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<mods:title id="3FC6306E9ADBD60200D7F87B5D485604">Five new species of the genus Rhyacophila Stephens (Trichoptera, Rhyacophilidae) from southwestern Hokkaido, northern Japan</mods:title>
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<mods:namePart id="01A373F5383B9A6499E8C9240D58E21A">Ito, Tomiko</mods:namePart>
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<taxonomicName id="A6874D1B5226FFE7D8D16280FD68FD3C" ID-CoL="4SNMH" authority="Lukyanchenko 1993" authorityName="Lukyanchenko" authorityYear="1993" box="[151,654,726,753]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="species" species="arefini">
<emphasis id="53F3EA8A5226FFE7D8D16280FD68FD3C" bold="true" box="[151,654,726,753]" pageId="1" pageNumber="552">
<emphasis id="53F3EA8A5226FFE7D8D16280FE6FFD3C" bold="true" box="[151,393,726,753]" italics="true" pageId="1" pageNumber="552">Rhyacophila arefini</emphasis>
<bibRefCitation id="05164B695226FFE7D9D76281FD68FD3C" author="Lukyanchenko, T. I." box="[401,654,726,753]" pageId="1" pageNumber="552" pagination="5 - 6" refId="ref13054" refString="Lukyanchenko, T. I. (1993) A new species of caddisfly of the genus Rhyacophila Pictet (Trichoptera: Rhyacophilidae). Braueria, 20, 5 - 6." type="journal article" year="1993">Lukyanchenko 1993</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="299D65135226FFE7D8D162ADFED4FCD9" box="[151,306,763,789]" pageId="1" pageNumber="552" type="description">
<paragraph id="613836985226FFE7D8D162ADFED4FCD9" blockId="1.[151,654,726,789]" box="[151,306,763,789]" pageId="1" pageNumber="552">
(
<figureCitation id="F9BC2A1D5226FFE7D8D962ADFECFFCD9" box="[159,297,763,789]" captionStart="FIGURE 1" captionStartId="2.[151,250,1655,1679]" captionTargetBox="[165,1422,186,1625]" captionTargetId="figure-29@2.[151,1436,179,1630]" captionTargetPageId="2" captionText="FIGURE 1. Recorded collection sites of Rhyacophila species. 1A, location of Hokkaido and recorded sites of known species discussed in present study. 1B1E, recorded and collected sites in Hokkaido: 1B, 1C, R. eniwa sp. nov. and R. arefini Lukyanchenko; 1D, R. bifue sp. nov., R. imakane sp. nov., R. misago sp. nov., and R. hamata Hattori &amp; Ohkawa; 1E, R. shimamaki sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14239182" httpUri="https://zenodo.org/record/14239182/files/figure.png" pageId="1" pageNumber="552">Figs 1A1C</figureCitation>
)
</paragraph>
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<paragraph id="613836985226FFE7D8D16369FABCFCBA" blockId="1.[151,1436,831,886]" pageId="1" pageNumber="552">
<taxonomicName id="A6874D1B5226FFE7D8D16369FD46FC9B" ID-CoL="4SNMH" authority="Lukyanchenko 1993, pp. 5 - 6" authorityName="Lukyanchenko" authorityPageNumber="5 - 6" authorityYear="1993" box="[151,672,831,855]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="species" species="arefini">
<emphasis id="53F3EA8A5226FFE7D8D16369FE82FC9B" box="[151,356,831,855]" italics="true" pageId="1" pageNumber="552">Rhyacophila arefini</emphasis>
<bibRefCitation id="05164B695226FFE7D92C6369FDA6FC9B" author="Lukyanchenko, T. I." box="[362,576,831,855]" pageId="1" pageNumber="552" pagination="5 - 6" refId="ref13054" refString="Lukyanchenko, T. I. (1993) A new species of caddisfly of the genus Rhyacophila Pictet (Trichoptera: Rhyacophilidae). Braueria, 20, 5 - 6." type="journal article" year="1993">Lukyanchenko 1993</bibRefCitation>
, pp. 56
</taxonomicName>
, male, female, pupa, larva, South Kuril (Kunashir),
<collectingRegion id="A343F87A5226FFE7DC8F6369FACBFC9B" box="[1225,1325,831,855]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
(Ishikari);
<bibRefCitation id="05164B695226FFE7D8816308FEB6FCBA" author="Arefina, T. I." box="[199,336,862,886]" pageId="1" pageNumber="552" pagination="21 - 44" refId="ref12098" refString="Arefina, T. I. (2001) An outline of females of the genus Rhyacophila (Trichoptera: Rhyacophilidae) from eastern Asia. In: Bae, Y. J. (Ed.), The 21 th Century and Aquatic Entomology in East Asia. Proceedings of the 1 st Symposium of AESEA. Korean Society of Aquatic Entomology, Seoul, pp. 21 - 44." type="book chapter" year="2001">Arefina 2001</bibRefCitation>
, pp. 28, 41, female, South Kuril (Shikotan);
<bibRefCitation id="05164B695226FFE7DB556308FC71FCBA" author="Hattori, T." box="[787,919,862,886]" pageId="1" pageNumber="552" pagination="414 - 437" refId="ref12171" refString="Hattori, T. (2005) Rhyacophilidae. In: Kawai, T. &amp; Tanida, K. (Eds), Aquatic Insects of Japan: Manual with Keys and Illustrations. Tokai University Press, Kanagawa, pp. 414 - 437. [in Japanese]" type="book chapter" year="2005">Hattori 2005</bibRefCitation>
, pp. 427, 434435, male, female,
<collectingRegion id="A343F87A5226FFE7DCA86308FAB3FCBA" box="[1262,1365,862,886]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
.
</paragraph>
<paragraph id="613836985226FFE7D8D163F2FAA3FBBE" blockId="1.[151,1437,932,1787]" pageId="1" pageNumber="552">
<emphasis id="53F3EA8A5226FFE7D8D163F2FE46FC72" bold="true" box="[151,416,932,958]" pageId="1" pageNumber="552">Records in references</emphasis>
.
<materialsCitation id="D1EF3CC55226FFE7D9F463F2FBF4FC72" box="[434,1042,932,959]" location="Ishikari" pageId="1" pageNumber="552" specimenCount="1" stateProvince="Hokkaido">
<bibRefCitation id="05164B695226FFE7D9F463F2FD72FC72" author="Kuhara, N. &amp; Ito, T." box="[434,660,932,959]" pageId="1" pageNumber="552" pagination="18 - 24" refId="ref12868" refString="Kuhara, N. &amp; Ito, T. (1994) Caddisfly fauna of a spring stream, Kutanida Creek, Eniwa, Hokkaido, northern Japan. Biology of Inland Waters, 9, 18 - 24. [in Japanese with English abstract]" type="journal article" year="1994">Kuhara &amp; Ito 1994</bibRefCitation>
, pp. 20, 23,
<collectingRegion id="A343F87A5226FFE7DB6B63F2FC7DFC72" box="[813,923,932,958]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
(
<location id="645860435226FFE7DBE863F2FBECFC72" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DBE863F2FBECFC72" box="[942,1034,932,958]" name="Ishikari" pageId="1" pageNumber="552" stateProvince="Hokkaido">Ishikari</location>
)
</materialsCitation>
;
<materialsCitation id="D1EF3CC55226FFE7DC6663F2FF1BFC2E" collectorName="Vshivkova &amp; Kolin" location="Sakhalin" pageId="1" pageNumber="552" specimenCount="1" stateProvince="Sakhalin">
<collectorName id="CC72534E5226FFE7DC6663F2FB7FFC72" box="[1056,1177,932,958]" pageId="1" pageNumber="552">Vshivkova</collectorName>
&amp;
<collectorName id="CC72534E5226FFE7DC8363F2FAE3FC72" box="[1221,1285,932,958]" pageId="1" pageNumber="552">Kolin</collectorName>
1997, p. 60,
<collectingRegion id="A343F87A5226FFE7D8D1639EFF1BFC2E" box="[151,253,968,994]" country="Russia" name="Sakhalin" pageId="1" pageNumber="552">Sakhalin</collectingRegion>
</materialsCitation>
;
<materialsCitation id="D1EF3CC55226FFE7D940639FFC6EFC2E" box="[262,904,968,995]" location="Nemuro" pageId="1" pageNumber="552" specimenCount="1" stateProvince="Hokkaido">
<bibRefCitation id="05164B695226FFE7D940639FFE42FC2E" author="Ito, T. &amp; Kamei, H. &amp; Ohkawa, A. &amp; Kuhara, N. &amp; Nishimoto, H." box="[262,420,968,995]" pageId="1" pageNumber="552" pagination="1 - 17" refId="ref12365" refString="Ito, T., Kamei, H., Ohkawa, A., Kuhara, N. &amp; Nishimoto, H. (1998) Caddisfly fauna of the Shibetsu District and the Shiretoko Pass, easternmost part of Hokkaido, northern Japan. Biology of Inland Waters, 13, 1 - 17. [in Japanese with English abstract]" type="journal article" year="1998">
Ito
<emphasis id="53F3EA8A5226FFE7D96B639FFEBAFC2E" box="[301,348,968,994]" italics="true" pageId="1" pageNumber="552">et al</emphasis>
. 1998
</bibRefCitation>
, pp. 10, 1214, 16, 17,
<collectingRegion id="A343F87A5226FFE7DAE4639EFCF6FC2E" box="[674,784,968,994]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
(
<location id="645860435226FFE7DB58639EFC66FC2E" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DB58639EFC66FC2E" box="[798,896,968,994]" name="Nemuro" pageId="1" pageNumber="552" stateProvince="Hokkaido">Nemuro</location>
)
</materialsCitation>
;
<materialsCitation id="D1EF3CC55226FFE7DBD4639EFA7EFC2E" box="[914,1432,968,995]" location="Tokachi" pageId="1" pageNumber="552" specimenCount="1" stateProvince="Hokkaido">
<bibRefCitation id="05164B695226FFE7DBD4639EFBC9FC2E" author="Ohkawa, A." box="[914,1071,968,994]" pageId="1" pageNumber="552" pagination="35 - 44" refId="ref13332" refString="Ohkawa, A. (1999) Caddisfly fauna of a spring stream, Kikanko Creek, Tokachi, Hokkaido, northern Japan. Biology of Inland Waters, 14, 35 - 44. [in Japanese with English abstract]" type="journal article" year="1999">Ohkawa 1999</bibRefCitation>
, pp. 4044,
<collectingRegion id="A343F87A5226FFE7DCF2639EFAC4FC2E" box="[1204,1314,968,994]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
(
<location id="645860435226FFE7DD76639EFA77FC2E" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DD76639EFA77FC2E" box="[1328,1425,968,994]" name="Tokachi" pageId="1" pageNumber="552" stateProvince="Hokkaido">Tokachi</location>
)
</materialsCitation>
;
<materialsCitation id="D1EF3CC55226FFE7D8D163BAFCE0FBCA" box="[151,774,1004,1031]" location="Rishiri Island" pageId="1" pageNumber="552" specimenCount="1" stateProvince="Hokkaido">
<bibRefCitation id="05164B695226FFE7D8D163BAFE61FBCA" author="Kuhara, N. &amp; Sato, M." box="[151,391,1004,1030]" pageId="1" pageNumber="552" pagination="63 - 68" refId="ref12915" refString="Kuhara, N. &amp; Sato, M. (2001) Caddisflies of Rishiri Island, northern Japan. Rishiri Studies, 20, 63 - 68. [in Japanese with English abstract]" type="journal article" year="2001">Kuhara &amp; Sato 2001</bibRefCitation>
, p. 64,
<collectingRegion id="A343F87A5226FFE7D99963BAFDABFBCA" box="[479,589,1004,1030]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
(
<location id="645860435226FFE7DA1863BAFD18FBCA" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DA1863BAFD18FBCA" box="[606,766,1004,1030]" name="Rishiri Island" pageId="1" pageNumber="552" stateProvince="Hokkaido">Rishiri Island</location>
)
</materialsCitation>
;
<materialsCitation id="D1EF3CC55226FFE7DB5563BAFA7EFBCA" box="[787,1432,1004,1031]" location="Kamikawa" pageId="1" pageNumber="552" specimenCount="1" stateProvince="Hokkaido">
<bibRefCitation id="05164B695226FFE7DB5563BAFBCAFBCA" author="Ohkawa, A. &amp; Hattori, T." box="[787,1068,1004,1030]" pageId="1" pageNumber="552" pagination="263 - 274" refId="ref13374" refString="Ohkawa, A. &amp; Hattori, T. (2007) The caddisflies (Trichoptera) of the University Forest in Hokkaido, the University of Tokyo. I. Rhyacophilidae. In: Bueno-Soria, J., Barba-Alvarez, R. &amp; Armitage, B. (Eds.), Proceedings of the 12 th International Symposium on Trichoptera. The Caddis Press, Columbus, Ohio, pp. 263 - 274." type="book chapter" year="2007">Ohkawa &amp; Hattori 2007</bibRefCitation>
, p. 266,
<collectingRegion id="A343F87A5226FFE7DCD763BAFB19FBCA" box="[1169,1279,1004,1030]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
(
<location id="645860435226FFE7DD5663BAFA76FBCA" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DD5663BAFA76FBCA" box="[1296,1424,1004,1030]" name="Kamikawa" pageId="1" pageNumber="552" stateProvince="Hokkaido">Kamikawa</location>
)
</materialsCitation>
;
<materialsCitation id="D1EF3CC55226FFE7D8D16447FD73FBE6" box="[151,661,1040,1067]" location="Okhotsk" pageId="1" pageNumber="552" specimenCount="1" stateProvince="Hokkaido">
<bibRefCitation id="05164B695226FFE7D8D16447FEBAFBE6" author="Muramatsu, H." box="[151,348,1040,1067]" pageId="1" pageNumber="552" pagination="1 - 12" refId="ref13210" refString="Muramatsu, H. (2008) Abashiri-shicho kannai-de saishusareta tobikera-moku no konchu (III) [Caddisflies (Trichoptera) collected in Abashiri district (III)]. Okhotsk no Konchu, 5, 1 - 12, tabs. 1 - 4. [in Japanese]" type="journal article" year="2008">Muramatsu 2008</bibRefCitation>
, p. 3,
<collectingRegion id="A343F87A5226FFE7D9E16446FDF3FBE6" box="[423,533,1040,1066]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
(
<location id="645860435226FFE7DA616446FD6BFBE6" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DA616446FD6BFBE6" box="[551,653,1040,1066]" name="Okhotsk" pageId="1" pageNumber="552" stateProvince="Hokkaido">Okhotsk</location>
)
</materialsCitation>
;
<materialsCitation id="D1EF3CC55226FFE7DAE56447FB64FBE6" box="[675,1154,1040,1067]" location="Okhotsk" pageId="1" pageNumber="552" specimenCount="1" stateProvince="Hokkaido">
<bibRefCitation id="05164B695226FFE7DAE56447FCAFFBE6" author="Ito, T. &amp; Nakatani, M. &amp; Matsumoto, K." box="[675,841,1040,1067]" pageId="1" pageNumber="552" pagination="5 - 10" refId="ref12483" refString="Ito, T., Nakatani, M. &amp; Matsumoto, K. (2009) Trichoptera fauna of Kamenoko-ike, Kiyosato-cho, Hokkaido, northern Japan. Sylvicola, 27, 5 - 10. [in Japanese with English abstract]" type="journal article" year="2009">
Ito
<emphasis id="53F3EA8A5226FFE7DA8F6447FD1BFBE6" box="[713,765,1040,1066]" italics="true" pageId="1" pageNumber="552">et al</emphasis>
. 2009
</bibRefCitation>
, p. 8,
<collectingRegion id="A343F87A5226FFE7DBD26446FBE4FBE6" box="[916,1026,1040,1066]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
(
<location id="645860435226FFE7DC526446FB9CFBE6" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DC526446FB9CFBE6" box="[1044,1146,1040,1066]" name="Okhotsk" pageId="1" pageNumber="552" stateProvince="Hokkaido">Okhotsk</location>
)
</materialsCitation>
;
<materialsCitation id="D1EF3CC55226FFE7DCD66447FE66FB82" location="Tokachi" pageId="1" pageNumber="552" specimenCount="1" stateProvince="Hokkaido">
<bibRefCitation id="05164B695226FFE7DCD66447FABDFBE6" author="Ito, T. &amp; Kubo, E." box="[1168,1371,1040,1067]" pageId="1" pageNumber="552" pagination="1 - 12" refId="ref12279" refString="Ito, T. &amp; Kubo, E. (2011) Caddisflies (Trichoptera) of Toyoni-gawa River system, Tokachi, Hokkaido, northern Japan. Sylvicola, 29, 1 - 12. [in Japanese with English abstract]" type="journal article" year="2011">Ito &amp; Kubo 2011</bibRefCitation>
, p. 4,
<collectingRegion id="A343F87A5226FFE7D8D16462FEE3FB82" box="[151,261,1076,1102]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
(
<location id="645860435226FFE7D9516462FE9EFB82" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7D9516462FE9EFB82" box="[279,376,1076,1102]" name="Tokachi" pageId="1" pageNumber="552" stateProvince="Hokkaido">Tokachi</location>
)
</materialsCitation>
;
<materialsCitation id="D1EF3CC55226FFE7D9CB6462FBCCFB82" box="[397,1066,1076,1103]" location="Hidaka" municipality="Sorachi" pageId="1" pageNumber="552" specimenCount="1" stateProvince="Hokkaido">
<bibRefCitation id="05164B695226FFE7D9CB6462FDC5FB82" author="Kuhara, N." box="[397,547,1076,1102]" pageId="1" pageNumber="552" pagination="47 - 76" refId="ref12824" refString="Kuhara, N. (2011) Species composition and flight periods of caddisflies (Trichoptera) at headwater streams in Hokkaido, northern Japan. Biology of Inland Waters, 26, 47 - 76. [in Japanese with English abstract]" type="journal article" year="2011">Kuhara 2011</bibRefCitation>
, pp. 52, 57, 61,
<collectingRegion id="A343F87A5226FFE7DAA26462FCB4FB82" box="[740,850,1076,1102]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
(
<collectingMunicipality id="815CACE25226FFE7DB226462FC58FB82" box="[868,958,1076,1102]" pageId="1" pageNumber="552">Sorachi</collectingMunicipality>
,
<location id="645860435226FFE7DB8A6462FBC4FB82" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DB8A6462FBC4FB82" box="[972,1058,1076,1102]" municipality="Sorachi" name="Hidaka" pageId="1" pageNumber="552" stateProvince="Hokkaido">Hidaka</location>
)
</materialsCitation>
;
<materialsCitation id="D1EF3CC55226FFE7DC7E6462FE64FBBE" location="Okhotsk" pageId="1" pageNumber="552" specimenCount="1" stateProvince="Hokkaido">
<bibRefCitation id="05164B695226FFE7DC7E6462FAABFB82" author="Iijima, K. &amp; Tsuchiya, K." box="[1080,1357,1076,1102]" pageId="1" pageNumber="552" pagination="27 - 35" refId="ref12239" refString="Iijima, K. &amp; Tsuchiya, K. (2014) Catalogue of caddisflies (Trichoptera) collected by Kazuo IIJIMA. Memoirs of the Kushiro City Museum, 35, 27 - 35. [in Japanese]" type="journal article" year="2014">Iijima &amp; Tsuchiya 2014</bibRefCitation>
, p. 27,
<collectingRegion id="A343F87A5226FFE7D8D1640EFEE3FBBE" box="[151,261,1112,1138]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
(
<location id="645860435226FFE7D952640EFE9CFBBE" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7D952640EFE9CFBBE" box="[276,378,1112,1138]" name="Okhotsk" pageId="1" pageNumber="552" stateProvince="Hokkaido">Okhotsk</location>
)
</materialsCitation>
;
<materialsCitation id="D1EF3CC55226FFE7D9CB640FFBEEFBBE" box="[397,1032,1112,1139]" location="Kamikawa" municipality="Soya" pageId="1" pageNumber="552" specimenCount="1" stateProvince="Hokkaido">
<bibRefCitation id="05164B695226FFE7D9CB640FFD8DFBBE" author="Ito, T. &amp; Kuhara, N." box="[397,619,1112,1139]" pageId="1" pageNumber="552" pagination="51 - 88" refId="ref12322" refString="Ito, T. &amp; Kuhara, N. (2023) Caddisflies (Trichoptera) of northern part of Hokkaido, northernmost Japan. Biology of Inland Waters, 38, 51 - 88. [in Japanese with English abstract]" type="journal article" year="2023">Ito &amp; Kuhara, 2023</bibRefCitation>
, p. 63,
<collectingRegion id="A343F87A5226FFE7DAFA640EFCCCFBBE" box="[700,810,1112,1138]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
(
<collectingMunicipality id="815CACE25226FFE7DB7C640EFC90FBBE" box="[826,886,1112,1138]" pageId="1" pageNumber="552">Soya</collectingMunicipality>
,
<location id="645860435226FFE7DBC7640EFBE6FBBE" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DBC7640EFBE6FBBE" box="[897,1024,1112,1138]" municipality="Soya" name="Kamikawa" pageId="1" pageNumber="552" stateProvince="Hokkaido">Kamikawa</location>
)
</materialsCitation>
;
<materialsCitation id="D1EF3CC55226FFE7DC54640FFAA4FBBE" box="[1042,1346,1112,1139]" collectorName="Ito" location="Sakhalin" pageId="1" pageNumber="552" specimenCount="1" stateProvince="Sakhalin">
<collectorName id="CC72534E5226FFE7DC54640FFBD7FBBF" box="[1042,1073,1113,1139]" pageId="1" pageNumber="552">Ito</collectorName>
<emphasis id="53F3EA8A5226FFE7DC7E640FFB8CFBBE" box="[1080,1130,1112,1138]" italics="true" pageId="1" pageNumber="552">et al</emphasis>
. in press,
<collectingRegion id="A343F87A5226FFE7DC9A640EFAA4FBBE" box="[1244,1346,1112,1138]" country="Russia" name="Sakhalin" pageId="1" pageNumber="552">Sakhalin</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="299D65135226FFE7D881642AFF2DF9EE" pageId="1" pageNumber="552" type="materials_examined">
<paragraph id="613836985226FFE7D881642AFF2DF9EE" blockId="1.[151,1437,932,1787]" pageId="1" pageNumber="552">
<emphasis id="53F3EA8A5226FFE7D881642AFE4FFB5A" bold="true" box="[199,425,1148,1174]" pageId="1" pageNumber="552">Material examined</emphasis>
.
<materialsCitation id="D1EF3CC55226FFE7D9F1642AFBFFFAEB" collectingDate="1977-07-23" collectingDateMax="2016-07-12" collectingDateMin="1977-07-23" collectionCode="TI" collectorName="Y. Nagasaka" location="Akabira-shi" municipality="Sorachi" pageId="1" pageNumber="552" specimenCount="119" specimenCount-female="39" specimenCount-male="80" stateProvince="Hokkaido">
<emphasis id="53F3EA8A5226FFE7D9F1642AFD7EFB5A" bold="true" box="[439,664,1148,1174]" pageId="1" pageNumber="552">
<collectingRegion id="A343F87A5226FFE7D9F1642AFDC9FB5A" box="[439,559,1148,1174]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
,
<collectingMunicipality id="815CACE25226FFE7DA7A642AFD7EFB5A" box="[572,664,1148,1174]" pageId="1" pageNumber="552">Sorachi</collectingMunicipality>
</emphasis>
:
<location id="645860435226FFE7DAE3642AFCCAFB5A" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DAE3642AFCCAFB5A" box="[677,812,1148,1174]" municipality="Sorachi" name="Akabira-shi" pageId="1" pageNumber="552" stateProvince="Hokkaido">Akabira-shi</location>
, Penke-shubushinai-zawa,
<specimenCount id="7781FD115226FFE7DC1D642AFB24FB5A" box="[1115,1218,1148,1174]" count="67" pageId="1" pageNumber="552" type="male">67 males</specimenCount>
,
<specimenCount id="7781FD115226FFE7DC8B642AFAACFB5A" box="[1229,1354,1148,1174]" count="38" pageId="1" pageNumber="552" type="female">38 females</specimenCount>
,
<date id="153910585226FFE7DD13642AFEFFFB76" pageId="1" pageNumber="552" value="2016-06-29" valueMax="2016-07-12" valueMin="2016-06-29">
<collectingDate id="057DE9B05226FFE7DD13642AFEFFFB76" pageId="1" pageNumber="552" value="2016-06-29" valueMax="2016-07-12" valueMin="2016-06-29">29.vi 12.vii.2016</collectingDate>
</date>
, leg.
<collectorName id="CC72534E5226FFE7D91164F7FE02FB76" box="[343,484,1184,1211]" pageId="1" pageNumber="552">Y. Nagasaka</collectorName>
<emphasis id="53F3EA8A5226FFE7D9AA64F7FDF9FB76" box="[492,543,1184,1210]" italics="true" pageId="1" pageNumber="552">et al</emphasis>
., M (
<collectionCode id="0796AE5D5226FFE7DA1964F6FD98FB76" box="[607,638,1184,1210]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
). Ashibetsu-shi, Nakanosawa-gawa,
<specimenCount id="7781FD115226FFE7DC5B64F6FB9EFB76" box="[1053,1144,1184,1210]" count="9" pageId="1" pageNumber="552" type="male">9 males</specimenCount>
,
<date id="153910585226FFE7DCC264F6FAA1FB76" box="[1156,1351,1184,1210]" pageId="1" pageNumber="552" value="2013-06-13" valueMax="2013-08-08" valueMin="2013-06-13">
<collectingDate id="057DE9B05226FFE7DCC264F6FAA1FB76" box="[1156,1351,1184,1210]" pageId="1" pageNumber="552" value="2013-06-13" valueMax="2013-08-08" valueMin="2013-06-13">13.vi8.viii.2013</collectingDate>
</date>
, leg.
<collectorName id="CC72534E5226FFE7DDC364F7FEE2FB12" pageId="1" pageNumber="552">Y. Nagasaka</collectorName>
<emphasis id="53F3EA8A5226FFE7D9486493FEA4FB12" box="[270,322,1220,1246]" italics="true" pageId="1" pageNumber="552">et al</emphasis>
., M (
<collectionCode id="0796AE5D5226FFE7D9C36492FE42FB12" box="[389,420,1220,1246]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
). Ashibetsu-shi, Nisen-zawa,
<specimenCount id="7781FD115226FFE7DABF6492FCACFB12" box="[761,842,1220,1246]" count="1" pageId="1" pageNumber="552" type="male">1 male</specimenCount>
,
<date id="153910585226FFE7DB1E6492FBC6FB12" box="[856,1056,1220,1246]" pageId="1" pageNumber="552" value="2013-06-28" valueMax="2013-07-12" valueMin="2013-06-28">
<collectingDate id="057DE9B05226FFE7DB1E6492FBC6FB12" box="[856,1056,1220,1246]" pageId="1" pageNumber="552" value="2013-06-28" valueMax="2013-07-12" valueMin="2013-06-28">28.vi12.vii.2013</collectingDate>
</date>
, leg.
<collectorName id="CC72534E5226FFE7DC276493FB16FB12" box="[1121,1264,1220,1247]" pageId="1" pageNumber="552">Y. Nagasaka</collectorName>
<emphasis id="53F3EA8A5226FFE7DCBC6493FAC8FB12" box="[1274,1326,1220,1246]" italics="true" pageId="1" pageNumber="552">et al</emphasis>
., M (
<collectionCode id="0796AE5D5226FFE7DD376492FA76FB12" box="[1393,1424,1220,1246]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
). Ashibetsu-shi, Yoshida-no-sawa,
<specimenCount id="7781FD115226FFE7DA5464BEFD88FACE" box="[530,622,1256,1282]" count="2" pageId="1" pageNumber="552" type="male">2 males</specimenCount>
,
<specimenCount id="7781FD115226FFE7DA3A64BEFD04FACE" box="[636,738,1256,1282]" count="1" pageId="1" pageNumber="552" type="female">1 female</specimenCount>
,
<date id="153910585226FFE7DAB664BEFC55FACE" box="[752,947,1256,1282]" pageId="1" pageNumber="552" value="2013-06-07" valueMax="2013-08-23" valueMin="2013-06-07">
<collectingDate id="057DE9B05226FFE7DAB664BEFC55FACE" box="[752,947,1256,1282]" pageId="1" pageNumber="552" value="2013-06-07" valueMax="2013-08-23" valueMin="2013-06-07">7.vi23.viii.2013</collectingDate>
</date>
, leg.
<collectorName id="CC72534E5226FFE7DBB564BFFB67FACE" box="[1011,1153,1256,1283]" pageId="1" pageNumber="552">Y. Nagasaka</collectorName>
<emphasis id="53F3EA8A5226FFE7DCCC64BFFB58FACE" box="[1162,1214,1256,1282]" italics="true" pageId="1" pageNumber="552">et al</emphasis>
., M (
<collectionCode id="0796AE5D5226FFE7DD4764BEFAC6FACE" box="[1281,1312,1256,1282]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
).
<emphasis id="53F3EA8A5226FFE7DD7064BEFA73FACE" bold="true" box="[1334,1429,1256,1282]" pageId="1" pageNumber="552">Ishikari</emphasis>
: Sapporo-shi, Takino, Masuno-sawa, small tributary,
<specimenCount id="7781FD115226FFE7DAA5655AFCD4FAEA" box="[739,818,1292,1318]" count="1" pageId="1" pageNumber="552" type="male">1 male</specimenCount>
,
<date id="153910585226FFE7DB7B655AFC59FAEA" box="[829,959,1292,1318]" pageId="1" pageNumber="552" value="1977-07-23">
<collectingDate id="057DE9B05226FFE7DB7B655AFC59FAEA" box="[829,959,1292,1318]" pageId="1" pageNumber="552" value="1977-07-23">23.vii.1977</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5226FFE7DBBA655BFBFFFAEB" box="[1020,1049,1293,1319]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55226FFE7DC62655AFF05FA87" collectingDate="1981-06-06" collectingDateMax="1981-07-18" collectingDateMin="1981-06-06" collectionCode="TI" pageId="1" pageNumber="552" specimenCount="3" specimenCount-male="3" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5226FFE7DC02655AFB85FAEA" box="[1092,1123,1292,1318]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
);
<specimenCount id="7781FD115226FFE7DC3E655AFB37FAEA" box="[1144,1233,1292,1318]" count="3" pageId="1" pageNumber="552" type="male">3 males</specimenCount>
,
<date id="153910585226FFE7DC9B655AFA7EFAEA" box="[1245,1432,1292,1318]" pageId="1" pageNumber="552" value="1981-06-06" valueMax="1981-07-18" valueMin="1981-06-06">
<collectingDate id="057DE9B05226FFE7DC9B655AFA7EFAEA" box="[1245,1432,1292,1318]" pageId="1" pageNumber="552" value="1981-06-06" valueMax="1981-07-18" valueMin="1981-06-06">6.vi18.vii.1981</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5226FFE7D8806567FF05FA87" box="[198,227,1329,1355]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55226FFE7D8AB6566FD82FA87" box="[237,612,1328,1355]" collectingDate="1984-06-22" collectionCode="TI" pageId="1" pageNumber="552" specimenCount="1" specimenCount-male="1" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5226FFE7D94D6566FECCFA86" box="[267,298,1328,1354]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
);
<specimenCount id="7781FD115226FFE7D97B6566FE6CFA86" box="[317,394,1328,1354]" count="1" pageId="1" pageNumber="552" type="male">1 male</specimenCount>
,
<date id="153910585226FFE7D9D26566FDE9FA86" box="[404,527,1328,1354]" pageId="1" pageNumber="552" value="1984-06-22">
<collectingDate id="057DE9B05226FFE7D9D26566FDE9FA86" box="[404,527,1328,1354]" pageId="1" pageNumber="552" value="1984-06-22">22.vi.1984</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5226FFE7DA016567FD82FA87" box="[583,612,1329,1355]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55226FFE7DA296566FC08FA87" box="[623,1006,1328,1355]" collectingDate="1986-07-18" collectionCode="TI" pageId="1" pageNumber="552" specimenCount="1" specimenCount-male="1" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5226FFE7DACB6566FD4AFA86" box="[653,684,1328,1354]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
);
<specimenCount id="7781FD115226FFE7DAF96566FCEAFA86" box="[703,780,1328,1354]" count="1" pageId="1" pageNumber="552" type="male">1 male</specimenCount>
,
<date id="153910585226FFE7DB506566FC7EFA86" box="[790,920,1328,1354]" pageId="1" pageNumber="552" value="1986-07-18">
<collectingDate id="057DE9B05226FFE7DB506566FC7EFA86" box="[790,920,1328,1354]" pageId="1" pageNumber="552" value="1986-07-18">18.vii.1986</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5226FFE7DB976567FC08FA87" box="[977,1006,1329,1355]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55226FFE7DBBE6566FD93FAA3" collectingDate="1982-08-08" collectionCode="TI" county="Sapporo-shi" elevation="1200" location="Yoichidake" municipality="Jozankei" pageId="1" pageNumber="552" specimenCount="1" specimenCount-male="1" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5226FFE7DC506566FBD3FA86" box="[1046,1077,1328,1354]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
).
<collectingCounty id="88594E145226FFE7DC0E6566FB34FA86" box="[1096,1234,1328,1354]" pageId="1" pageNumber="552">Sapporo-shi</collectingCounty>
,
<collectingMunicipality id="815CACE25226FFE7DC9A6566FAA7FA86" box="[1244,1345,1328,1354]" pageId="1" pageNumber="552">Jôzankei</collectingMunicipality>
,
<location id="645860435226FFE7DD0C6566FF29FAA2" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DD0C6566FF29FAA2" county="Sapporo-shi" municipality="Jozankei" name="Yoichidake" pageId="1" pageNumber="552" stateProvince="Hokkaido">Yoichidake</location>
,
<quantity id="A67F9B7D5226FFE7D89D6502FEDFFAA3" box="[219,313,1364,1391]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="552" unit="m" value="1200.0">
<elevation id="EAAAD1AB5226FFE7D89D6502FEDFFAA3" box="[219,313,1364,1391]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="552" unit="m" value="1200.0">1,200 m</elevation>
</quantity>
,
<specimenCount id="7781FD115226FFE7D9006502FE72FAA2" box="[326,404,1364,1390]" count="1" pageId="1" pageNumber="552" type="male">1 male</specimenCount>
,
<date id="153910585226FFE7D9E66502FDFAFAA2" box="[416,540,1364,1390]" pageId="1" pageNumber="552" value="1982-08-08">
<collectingDate id="057DE9B05226FFE7D9E66502FDFAFAA2" box="[416,540,1364,1390]" pageId="1" pageNumber="552" value="1982-08-08">8.viii.1982</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5226FFE7DA1E6503FD93FAA3" box="[600,629,1365,1391]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55226FFE7DAC46502FE8EFA5F" collectingDate="2013-06-05" collectingDateMax="2013-06-10" collectingDateMin="2013-06-05" collectionCode="TI" county="Eniwa-shi" location="Saiwai-bashi" municipality="Moizari-gawa" pageId="1" pageNumber="552" specimenCount="12" specimenCount-female="7" specimenCount-male="5" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5226FFE7DAE76502FD26FAA2" box="[673,704,1364,1390]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
).
<collectingCounty id="88594E145226FFE7DA926502FCAEFAA2" box="[724,840,1364,1390]" pageId="1" pageNumber="552">Eniwa-shi</collectingCounty>
,
<collectingMunicipality id="815CACE25226FFE7DB136502FC13FAA2" box="[853,1013,1364,1390]" pageId="1" pageNumber="552">Moizari-gawa</collectingMunicipality>
,
<location id="645860435226FFE7DC466502FB72FAA2" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DC466502FB72FAA2" box="[1024,1172,1364,1390]" county="Eniwa-shi" municipality="Moizari-gawa" name="Saiwai-bashi" pageId="1" pageNumber="552" stateProvince="Hokkaido">Saiwai-bashi</location>
,
<specimenCount id="7781FD115226FFE7DCD96502FB1FFAA2" box="[1183,1273,1364,1391]" count="5" pageId="1" pageNumber="552" type="male">5 males</specimenCount>
,
<specimenCount id="7781FD115226FFE7DD436503FA93FAA2" box="[1285,1397,1364,1391]" count="7" pageId="1" pageNumber="552" type="female">7 females</specimenCount>
,
<date id="153910585226FFE7DDC66502FEF4FA5E" pageId="1" pageNumber="552" value="2013-06-05" valueMax="2013-06-10" valueMin="2013-06-05">
<collectingDate id="057DE9B05226FFE7DDC66502FEF4FA5E" pageId="1" pageNumber="552" value="2013-06-05" valueMax="2013-06-10" valueMin="2013-06-05">5 10.vi.2013</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5226FFE7D90D652FFE8EFA5F" box="[331,360,1401,1427]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55226FFE7D934652FFF04FA7B" collectingDate="1996-05-28" collectionCode="TI" county="Eniwa-shi" location="Fishery Experimental Station" municipality="Kita-kashiwagi" pageId="1" pageNumber="552" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Hokkaido">
L (
<collectionCode id="0796AE5D5226FFE7D9D7652EFE56FA5E" box="[401,432,1400,1426]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
).
<collectingCounty id="88594E145226FFE7D985652EFDD1FA5E" box="[451,567,1400,1426]" pageId="1" pageNumber="552">Eniwa-shi</collectingCounty>
,
<collectingMunicipality id="815CACE25226FFE7DA07652EFD09FA5E" box="[577,751,1400,1426]" pageId="1" pageNumber="552">Kita-kashiwagi</collectingMunicipality>
,
<location id="645860435226FFE7DABF652EFBA4FA5E" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DABF652EFBA4FA5E" box="[761,1090,1400,1426]" county="Eniwa-shi" municipality="Kita-kashiwagi" name="Fishery Experimental Station" pageId="1" pageNumber="552" stateProvince="Hokkaido">Fishery Experimental Station</location>
,
<specimenCount id="7781FD115226FFE7DC0A652EFB43FA5E" box="[1100,1189,1400,1426]" count="2" pageId="1" pageNumber="552" type="male">2 males</specimenCount>
,
<specimenCount id="7781FD115226FFE7DCE9652EFAFBFA5E" box="[1199,1309,1400,1426]" count="2" pageId="1" pageNumber="552" type="female">2 females</specimenCount>
,
<date id="153910585226FFE7DD61652EFA7EFA5E" box="[1319,1432,1400,1426]" pageId="1" pageNumber="552" value="1996-05-28">
<collectingDate id="057DE9B05226FFE7DD61652EFA7EFA5E" box="[1319,1432,1400,1426]" pageId="1" pageNumber="552" value="1996-05-28">28.v.1996</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5226FFE7D88365CBFF04FA7B" box="[197,226,1437,1463]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55226FFE7D8AA65CAFC17FA7B" box="[236,1009,1436,1463]" collectingDate="1996-07-19" collectionCode="TI" location="Eniwa-koen" municipality="Eniwa-shi" pageId="1" pageNumber="552" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5226FFE7D94C65CAFECFFA7A" box="[266,297,1436,1462]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
).
<collectingMunicipality id="815CACE25226FFE7D97D65CAFE49FA7A" box="[315,431,1436,1462]" pageId="1" pageNumber="552">Eniwa-shi</collectingMunicipality>
,
<location id="645860435226FFE7D9FF65CAFDA5FA7A" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7D9FF65CAFDA5FA7A" box="[441,579,1436,1462]" municipality="Eniwa-shi" name="Eniwa-koen" pageId="1" pageNumber="552" stateProvince="Hokkaido">Eniwa-kôen</location>
,
<specimenCount id="7781FD115226FFE7DA0A65CAFD42FA7A" box="[588,676,1436,1462]" count="2" pageId="1" pageNumber="552" type="male">2 males</specimenCount>
,
<specimenCount id="7781FD115226FFE7DAE865CAFCF6FA7A" box="[686,784,1436,1462]" count="1" pageId="1" pageNumber="552" type="female">1 female</specimenCount>
,
<date id="153910585226FFE7DB5C65CAFC7AFA7A" box="[794,924,1436,1462]" pageId="1" pageNumber="552" value="1996-07-19">
<collectingDate id="057DE9B05226FFE7DB5C65CAFC7AFA7A" box="[794,924,1436,1462]" pageId="1" pageNumber="552" value="1996-07-19">19.vii.1996</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5226FFE7DB9265CBFC17FA7B" box="[980,1009,1437,1463]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55226FFE7DBBD65CAFD2BFA17" collectingDate="1980-06-05" collectionCode="TI" location="Rankoshi" municipality="Chitose-shi" pageId="1" pageNumber="552" specimenCount="1" specimenCount-male="1" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5226FFE7DC5E65CAFBD1FA7A" box="[1048,1079,1436,1462]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
).
<collectingMunicipality id="815CACE25226FFE7DC0F65CAFB2DFA7A" box="[1097,1227,1436,1462]" pageId="1" pageNumber="552">Chitose-shi</collectingMunicipality>
,
<location id="645860435226FFE7DC9365CAFAA4FA7A" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DC9365CAFAA4FA7A" box="[1237,1346,1436,1462]" municipality="Chitose-shi" name="Rankoshi" pageId="1" pageNumber="552" stateProvince="Hokkaido">Rankoshi</location>
, salmon hatchery, sprimg stream,
<specimenCount id="7781FD115226FFE7D9F66596FE18FA16" box="[432,510,1472,1498]" count="1" pageId="1" pageNumber="552" type="male">1 male</specimenCount>
,
<date id="153910585226FFE7DA4F6596FD93FA17" box="[521,629,1472,1499]" pageId="1" pageNumber="552" value="1980-06-05">
<collectingDate id="057DE9B05226FFE7DA4F6596FD93FA17" box="[521,629,1472,1499]" pageId="1" pageNumber="552" value="1980-06-05">5.vi.1980</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5226FFE7DAF66597FD2BFA17" box="[688,717,1473,1499]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55226FFE7DA9E6596FDAFFA33" collectingDate="1980-06-05" collectionCode="TI" county="Chitose-shi" location="Fukajo-bashi" municipality="Rankoshi" pageId="1" pageNumber="552" specimenCount="9" specimenCount-female="4" specimenCount-male="5" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5226FFE7DAB16596FCF0FA16" box="[759,790,1472,1498]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
).
<collectingCounty id="88594E145226FFE7DB6F6596FC4DFA16" box="[809,939,1472,1498]" pageId="1" pageNumber="552">Chitose-shi</collectingCounty>
,
<collectingMunicipality id="815CACE25226FFE7DBF16596FBC2FA16" box="[951,1060,1472,1498]" pageId="1" pageNumber="552">Rankoshi</collectingMunicipality>
, spring stream near
<location id="645860435226FFE7DD456596FA7EFA16" LSID="urn:lsid:plazi:treatment:E92E878E5226FFE7D8D16280FD75F936:645860435226FFE7DD456596FA7EFA16" box="[1283,1432,1472,1498]" county="Chitose-shi" municipality="Rankoshi" name="Fukajo-bashi" pageId="1" pageNumber="552" stateProvince="Hokkaido">Fukajô-bashi</location>
,
<specimenCount id="7781FD115226FFE7D8D165B2FF15FA32" box="[151,243,1508,1535]" count="5" pageId="1" pageNumber="552" type="male">5 males</specimenCount>
,
<specimenCount id="7781FD115226FFE7D94765B2FE94FA32" box="[257,370,1508,1534]" count="4" pageId="1" pageNumber="552" type="female">4 females</specimenCount>
,
<date id="153910585226FFE7D9C665B2FE0AFA33" box="[384,492,1508,1535]" pageId="1" pageNumber="552" value="1980-06-05">
<collectingDate id="057DE9B05226FFE7D9C665B2FE0AFA33" box="[384,492,1508,1535]" pageId="1" pageNumber="552" value="1980-06-05">5.vi.1980</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5226FFE7DA6A65B3FDAFFA33" box="[556,585,1509,1535]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55226FFE7DA1165B2FC0FFA33" box="[599,1001,1508,1535]" collectingDate="2004-07-09" collectionCode="TI" pageId="1" pageNumber="552" specimenCount="2" specimenCount-male="2" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5226FFE7DA3F65B2FD7EFA32" box="[633,664,1508,1534]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
);
<specimenCount id="7781FD115226FFE7DAE965B2FCECFA32" box="[687,778,1508,1534]" count="2" pageId="1" pageNumber="552" type="male">2 males</specimenCount>
,
<date id="153910585226FFE7DB5E65B2FC6AFA32" box="[792,908,1508,1534]" pageId="1" pageNumber="552" value="2004-07-09">
<collectingDate id="057DE9B05226FFE7DB5E65B2FC6AFA32" box="[792,908,1508,1534]" pageId="1" pageNumber="552" value="2004-07-09">9.vii.2004</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5226FFE7DB8A65B3FC0FFA33" box="[972,1001,1509,1535]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
</materialsCitation>
,
<materialsCitation id="D1EF3CC55226FFE7DBB165B2FA99FA33" box="[1015,1407,1508,1535]" collectingDate="2017-07-06" collectionCode="TI" pageId="1" pageNumber="552" specimenCount="1" specimenCount-male="1" stateProvince="Hokkaido">
S (
<collectionCode id="0796AE5D5226FFE7DC5F65B2FBDEFA32" box="[1049,1080,1508,1534]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
);
<specimenCount id="7781FD115226FFE7DC0965B2FB79FA32" box="[1103,1183,1508,1534]" count="1" pageId="1" pageNumber="552" type="male">1 male</specimenCount>
,
<date id="153910585226FFE7DCEB65B2FAC7FA32" box="[1197,1313,1508,1534]" pageId="1" pageNumber="552" value="2017-07-06">
<collectingDate id="057DE9B05226FFE7DCEB65B2FAC7FA32" box="[1197,1313,1508,1534]" pageId="1" pageNumber="552" value="2017-07-06">6.vii.2017</collectingDate>
</date>
, leg.
<collectionCode id="0796AE5D5226FFE7DD2465B3FA99FA33" box="[1378,1407,1509,1535]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
</materialsCitation>
, S (
<collectionCode id="0796AE5D5226FFE7D8E6665EFF59F9EE" box="[160,191,1544,1570]" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="1" pageNumber="552">TI</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="299D65135226FFE7D881667AFD75F936" pageId="1" pageNumber="552" type="distribution">
<paragraph id="613836985226FFE7D881667AFEE7F9A6" blockId="1.[151,1437,932,1787]" pageId="1" pageNumber="552">
<emphasis id="53F3EA8A5226FFE7D881667AFEBFF98A" bold="true" box="[199,345,1580,1606]" pageId="1" pageNumber="552">Distribution</emphasis>
(
<figureCitation id="F9BC2A1D5226FFE7D936667AFDE4F98A" box="[368,514,1580,1606]" captionStart="FIGURE 1" captionStartId="2.[151,250,1655,1679]" captionTargetBox="[165,1422,186,1625]" captionTargetId="figure-29@2.[151,1436,179,1630]" captionTargetPageId="2" captionText="FIGURE 1. Recorded collection sites of Rhyacophila species. 1A, location of Hokkaido and recorded sites of known species discussed in present study. 1B1E, recorded and collected sites in Hokkaido: 1B, 1C, R. eniwa sp. nov. and R. arefini Lukyanchenko; 1D, R. bifue sp. nov., R. imakane sp. nov., R. misago sp. nov., and R. hamata Hattori &amp; Ohkawa; 1E, R. shimamaki sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14239182" httpUri="https://zenodo.org/record/14239182/files/figure.png" pageId="1" pageNumber="552">Figs 1A1C</figureCitation>
, black circles).
<collectingRegion id="A343F87A5226FFE7DA8F667AFCD1F98A" box="[713,823,1580,1606]" country="Japan" name="Hokkaido" pageId="1" pageNumber="552">Hokkaido</collectingRegion>
(northern, eastern and central areas), South Kuril,
<collectingRegion id="A343F87A5226FFE7D8D16606FF1BF9A6" box="[151,253,1616,1642]" country="Russia" name="Sakhalin" pageId="1" pageNumber="552">Sakhalin</collectingRegion>
.
</paragraph>
<paragraph id="613836985226FFE7D8816622FE5BF97E" blockId="1.[151,1437,932,1787]" pageId="1" pageNumber="552">
<emphasis id="53F3EA8A5226FFE7D8816622FEC5F942" bold="true" box="[199,291,1652,1678]" pageId="1" pageNumber="552">Habitat</emphasis>
. The adults of the species were collected in mountain areas near springs, waterfalls, and small streams up to several meters wide.
</paragraph>
<paragraph id="613836985226FFE7D88166EAFA89F91A" blockId="1.[151,1437,932,1787]" box="[199,1391,1724,1751]" pageId="1" pageNumber="552">
<emphasis id="53F3EA8A5226FFE7D88166EAFE8AF91A" bold="true" box="[199,364,1724,1750]" pageId="1" pageNumber="552">Species group</emphasis>
. This species was placed in the
<taxonomicName id="A6874D1B5226FFE7DA9566EAFC58F91A" authorityName="McLachlan" authorityYear="1879" box="[723,958,1724,1750]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="552" phylum="Arthropoda" rank="species" species="sibirica">
<emphasis id="53F3EA8A5226FFE7DA9566EAFC58F91A" box="[723,958,1724,1750]" italics="true" pageId="1" pageNumber="552">Rhyacophila sibirica</emphasis>
</taxonomicName>
Species Group (
<bibRefCitation id="05164B695226FFE7DC3166EAFA84F91A" author="Lukyanchenko, T. I." box="[1143,1378,1724,1750]" pageId="1" pageNumber="552" pagination="5 - 6" refId="ref13054" refString="Lukyanchenko, T. I. (1993) A new species of caddisfly of the genus Rhyacophila Pictet (Trichoptera: Rhyacophilidae). Braueria, 20, 5 - 6." type="journal article" year="1993">Lukyanchenko 1993</bibRefCitation>
).
</paragraph>
<paragraph id="613836985226FFE7D88166B6FD75F936" blockId="1.[151,1437,932,1787]" box="[199,659,1760,1787]" pageId="1" pageNumber="552">
<emphasis id="53F3EA8A5226FFE7D88166B6FE9AF936" bold="true" box="[199,380,1760,1786]" pageId="1" pageNumber="552">Japanese name</emphasis>
. Arefin-nagare-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>