption id="35F86610522BFFEAD8D166F8FEE0F8C2" ID-DOI="http://doi.org/10.5281/zenodo.14239192" ID-Zenodo-Dep="14239192" httpUri="https://zenodo.org/record/14239192/files/figure.png" pageId="12" pageNumber="563" startId="12.[151,250,1710,1734]" targetBox="[158,1426,191,1678]" targetPageId="12" targetType="figure">
<paragraph id="61383698522BFFEAD8D166F8FEE0F8C2" blockId="12.[151,1436,1710,1806]" pageId="12" pageNumber="563">
<emphasis id="53F3EA8A522BFFEAD8D166F8FEF6F90A" bold="true" box="[151,272,1710,1734]" pageId="12" pageNumber="563">FIGURE 6</emphasis>
. Larva of
<taxonomicName id="A6874D1B522BFFEAD9C466F8FDB3F90A" authorityName="Ito &amp; Kuhara &amp; Malicky" authorityYear="2024" box="[386,597,1710,1734]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="563" phylum="Arthropoda" rank="species" species="misago" status="sp. nov.">
<emphasis id="53F3EA8A522BFFEAD9C466F8FDB3F90A" box="[386,597,1710,1734]" italics="true" pageId="12" pageNumber="563">Rhyacophila misago</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48C057F1522BFFEADA1866F8FD52F909" box="[606,692,1710,1734]" pageId="12" pageNumber="563" rank="species">
<emphasis id="53F3EA8A522BFFEADA1866F8FD4BF90A" bold="true" box="[606,685,1710,1734]" pageId="12" pageNumber="563">sp. nov</emphasis>
.
</taxonomicNameLabel>
6A, habitus, left lateral; 6B, head, dorsal; 6C, mandibles, ventral; 6D, pronotum, dorsal; 6E, pleuron and right foreleg, right lateral; 6F, pleuron and right hind leg, right lateral; 6G, left anal proleg, left lateral.
</paragraph>
</caption>
<paragraph id="61383698522BFFEAD8816736FE0DF80F" blockId="12.[151,1436,1888,1987]" pageId="12" pageNumber="563">
The larva of this species is similar to that of
<taxonomicName id="A6874D1B522BFFEADAF36737FBD9F8B6" authority="Hattori &amp; Ohkawa 2007" authorityName="Hattori &amp; Ohkawa" authorityYear="2007" box="[693,1087,1888,1914]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="563" phylum="Arthropoda" rank="species" species="hamata">
<emphasis id="53F3EA8A522BFFEADAF36737FCC1F8B6" box="[693,807,1888,1914]" italics="true" pageId="12" pageNumber="563">R. hamata</emphasis>
Hattori &amp;
<bibRefCitation id="05164B69522BFFEADBE66736FBD9F8B6" author="Ohkawa, A. &amp; Hattori, T." box="[928,1087,1888,1914]" pageId="12" pageNumber="563" pagination="263 - 274" refId="ref13374" refString="Ohkawa, A. &amp; Hattori, T. (2007) The caddisflies (Trichoptera) of the University Forest in Hokkaido, the University of Tokyo. I. Rhyacophilidae. In: Bueno-Soria, J., Barba-Alvarez, R. &amp; Armitage, B. (Eds.), Proceedings of the 12 th International Symposium on Trichoptera. The Caddis Press, Columbus, Ohio, pp. 263 - 274." type="book chapter" year="2007">Ohkawa 2007</bibRefCitation>
</taxonomicName>
, but different from the latter by the presence of a rectangular, light color pattern on the frontoclypeal apotome of the head, whereas a rectangular pattern is absent in
<taxonomicName id="A6874D1B522BFFEAD93767FFFE02F80E" box="[369,484,1960,1986]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="563" phylum="Arthropoda" rank="species" species="hamata">
<emphasis id="53F3EA8A522BFFEAD93767FFFE02F80E" box="[369,484,1960,1986]" italics="true" pageId="12" pageNumber="563">R. hamata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="61383698522AFFEBD88160C0FD6AFE61" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="564">
<emphasis id="53F3EA8A522AFFEBD88160C0FEEDFF7C" bold="true" box="[199,267,150,176]" pageId="13" pageNumber="564">Adult</emphasis>
. Male: Each forewing
<quantity id="A67F9B7D522AFFEBDA4F60C0FD69FF7D" box="[521,655,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.3" metricValueMin="5.9" pageId="13" pageNumber="564" unit="mm" value="6.1" valueMax="6.3" valueMin="5.9">5.96.3 mm</quantity>
long (mean =
<quantity id="A67F9B7D522AFFEBDB6860C1FC63FF7D" box="[814,901,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="13" pageNumber="564" unit="mm" value="6.1">6.1 mm</quantity>
, n = 5) and each hind wing
<quantity id="A67F9B7D522AFFEBDCF360C0FADDFF7D" box="[1205,1339,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="5.8" metricValueMin="5.4" pageId="13" pageNumber="564" unit="mm" value="5.6" valueMax="5.8" valueMin="5.4">5.45.8 mm</quantity>
(mean =
<quantity id="A67F9B7D522AFFEBD8D160ECFF08FF19" box="[151,238,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="13" pageNumber="564" unit="mm" value="5.6">5.6 mm</quantity>
, n = 5). Female: Each forewing
<quantity id="A67F9B7D522AFFEBDA1560EDFD3FFF19" box="[595,729,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="6.9" metricValueMin="6.6" pageId="13" pageNumber="564" unit="mm" value="6.75" valueMax="6.9" valueMin="6.6">6.66.9 mm</quantity>
(mean =
<quantity id="A67F9B7D522AFFEBDB0760EDFC7EFF19" box="[833,920,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="13" pageNumber="564" unit="mm" value="6.7">6.7 mm</quantity>
, n = 3) and each hind wing
<quantity id="A67F9B7D522AFFEBDC8C60ECFAB6FF19" box="[1226,1360,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.2" metricValueMin="5.8" pageId="13" pageNumber="564" unit="mm" value="6.0" valueMax="6.2" valueMin="5.8">5.86.2 mm</quantity>
(mean =
<quantity id="A67F9B7D522AFFEBD8E96089FEE1FF35" box="[175,263,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="13" pageNumber="564" unit="mm" value="6.1">6.1 mm</quantity>
, n = 3). Head dark brown, warts light brown with brown setae, antennae dark brown, palpi light brown. Thorax dark brown dorsally. Legs light brown with dark brown spurs. Wings brown with darker veins and fulvous pterostigma. Abdomen with dark brown tergites and light brown sternites, dark pigments scattered dorsally; scent glands of sternite V open on anterolateral-angle mounds well encircled by sutures; small mid-ventral process on each of sternites VI and VII (or on only VII) in the male and on sternite VI in the female (process on sternite VI of male often very small or completely absent).
</paragraph>
</subSubSection>
<subSubSection id="299D6513522AFFEBD88161E1FE4BF92D" pageId="13" pageNumber="564" type="description">
<paragraph id="61383698522AFFEBD88161E1FD8FFCD9" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="564">
<emphasis id="53F3EA8A522AFFEBD88161E1FE92FE1C" bold="true" box="[199,372,438,465]" pageId="13" pageNumber="564">Male genitalia</emphasis>
(
<figureCitation id="F9BC2A1D522AFFEBD9C261E1FDF3FE1D" box="[388,533,438,465]" captionStart="FIGURE 5" captionStartId="11.[151,250,1787,1811]" captionTargetBox="[254,1333,184,1758]" captionTargetId="figure-20@11.[241,1346,179,1761]" captionTargetPageId="11" captionText="FIGURE 5. Adult of Rhyacophila misago sp. nov. 5A5D, male: 5A, genitalia, left lateral, omitting left inferior appendage; 5B, same, dorsal; 5C, right inferior appendage, dorsal; 5D, right inferior appendage and phallic apparatus, ventral. 5E5I, female: 5E, abdominal segments VIIIXI, left lateral; 5F, segment VIII, dorsal; 5G, same, ventral; 5H, vaginal apparatus, left lateral; 5I, same, ventral.Abbreviations:VIIIXI = abdominal segments VIIIXI; a inf app = apical segment of inferior appendage (paired); ae = aedeagus; an sc = anal sclerite (paired); ap ba = apical band (paired); b inf app = basal segment of inferior appendage (paired); do me pr = dorsomesal process (paired); par = paramere (paired); pp = posterior process; ps = processus spermathecae; ru po IX = rugged posterior part of segment IX; sl cu pr = slender curved process (paired); te st = tergal strap (paired)." figureDoi="http://doi.org/10.5281/zenodo.14239190" httpUri="https://zenodo.org/record/14239190/files/figure.png" pageId="13" pageNumber="564">Figs 5A5D</figureCitation>
). Dorsal margin of segment IX (IX) about 3 times as long as ventral margin in lateral view. Rugged dorsal posterior part of segment IX (ru po IX) with several short acute processes near middle and pair of slender curved processes (sl cu pr) laterally; each slender process very long, directed caudad at first, curved dorsad at right angle near mid-length in lateral view and configured as semicircle in dorsal view. Segment X (X) depressed; subtriangular with incised apical margin in dorsal view; slender bar-like with slightly thickened apices in lateral view. Anal sclerite (an sc) strongly sclerotized, wide basally, gradually narrowing posteriorly with short longitudinal slit on midline apically in dorsal view; subrectangular with large, tall, subquadrate process basally in lateral view. Tergal strap (te st) and apical band (ap ba) membranous, vertically high in lateral view. Aedeagus (ae) long and slender. Each paramere (par) semicircular in ventral view, slender basally, gradually thickened to mid-length, and round apically in lateral view.
</paragraph>
<paragraph id="61383698522AFFEBD8816349FAEAFC05" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="564">Basal segment of each inferior appendage (b inf app) long, 2.5 times as long as apical segment (a inf app), in lateral view with dorsal and ventral margins parallel with each other; dorsomesal process (do me pr) arising from mesal side near dorso-caudal corner of basal segment, heavily sclerotized, hook-like in dorsal view, about 1/4 as long as basal segment; apical segment of each inferior appendage (a inf app) oval, diagonally positioned in lateral view, oblong in dorsal and ventral views, mesal surface dorsally and apically with numerous spinules.</paragraph>
<paragraph id="61383698522AFFEBD8816385FCF7FBB1" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="564">
<emphasis id="53F3EA8A522AFFEBD8816385FE74FC20" bold="true" box="[199,402,978,1005]" pageId="13" pageNumber="564">Female genitalia</emphasis>
(
<figureCitation id="F9BC2A1D522AFFEBD9E16385FDC9FC21" box="[423,559,978,1005]" captionStart="FIGURE 5" captionStartId="11.[151,250,1787,1811]" captionTargetBox="[254,1333,184,1758]" captionTargetId="figure-20@11.[241,1346,179,1761]" captionTargetPageId="11" captionText="FIGURE 5. Adult of Rhyacophila misago sp. nov. 5A5D, male: 5A, genitalia, left lateral, omitting left inferior appendage; 5B, same, dorsal; 5C, right inferior appendage, dorsal; 5D, right inferior appendage and phallic apparatus, ventral. 5E5I, female: 5E, abdominal segments VIIIXI, left lateral; 5F, segment VIII, dorsal; 5G, same, ventral; 5H, vaginal apparatus, left lateral; 5I, same, ventral.Abbreviations:VIIIXI = abdominal segments VIIIXI; a inf app = apical segment of inferior appendage (paired); ae = aedeagus; an sc = anal sclerite (paired); ap ba = apical band (paired); b inf app = basal segment of inferior appendage (paired); do me pr = dorsomesal process (paired); par = paramere (paired); pp = posterior process; ps = processus spermathecae; ru po IX = rugged posterior part of segment IX; sl cu pr = slender curved process (paired); te st = tergal strap (paired)." figureDoi="http://doi.org/10.5281/zenodo.14239190" httpUri="https://zenodo.org/record/14239190/files/figure.png" pageId="13" pageNumber="564">Figs 5E5I</figureCitation>
). Segment VIII (VIII) short, 0.8 times as long as basal height, gradually narrowing posteriorly in lateral, dorsal, and ventral views; posterolateral margins gently convex in lateral view, widely incised as V-shaped in dorsal view, shallowly and widely incised in ventral view. Vaginal apparatus long; processus spermathecae (ps) round in ventral view, semi-membranous; posterior process (pp) sclerotized, long, 1.5 times as long as processus spermathecae, round apically.
</paragraph>
<paragraph id="61383698522AFFEBD88164D0FB5DFB6D" blockId="13.[151,1437,150,2013]" box="[199,1211,1158,1185]" pageId="13" pageNumber="564">
<emphasis id="53F3EA8A522AFFEBD88164D0FE43FB6D" bold="true" box="[199,421,1158,1185]" pageId="13" pageNumber="564">Final Instar Larva</emphasis>
. (
<figureCitation id="F9BC2A1D522AFFEBD9FD64D1FDE6FB6D" box="[443,512,1159,1185]" captionStart="FIGURE 6" captionStartId="12.[151,250,1710,1734]" captionTargetBox="[158,1426,191,1678]" captionTargetId="figure-29@12.[151,1436,181,1686]" captionTargetPageId="12" captionText="FIGURE 6. Larva of Rhyacophila misago sp. nov. 6A, habitus, left lateral; 6B, head, dorsal; 6C, mandibles, ventral; 6D, pronotum, dorsal; 6E, pleuron and right foreleg, right lateral; 6F, pleuron and right hind leg, right lateral; 6G, left anal proleg, left lateral." figureDoi="http://doi.org/10.5281/zenodo.14239192" httpUri="https://zenodo.org/record/14239192/files/figure.png" pageId="13" pageNumber="564">Fig. 6</figureCitation>
). Length up to
<quantity id="A67F9B7D522AFFEBDAED64D0FD1AFB6D" box="[683,764,1158,1185]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="564" unit="mm" value="15.0">15 mm</quantity>
, width up to
<quantity id="A67F9B7D522AFFEBDBC864D0FC00FB6D" box="[910,998,1158,1185]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="564" unit="mm" value="2.5">2.5 mm</quantity>
, relatively stumpy.
</paragraph>
<paragraph id="61383698522AFFEBD88164FDFE32FAB5" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="564">
<emphasis id="53F3EA8A522AFFEBD88164FDFEE2FB09" box="[199,260,1195,1221]" italics="true" pageId="13" pageNumber="564">Head</emphasis>
(
<figureCitation id="F9BC2A1D522AFFEBD95264FDFE44FB09" box="[276,418,1195,1221]" captionStart="FIGURE 6" captionStartId="12.[151,250,1710,1734]" captionTargetBox="[158,1426,191,1678]" captionTargetId="figure-29@12.[151,1436,181,1686]" captionTargetPageId="12" captionText="FIGURE 6. Larva of Rhyacophila misago sp. nov. 6A, habitus, left lateral; 6B, head, dorsal; 6C, mandibles, ventral; 6D, pronotum, dorsal; 6E, pleuron and right foreleg, right lateral; 6F, pleuron and right hind leg, right lateral; 6G, left anal proleg, left lateral." figureDoi="http://doi.org/10.5281/zenodo.14239192" httpUri="https://zenodo.org/record/14239192/files/figure.png" pageId="13" pageNumber="564">Figs 6A6C</figureCitation>
). Width
<quantity id="A67F9B7D522AFFEBDA4564FDFD40FB09" box="[515,678,1195,1221]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" metricValueMax="9.6" metricValueMin="8.6" pageId="13" pageNumber="564" unit="mm" value="0.9099999999999999" valueMax="0.96" valueMin="0.86">0.860.96 mm</quantity>
(mean =
<quantity id="A67F9B7D522AFFEBDB5564FDFC9CFB09" box="[787,890,1195,1221]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="564" unit="mm" value="0.9">0.90 mm</quantity>
, n = 4); width: length: depth about 1.0: 1.6: 1.5. Brown with dark brown dots on parietals; frontoclypeal apotome dark brown with light colored rectangular pattern at middle separating darker regions laterally and anteriorly and with transverse pattern of four muscle scars at posterior part. Eyes rectangular. Labrum light brown, subquadrate with slightly incised anterior margin and 5 pairs of setae. Mandibles robust, deep brown, asymmetrical; left one with acute apex and mesal blade; right one with acute apex and mesal hump.
</paragraph>
<paragraph id="61383698522AFFEBD88165D5FAEDF9C5" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="564">
<emphasis id="53F3EA8A522AFFEBD88165D5FE7AFA50" box="[199,412,1411,1437]" italics="true" pageId="13" pageNumber="564">Thoracic segments</emphasis>
(
<figureCitation id="F9BC2A1D522AFFEBD9EB65D5FD8AFA51" box="[429,620,1411,1437]" captionStart="FIGURE 6" captionStartId="12.[151,250,1710,1734]" captionTargetBox="[158,1426,191,1678]" captionTargetId="figure-29@12.[151,1436,181,1686]" captionTargetPageId="12" captionText="FIGURE 6. Larva of Rhyacophila misago sp. nov. 6A, habitus, left lateral; 6B, head, dorsal; 6C, mandibles, ventral; 6D, pronotum, dorsal; 6E, pleuron and right foreleg, right lateral; 6F, pleuron and right hind leg, right lateral; 6G, left anal proleg, left lateral." figureDoi="http://doi.org/10.5281/zenodo.14239192" httpUri="https://zenodo.org/record/14239192/files/figure.png" pageId="13" pageNumber="564">Figs 6A, 6D6F</figureCitation>
). Pronotum completely covered with 2 light brown plates, anterolateral corners round, slightly narrowing posteriorly, incised mesally and rugged and darker in color along posterior margin; 1315 setae on setal areas 13 (
<emphasis id="53F3EA8A522AFFEBDA12659DFD74FA28" box="[596,658,1483,1508]" italics="true" pageId="13" pageNumber="564">sensu</emphasis>
<bibRefCitation id="05164B69522AFFEBDADF659DFCDFFA29" author="Wiggins, G. B." box="[665,825,1483,1509]" pageId="13" pageNumber="564" refId="ref13697" refString="Wiggins, G. B. (1996) Larvae of the North American Caddisfly Genera. Second Edition. University of Toronto Press, Toronto, 457 pp. https: // doi. org / 10.3138 / 9781442623606" type="book" year="1996">Wiggins 1996</bibRefCitation>
), many slender short setae on anterior edge. Meso- and metanota membranous; dark brown with several light color markings dorsally, almost white ventrally.
</paragraph>
<paragraph id="61383698522AFFEBD8816645FD8AF99D" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="564">Foretrochantin of prothorax and pleura of meso- and metathoraxes well developed. Forelegs short and somewhat stout, middle and hind legs rather slender.</paragraph>
<paragraph id="61383698522AFFEBD881660DFE4BF92D" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="564">
<emphasis id="53F3EA8A522AFFEBD881660DFED7F9B9" box="[199,305,1627,1653]" italics="true" pageId="13" pageNumber="564">Abdomen</emphasis>
(
<figureCitation id="F9BC2A1D522AFFEBD906660DFE36F9B9" box="[320,464,1627,1653]" captionStart="FIGURE 6" captionStartId="12.[151,250,1710,1734]" captionTargetBox="[158,1426,191,1678]" captionTargetId="figure-29@12.[151,1436,181,1686]" captionTargetPageId="12" captionText="FIGURE 6. Larva of Rhyacophila misago sp. nov. 6A, habitus, left lateral; 6B, head, dorsal; 6C, mandibles, ventral; 6D, pronotum, dorsal; 6E, pleuron and right foreleg, right lateral; 6F, pleuron and right hind leg, right lateral; 6G, left anal proleg, left lateral." figureDoi="http://doi.org/10.5281/zenodo.14239192" httpUri="https://zenodo.org/record/14239192/files/figure.png" pageId="13" pageNumber="564">Figs 6A, 6G</figureCitation>
). Moderately depressed dorsoventrally, gills absent. Segments IVIII dark brown with several light color markings and 13 setae dorsally; near white ventrally. Dorsal ellipsoidal sclerite on segment IX. Anal legs each with freely projecting lateral basoventral hook; apical claw with two ventral teeth, basal tooth smaller than distal tooth.
</paragraph>
</subSubSection>
<subSubSection id="299D6513522AFFE8D88166BDFE3BFE45" lastPageId="14" lastPageNumber="565" pageId="13" pageNumber="564" type="materials_examined">
<paragraph id="61383698522AFFEBD88166BDFD5DF8E5" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="564">
<materialsCitation id="D1EF3CC5522AFFEBD88166BDFD5DF8E5" collectingDate="2006-08-04" collectionCode="TI" county="Male" elevation="5" latitude="42.5022" location="Fukushima-cho" longLatPrecision="6" longitude="140.3269" municipality="Oshima" pageId="13" pageNumber="564" specimenCode="CBM-ZI 0189176" specimenCount="1" stateProvince="Hokkaido" typeStatus="holotype">
<emphasis id="53F3EA8A522AFFEBD88166BDFED2F8C9" bold="true" box="[199,308,1771,1797]" pageId="13" pageNumber="564">
<typeStatus id="BE3C883A522AFFEBD88166BDFED2F8C9" box="[199,308,1771,1797]" pageId="13" pageNumber="564" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingCounty id="88594E14522AFFEBD90266BDFE66F8C9" box="[324,384,1771,1797]" pageId="13" pageNumber="564">Male</collectingCounty>
,
<emphasis id="53F3EA8A522AFFEBD9C866BCFDE2F8C8" bold="true" box="[398,516,1770,1796]" pageId="13" pageNumber="564">
<collectingRegion id="A343F87A522AFFEBD9C866BCFDE2F8C8" box="[398,516,1770,1796]" country="Japan" name="Hokkaido" pageId="13" pageNumber="564">Hokkaido</collectingRegion>
</emphasis>
,
<emphasis id="53F3EA8A522AFFEBDA5366BCFD94F8C8" bold="true" box="[533,626,1770,1796]" pageId="13" pageNumber="564">
<collectingMunicipality id="815CACE2522AFFEBDA5366BCFD94F8C8" box="[533,626,1770,1796]" pageId="13" pageNumber="564">Oshima</collectingMunicipality>
</emphasis>
,
<location id="64586043522AFFEBDAC566BDFCD2F8C9" LSID="urn:lsid:plazi:treatment:E92E878E522DFFE8D8D16540FD79FDD5:64586043522AFFEBDAC566BDFCD2F8C9" box="[643,820,1771,1797]" county="Male" latitude="42.5022" longLatPrecision="6" longitude="140.3269" municipality="Oshima" name="Fukushima-cho" pageId="13" pageNumber="564" stateProvince="Hokkaido">Fukushima-chô</location>
, Misago-no-taki (
<geoCoordinate id="04B3505F522AFFEBDBB966BCFB98F8C9" box="[1023,1150,1770,1797]" degrees="42.5022" direction="north" orientation="latitude" pageId="13" pageNumber="564" precision="5" value="42.5022">42.5022°N</geoCoordinate>
,
<geoCoordinate id="04B3505F522AFFEBDCCA66BDFAF2F8C9" box="[1164,1300,1771,1797]" degrees="140.3269" direction="east" orientation="longitude" pageId="13" pageNumber="564" precision="5" value="140.3269">140.3269°E</geoCoordinate>
,
<elevation id="EAAAD1AB522AFFEBDD6766BCFA6AF8C9" box="[1313,1420,1770,1797]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="564" unit="m" value="5.0">
<quantity id="A67F9B7D522AFFEBDD6766BCFAA9F8C9" box="[1313,1359,1770,1797]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="564" unit="m" value="5.0">5 m</quantity>
a.s.l.
</elevation>
),
<date id="15391058522AFFEBD8D16759FEF5F8E5" box="[151,275,1807,1833]" pageId="13" pageNumber="564" value="2006-08-04">
<collectingDate id="057DE9B0522AFFEBD8D16759FEF5F8E5" box="[151,275,1807,1833]" pageId="13" pageNumber="564" value="2006-08-04">4.viii.2006</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D522AFFEBD9086759FE8EF8E5" box="[334,360,1807,1833]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="13" pageNumber="564">TI</collectionCode>
&amp; TH, S (
<specimenCode id="31219EE3522AFFEBD99B6759FD49F8E5" box="[477,687,1807,1833]" collectionCode="CBM-ZI" pageId="13" pageNumber="564">CBM-ZI 0189176</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="61383698522AFFEBD8816765FA6BF8BD" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="564">
<materialsCitation id="D1EF3CC5522AFFEBD8816765FC58F881" box="[199,958,1843,1869]" collectingDate="2006-08-04" collectionCode="SPMN-IS" county="Male" elevation="5" latitude="42.5022" location="Fukushima-cho" longLatPrecision="6" longitude="140.3269" municipality="Oshima" pageId="13" pageNumber="564" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Hokkaido" typeStatus="paratype">
<emphasis id="53F3EA8A522AFFEBD8816765FED9F881" bold="true" box="[199,319,1843,1869]" pageId="13" pageNumber="564">
<typeStatus id="BE3C883A522AFFEBD8816765FED9F881" box="[199,319,1843,1869]" pageId="13" pageNumber="564" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<specimenCount id="7781FD11522AFFEBD9166765FE47F881" box="[336,417,1843,1869]" count="1" pageId="13" pageNumber="564" type="male">1 male</specimenCount>
,
<specimenCount id="7781FD11522AFFEBD9E96765FDC7F881" box="[431,545,1843,1869]" count="2" pageId="13" pageNumber="564" type="female">2 females</specimenCount>
, same data as holotype (
<collectionCode id="0796AE5D522AFFEBDB076765FC50F881" box="[833,950,1843,1869]" pageId="13" pageNumber="564">SPMN-IS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D1EF3CC5522AFFEBDB8A6765FCEFF8BD" collectingDate="2007-08-04" collectingDateMax="2007-08-29" collectingDateMin="2007-08-04" collectionCode="TI, SPMN-IS" pageId="13" pageNumber="564" specimenCount="1" specimenCount-male="1" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCount id="7781FD11522AFFEBDB8A6765FBFBF881" box="[972,1053,1843,1869]" count="1" pageId="13" pageNumber="564" type="male">1 male</specimenCount>
, type locality,
<date id="15391058522AFFEBDC8F6765FAA4F881" box="[1225,1346,1843,1869]" pageId="13" pageNumber="564" value="2007-08-04">
<collectingDate id="057DE9B0522AFFEBDC8F6765FAA4F881" box="[1225,1346,1843,1869]" pageId="13" pageNumber="564" value="2007-08-04">4.viii.2007</collectingDate>
</date>
(pupa), reared and emerged on
<date id="15391058522AFFEBD9DB6701FDC1F8BD" box="[413,551,1879,1905]" pageId="13" pageNumber="564" value="2007-08-29">
<collectingDate id="057DE9B0522AFFEBD9DB6701FDC1F8BD" box="[413,551,1879,1905]" pageId="13" pageNumber="564" value="2007-08-29">29.viii.2007</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D522AFFEBDA246701FD9AF8BD" box="[610,636,1879,1905]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="13" pageNumber="564">TI</collectionCode>
(
<collectionCode id="0796AE5D522AFFEBDACA6701FCE7F8BD" box="[652,769,1879,1905]" pageId="13" pageNumber="564">SPMN-IS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D1EF3CC5522AFFEBDB526701FB04F8BD" box="[788,1250,1879,1905]" collectingDate="2007-09-30" collectionCode="TI" pageId="13" pageNumber="564" specimenCount="2" specimenCount-male="2" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCount id="7781FD11522AFFEBDB526701FC8BF8BD" box="[788,877,1879,1905]" count="2" pageId="13" pageNumber="564" type="male">2 males</specimenCount>
, type locality,
<date id="15391058522AFFEBDC496701FB6CF8BD" box="[1039,1162,1879,1905]" pageId="13" pageNumber="564" value="2007-09-30">
<collectingDate id="057DE9B0522AFFEBDC496701FB6CF8BD" box="[1039,1162,1879,1905]" pageId="13" pageNumber="564" value="2007-09-30">30.ix.2007</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D522AFFEBDC836701FB04F8BD" box="[1221,1250,1879,1905]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="13" pageNumber="564">TI</collectionCode>
</materialsCitation>
, S (
<collectionCode id="0796AE5D522AFFEBDD4A6701FA67F8BD" box="[1292,1409,1879,1905]" pageId="13" pageNumber="564">SPMN-IS</collectionCode>
).
</paragraph>
<paragraph id="61383698522AFFE8D881672CFE3BFE45" blockId="13.[151,1437,150,2013]" lastBlockId="14.[151,1436,151,537]" lastPageId="14" lastPageNumber="565" pageId="13" pageNumber="564">
<materialsCitation id="D1EF3CC5522AFFEBD881672CFACBF859" box="[199,1325,1914,1941]" collectingDate="1997-07-22" collectionCode="TI" location="Yakumo-cho" municipality="Oshima" pageId="13" pageNumber="564" specimenCount="6" specimenCount-male="6" stateProvince="Hokkaido">
<emphasis id="53F3EA8A522AFFEBD881672CFE76F858" bold="true" box="[199,400,1914,1940]" pageId="13" pageNumber="564">
<typeStatus id="BE3C883A522AFFEBD881672CFE76F858" box="[199,400,1914,1940]" isOtherTypeStatus="true" pageId="13" pageNumber="564">Other specimens</typeStatus>
</emphasis>
.
<emphasis id="53F3EA8A522AFFEBD9E7672CFDF1F858" bold="true" box="[417,535,1914,1940]" pageId="13" pageNumber="564">
<collectingRegion id="A343F87A522AFFEBD9E7672CFDF1F858" box="[417,535,1914,1940]" country="Japan" name="Hokkaido" pageId="13" pageNumber="564">Hokkaido</collectingRegion>
</emphasis>
,
<emphasis id="53F3EA8A522AFFEBDA61672CFD62F858" bold="true" box="[551,644,1914,1940]" pageId="13" pageNumber="564">
<collectingMunicipality id="815CACE2522AFFEBDA61672CFD62F858" box="[551,644,1914,1940]" pageId="13" pageNumber="564">Oshima</collectingMunicipality>
</emphasis>
:
<location id="64586043522AFFEBDAD2672DFCC0F859" LSID="urn:lsid:plazi:treatment:E92E878E522DFFE8D8D16540FD79FDD5:64586043522AFFEBDAD2672DFCC0F859" box="[660,806,1915,1941]" municipality="Oshima" name="Yakumo-cho" pageId="13" pageNumber="564" stateProvince="Hokkaido">Yakumo-chô</location>
, Goyô-no-sawa,
<specimenCount id="7781FD11522AFFEBDBAF672DFBA2F859" box="[1001,1092,1915,1941]" count="6" pageId="13" pageNumber="564" type="male">6 males</specimenCount>
,
<date id="15391058522AFFEBDC14672DFB32F859" box="[1106,1236,1915,1941]" pageId="13" pageNumber="564" value="1997-07-22">
<collectingDate id="057DE9B0522AFFEBDC14672DFB32F859" box="[1106,1236,1915,1941]" pageId="13" pageNumber="564" value="1997-07-22">22.vii.1997</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D522AFFEBDD55672DFACBF859" box="[1299,1325,1915,1941]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="13" pageNumber="564">TI</collectionCode>
</materialsCitation>
&amp;
<materialsCitation id="D1EF3CC5522AFFEBDD12672DFA7AF875" collectingDate="1997-07-23" collectionCode="SPMN-IS, TI" location="Yurappu-gawa" municipality="Yakumo-cho" pageId="13" pageNumber="564" specimenCount="6" specimenCount-female="1" specimenCount-larva="4" specimenCount-male="1" stateProvince="Hokkaido">
AO, S (
<collectionCode id="0796AE5D522AFFEBD8D967C9FEF2F875" box="[159,276,1951,1977]" pageId="13" pageNumber="564">SPMN-IS</collectionCode>
).
<collectingMunicipality id="815CACE2522AFFEBD96E67C9FE5CF875" box="[296,442,1951,1977]" pageId="13" pageNumber="564">Yakumo-chô</collectingMunicipality>
, small waterfall beside
<location id="64586043522AFFEBDA8C67C9FC95F875" LSID="urn:lsid:plazi:treatment:E92E878E522DFFE8D8D16540FD79FDD5:64586043522AFFEBDA8C67C9FC95F875" box="[714,883,1951,1977]" municipality="Yakumo-cho" name="Yurappu-gawa" pageId="13" pageNumber="564" stateProvince="Hokkaido">Yûrappu-gawa</location>
,
<specimenCount id="7781FD11522AFFEBDBC767C9FC34F875" box="[897,978,1951,1977]" count="1" pageId="13" pageNumber="564" type="male">1 male</specimenCount>
,
<specimenCount id="7781FD11522AFFEBDB9967C9FBA0F875" box="[991,1094,1951,1977]" count="1" pageId="13" pageNumber="564" type="female">1 female</specimenCount>
,
<specimenCount id="7781FD11522AFFEBDC1567C9FB54F875" box="[1107,1202,1951,1977]" count="4" pageId="13" pageNumber="564" type="larva">4 larvae</specimenCount>
,
<date id="15391058522AFFEBDC8667C9FAA7F875" box="[1216,1345,1951,1977]" pageId="13" pageNumber="564" value="1997-07-23">
<collectingDate id="057DE9B0522AFFEBDC8667C9FAA7F875" box="[1216,1345,1951,1977]" pageId="13" pageNumber="564" value="1997-07-23">23.vii.1997</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D522AFFEBDDC467C9FA7AF875" box="[1410,1436,1951,1977]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="13" pageNumber="564">TI</collectionCode>
</materialsCitation>
&amp;
<materialsCitation id="D1EF3CC5522AFFEBD8F56795FA67F811" box="[179,1409,1986,2013]" collectingDate="1995-06-23" collectionCode="SPMN-IS, TI" location="Yurappu-gawa" municipality="Yakumo-cho" pageId="13" pageNumber="564" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido">
AO, S (
<collectionCode id="0796AE5D522AFFEBD94F6795FE98F811" box="[265,382,1987,2013]" pageId="13" pageNumber="564">SPMN-IS</collectionCode>
).
<collectingMunicipality id="815CACE2522AFFEBD9D66795FDC4F811" box="[400,546,1987,2013]" pageId="13" pageNumber="564">Yakumo-chô</collectingMunicipality>
, small waterfall beside
<location id="64586043522AFFEBDB6A6795FC33F811" LSID="urn:lsid:plazi:treatment:E92E878E522DFFE8D8D16540FD79FDD5:64586043522AFFEBDB6A6795FC33F811" box="[812,981,1987,2013]" municipality="Yakumo-cho" name="Yurappu-gawa" pageId="13" pageNumber="564" stateProvince="Hokkaido">Yûrappu-gawa</location>
,
<specimenCount id="7781FD11522AFFEBDBA76795FBD6F811" box="[993,1072,1987,2013]" count="1" pageId="13" pageNumber="564" type="male">1 male</specimenCount>
,
<specimenCount id="7781FD11522AFFEBDC7A6795FB47F811" box="[1084,1185,1987,2013]" count="1" pageId="13" pageNumber="564" type="female">1 female</specimenCount>
,
<date id="15391058522AFFEBDCEB6794FAC1F811" box="[1197,1319,1986,2013]" pageId="13" pageNumber="564" value="1995-06-23">
<collectingDate id="057DE9B0522AFFEBDCEB6794FAC1F811" box="[1197,1319,1986,2013]" pageId="13" pageNumber="564" value="1995-06-23">23.vi.1995</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D522AFFEBDD226795FA67F811" box="[1380,1409,1987,2013]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="13" pageNumber="564">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC5522AFFE8DDCB6795FB01FF7D" collectingDate="2003-06-22" collectionCode="TI, HM" lastPageId="14" lastPageNumber="565" location="Namari-gawa" municipality="Yakumo-cho" pageId="13" pageNumber="564" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5229FFE8D8E660C1FF59FF7D" box="[160,191,151,177]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="14" pageNumber="565">TI</collectionCode>
).
<collectingMunicipality id="815CACE25229FFE8D89560C1FE83FF7D" box="[211,357,151,177]" pageId="14" pageNumber="565">Yakumo-chô</collectingMunicipality>
, small water flow beside
<location id="645860435229FFE8DACF60C1FCC2FF7D" LSID="urn:lsid:plazi:treatment:E92E878E522DFFE8D8D16540FD79FDD5:645860435229FFE8DACF60C1FCC2FF7D" box="[649,804,151,177]" municipality="Yakumo-cho" name="Namari-gawa" pageId="14" pageNumber="565" stateProvince="Hokkaido">Namari-gawa</location>
,
<specimenCount id="7781FD115229FFE8DB7660C1FC66FF7D" box="[816,896,151,177]" count="1" pageId="14" pageNumber="565" type="male">1 male</specimenCount>
,
<specimenCount id="7781FD115229FFE8DBCB60C1FC14FF7D" box="[909,1010,151,177]" count="1" pageId="14" pageNumber="565" type="female">1 female</specimenCount>
,
<date id="153910585229FFE8DBB960C1FB9EFF7D" box="[1023,1144,151,177]" pageId="14" pageNumber="565" value="2003-06-22">
<collectingDate id="057DE9B05229FFE8DBB960C1FB9EFF7D" box="[1023,1144,151,177]" pageId="14" pageNumber="565" value="2003-06-22">22.vi.2003</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5229FFE8DCF160C1FB01FF7D" box="[1207,1255,151,177]" country="0" name="Hastings Museum" pageId="14" pageNumber="565">HM</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55229FFE8DCB260C1FBBAFF19" collectingDate="1980-06-10" collectionCode="HM, TI" county="Mori-machi" location="Ojironai-gawa" municipality="Himekawa" pageId="14" pageNumber="565" specimenCount="2" specimenCount-male="2" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5229FFE8DD5260C1FAA1FF7D" box="[1300,1351,151,177]" country="0" name="Hastings Museum" pageId="14" pageNumber="565">HM</collectionCode>
).
<collectingCounty id="88594E145229FFE8DD1D60C1FF38FF19" pageId="14" pageNumber="565">Mori-machi</collectingCounty>
,
<collectingMunicipality id="815CACE25229FFE8D8AC60EDFE80FF19" box="[234,358,187,213]" pageId="14" pageNumber="565">Himekawa</collectingMunicipality>
, small waterfall beside
<location id="645860435229FFE8DA3460EDFCF1FF19" LSID="urn:lsid:plazi:treatment:E92E878E522DFFE8D8D16540FD79FDD5:645860435229FFE8DA3460EDFCF1FF19" box="[626,791,187,213]" county="Mori-machi" municipality="Himekawa" name="Ojironai-gawa" pageId="14" pageNumber="565" stateProvince="Hokkaido">Ojironai-gawa</location>
,
<specimenCount id="7781FD115229FFE8DB6460EDFC9AFF19" box="[802,892,187,213]" count="2" pageId="14" pageNumber="565" type="male">2 males</specimenCount>
,
<date id="153910585229FFE8DBCE60EDFBE4FF19" box="[904,1026,187,213]" pageId="14" pageNumber="565" value="1980-06-10">
<collectingDate id="057DE9B05229FFE8DBCE60EDFBE4FF19" box="[904,1026,187,213]" pageId="14" pageNumber="565" value="1980-06-10">10.vi.1980</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5229FFE8DC7960EDFBBAFF19" box="[1087,1116,187,213]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="14" pageNumber="565">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55229FFE8DC2E60EDFCD4FF35" collectingDate="2003-06-24" collectionCode="SPMN-IS, NK" location="Torisaki-gawa" municipality="Mori-machi" pageId="14" pageNumber="565" specimenCount="1" specimenCount-male="1" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5229FFE8DCC160EDFB1AFF19" box="[1159,1276,187,213]" pageId="14" pageNumber="565">SPMN-IS</collectionCode>
).
<collectingMunicipality id="815CACE25229FFE8DD4960EDFA7EFF19" box="[1295,1432,187,213]" pageId="14" pageNumber="565">Mori-machi</collectingMunicipality>
, waterfall beside
<location id="645860435229FFE8D9086089FE17FF35" LSID="urn:lsid:plazi:treatment:E92E878E522DFFE8D8D16540FD79FDD5:645860435229FFE8D9086089FE17FF35" box="[334,497,223,249]" municipality="Mori-machi" name="Torisaki-gawa" pageId="14" pageNumber="565" stateProvince="Hokkaido">Torisaki-gawa</location>
,
<specimenCount id="7781FD115229FFE8D9BD6089FDAEFF35" box="[507,584,223,249]" count="1" pageId="14" pageNumber="565" type="male">1 male</specimenCount>
,
<date id="153910585229FFE8DA146089FD2BFF35" box="[594,717,223,249]" pageId="14" pageNumber="565" value="2003-06-24">
<collectingDate id="057DE9B05229FFE8DA146089FD2BFF35" box="[594,717,223,249]" pageId="14" pageNumber="565" value="2003-06-24">24.vi.2003</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5229FFE8DB416089FCD4FF35" box="[775,818,223,249]" pageId="14" pageNumber="565">NK</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55229FFE8DB7A6089FE80FED1" collectingDate="2007-08-05" collectionCode="NK, TI" location="Torisaki Keikoku" municipality="Mori-machi" pageId="14" pageNumber="565" specimenCount="1" specimenCount-pupa="1" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5229FFE8DB1C6089FC6EFF35" box="[858,904,223,249]" pageId="14" pageNumber="565">NK</collectionCode>
).
<collectingMunicipality id="815CACE25229FFE8DBDC6089FBC5FF35" box="[922,1059,223,249]" pageId="14" pageNumber="565">Mori-machi</collectingMunicipality>
,
<location id="645860435229FFE8DC6A6089FB17FF35" LSID="urn:lsid:plazi:treatment:E92E878E522DFFE8D8D16540FD79FDD5:645860435229FFE8DC6A6089FB17FF35" box="[1068,1265,223,249]" municipality="Mori-machi" name="Torisaki Keikoku" pageId="14" pageNumber="565" stateProvince="Hokkaido">Torisaki Keikoku</location>
,
<specimenCount id="7781FD115229FFE8DCBD6089FAA0FF35" box="[1275,1350,223,249]" count="1" pageId="14" pageNumber="565" type="pupa">1 pupa</specimenCount>
(male),
<date id="153910585229FFE8D8D16154FEF5FED1" box="[151,275,258,285]" pageId="14" pageNumber="565" value="2007-08-05">
<collectingDate id="057DE9B05229FFE8D8D16154FEF5FED1" box="[151,275,258,285]" pageId="14" pageNumber="565" value="2007-08-05">5.viii.2007</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5229FFE8D90A6155FE80FED1" box="[332,358,259,285]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="14" pageNumber="565">TI</collectionCode>
</materialsCitation>
&amp;
<materialsCitation id="D1EF3CC55229FFE8D9CE6155FF00FE8D" collectingDate="2023-06-20" collectionCode="SPMN-IS, TI" location="Fukushima-cho" pageId="14" pageNumber="565" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido">
TH (
<collectionCode id="0796AE5D5229FFE8D9FD6155FDD6FED1" box="[443,560,259,285]" pageId="14" pageNumber="565">SPMN-IS</collectionCode>
).
<location id="645860435229FFE8DA076155FD14FED1" LSID="urn:lsid:plazi:treatment:E92E878E522DFFE8D8D16540FD79FDD5:645860435229FFE8DA076155FD14FED1" box="[577,754,259,285]" name="Fukushima-cho" pageId="14" pageNumber="565" stateProvince="Hokkaido">Fukushima-chô</location>
, Misago-no-taki (
<typeStatus id="BE3C883A5229FFE8DBF16155FC0FFED1" box="[951,1001,259,285]" pageId="14" pageNumber="565">type</typeStatus>
locality),
<specimenCount id="7781FD115229FFE8DC1F6155FB40FED1" box="[1113,1190,259,285]" count="1" pageId="14" pageNumber="565" type="male">1 male</specimenCount>
,
<specimenCount id="7781FD115229FFE8DCF66155FAF5FED1" box="[1200,1299,259,285]" count="1" pageId="14" pageNumber="565" type="female">1 female</specimenCount>
,
<date id="153910585229FFE8DD5B6155FA71FED1" box="[1309,1431,259,285]" pageId="14" pageNumber="565" value="2023-06-20">
<collectingDate id="057DE9B05229FFE8DD5B6155FA71FED1" box="[1309,1431,259,285]" pageId="14" pageNumber="565" value="2023-06-20">20.vi.2023</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5229FFE8D88F6171FF00FE8D" box="[201,230,295,321]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="14" pageNumber="565">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55229FFE8D8B26171FB68FE8D" box="[244,1166,295,321]" collectingDate="2007-09-30" collectionCode="TI, SPMN-IS" location="Fukushima-cho" pageId="14" pageNumber="565" specimenCount="1" specimenCount-larva="1" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5229FFE8D9506171FED3FE8D" box="[278,309,295,321]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="14" pageNumber="565">TI</collectionCode>
).
<location id="645860435229FFE8D90D6171FE1AFE8D" LSID="urn:lsid:plazi:treatment:E92E878E522DFFE8D8D16540FD79FDD5:645860435229FFE8D90D6171FE1AFE8D" box="[331,508,295,321]" name="Fukushima-cho" pageId="14" pageNumber="565" stateProvince="Hokkaido">Fukushima-chô</location>
, Shiraito-no-taki,
<specimenCount id="7781FD115229FFE8DA8D6171FCF8FE8D" box="[715,798,295,321]" count="1" pageId="14" pageNumber="565" type="larva">1 larva</specimenCount>
,
<date id="153910585229FFE8DB6D6171FC40FE8D" box="[811,934,295,321]" pageId="14" pageNumber="565" value="2007-09-30">
<collectingDate id="057DE9B05229FFE8DB6D6171FC40FE8D" box="[811,934,295,321]" pageId="14" pageNumber="565" value="2007-09-30">30.ix.2007</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5229FFE8DBA36171FC19FE8D" box="[997,1023,295,321]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="14" pageNumber="565">TI</collectionCode>
(
<collectionCode id="0796AE5D5229FFE8DC576171FB60FE8D" box="[1041,1158,295,321]" pageId="14" pageNumber="565">SPMN-IS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="D1EF3CC55229FFE8DCDD6171FD79FEA9" collectingDate="2006-09-12" collectionCode="TI" location="Fudono-taki" municipality="Fukushima-cho" pageId="14" pageNumber="565" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido">
<collectingMunicipality id="815CACE25229FFE8DCDD6171FAAAFE8D" box="[1179,1356,295,321]" pageId="14" pageNumber="565">Fukushima-chô</collectingMunicipality>
,
<location id="645860435229FFE8DD1F6171FF0FFEA9" LSID="urn:lsid:plazi:treatment:E92E878E522DFFE8D8D16540FD79FDD5:645860435229FFE8DD1F6171FF0FFEA9" municipality="Fukushima-cho" name="Fudono-taki" pageId="14" pageNumber="565" stateProvince="Hokkaido">Fudôno-taki</location>
,
<specimenCount id="7781FD115229FFE8D8B1611DFEA1FEA9" box="[247,327,331,357]" count="1" pageId="14" pageNumber="565" type="male">1 male</specimenCount>
,
<specimenCount id="7781FD115229FFE8D913611DFE5DFEA9" box="[341,443,331,357]" count="1" pageId="14" pageNumber="565" type="female">1 female</specimenCount>
,
<date id="153910585229FFE8D98E611DFDA5FEA9" box="[456,579,331,357]" pageId="14" pageNumber="565" value="2006-09-12">
<collectingDate id="057DE9B05229FFE8D98E611DFDA5FEA9" box="[456,579,331,357]" pageId="14" pageNumber="565" value="2006-09-12">12.ix.2006</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5229FFE8DAC4611DFD79FEA9" box="[642,671,331,357]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="14" pageNumber="565">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55229FFE8DAEB611DFE9EFE45" collectingDate="1995-06-21" collectionCode="SPMN-IS, NK" location="Iwabe-gawa" municipality="Fukushima-cho" pageId="14" pageNumber="565" specimenCount="1" specimenCount-male="1" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5229FFE8DA88611DFCA5FEA9" box="[718,835,331,357]" pageId="14" pageNumber="565">SPMN-IS</collectionCode>
).
<collectingMunicipality id="815CACE25229FFE8DB11611DFBEEFEA9" box="[855,1032,331,357]" pageId="14" pageNumber="565">Fukushima-chô</collectingMunicipality>
,
<location id="645860435229FFE8DC50611DFB44FEA9" LSID="urn:lsid:plazi:treatment:E92E878E522DFFE8D8D16540FD79FDD5:645860435229FFE8DC50611DFB44FEA9" box="[1046,1186,331,357]" municipality="Fukushima-cho" name="Iwabe-gawa" pageId="14" pageNumber="565" stateProvince="Hokkaido">Iwabe-gawa</location>
, near estuary,
<specimenCount id="7781FD115229FFE8DD0E611DFA7EFEA9" box="[1352,1432,331,357]" count="1" pageId="14" pageNumber="565" type="male">1 male</specimenCount>
,
<date id="153910585229FFE8D8D16138FEF7FE45" box="[151,273,366,393]" pageId="14" pageNumber="565" value="1995-06-21">
<collectingDate id="057DE9B05229FFE8D8D16138FEF7FE45" box="[151,273,366,393]" pageId="14" pageNumber="565" value="1995-06-21">21.vi.1995</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5229FFE8D90B6139FE9EFE45" box="[333,376,367,393]" pageId="14" pageNumber="565">NK</collectionCode>
</materialsCitation>
, S (
<collectionCode id="0796AE5D5229FFE8D9E46139FE36FE45" box="[418,464,367,393]" pageId="14" pageNumber="565">NK</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="299D65135229FFE8D88161C4FB1FFE1D" pageId="14" pageNumber="565" type="distribution">
<paragraph id="613836985229FFE8D88161C4FC05FE61" blockId="14.[151,1436,151,537]" box="[199,995,402,429]" pageId="14" pageNumber="565">
<emphasis id="53F3EA8A5229FFE8D88161C4FEBFFE60" bold="true" box="[199,345,402,428]" pageId="14" pageNumber="565">Distribution</emphasis>
(
<figureCitation id="F9BC2A1D5229FFE8D92F61C5FE27FE61" box="[361,449,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1655,1679]" captionTargetBox="[165,1422,186,1625]" captionTargetId="figure-29@2.[151,1436,179,1630]" captionTargetPageId="2" captionText="FIGURE 1. Recorded collection sites of Rhyacophila species. 1A, location of Hokkaido and recorded sites of known species discussed in present study. 1B1E, recorded and collected sites in Hokkaido: 1B, 1C, R. eniwa sp. nov. and R. arefini Lukyanchenko; 1D, R. bifue sp. nov., R. imakane sp. nov., R. misago sp. nov., and R. hamata Hattori &amp; Ohkawa; 1E, R. shimamaki sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14239182" httpUri="https://zenodo.org/record/14239182/files/figure.png" pageId="14" pageNumber="565">Fig. 1D</figureCitation>
, white triangles).
<collectingRegion id="A343F87A5229FFE8DACC61C5FD1EFE61" box="[650,760,403,429]" country="Japan" name="Hokkaido" pageId="14" pageNumber="565">Hokkaido</collectingRegion>
(southwestern area).
</paragraph>
<paragraph id="613836985229FFE8D88161E0FB1FFE1D" blockId="14.[151,1436,151,537]" box="[199,1273,438,465]" pageId="14" pageNumber="565">
<emphasis id="53F3EA8A5229FFE8D88161E0FEC5FE1C" bold="true" box="[199,291,438,464]" pageId="14" pageNumber="565">Habitat</emphasis>
. Adults of this species were collected beside waterfalls and small, fast flowing streams.
</paragraph>
</subSubSection>
<subSubSection id="299D65135229FFE8D881618DFD79FDD5" pageId="14" pageNumber="565" type="etymology">
<paragraph id="613836985229FFE8D881618DFAFBFE39" blockId="14.[151,1436,151,537]" box="[199,1309,475,501]" pageId="14" pageNumber="565">
<emphasis id="53F3EA8A5229FFE8D881618DFEAEFE39" bold="true" box="[199,328,475,501]" pageId="14" pageNumber="565">Etymology</emphasis>
. The name
<taxonomicName id="A6874D1B5229FFE8D98A618DFDCFFE38" authorityName="Ito &amp; Kuhara &amp; Malicky" authorityYear="2024" box="[460,553,475,501]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="565" phylum="Arthropoda" rank="species" species="misago">
<emphasis id="53F3EA8A5229FFE8D99E618DFDCFFE38" box="[472,553,475,500]" italics="true" pageId="14" pageNumber="565">misago</emphasis>
</taxonomicName>
” is a noun in apposition, coined from the name of the
<typeStatus id="BE3C883A5229FFE8DCCA618DFB5AFE39" box="[1164,1212,475,501]" pageId="14" pageNumber="565">type</typeStatus>
locality.
</paragraph>
<paragraph id="613836985229FFE8D88161A9FD79FDD5" blockId="14.[151,1436,151,537]" box="[199,671,511,537]" pageId="14" pageNumber="565">
<emphasis id="53F3EA8A5229FFE8D88161A9FE9AFDD5" bold="true" box="[199,380,511,537]" pageId="14" pageNumber="565">Japanese name</emphasis>
. Misago-nagare-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>