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<document id="84EF82CDCCC377D7ECB113D42CFA1B22" ENCODING="UTF8" ID-DOI="10.5281/zenodo.10150" ID-GBIF-Dataset="b8b04597-d671-4969-8198-e61b5a172d35" ID-HNS-PUB="2958" checkinTime="1401177633152" checkinUser="donat" docAuthor="Ward, Philip S." docDate="1993" docId="E9140597FBACEA19CBBCB792E0942BFB" docLanguage="en" docName="2958.xml" docOrigin="Journal of Hymenoptera Research 2" docTitle="Pseudomyrmex janzeni Ward 1993, sp. nov." docType="treatment" docVersion="14" lastPageNumber="143" masterDocId="5D2504842856413260C70E0149648476" masterDocTitle="Systematic studies on Pseudomyrmex acacia-ants (Hymenoptera: Formicidae: Pseudomyrmecinae)" masterLastPageNumber="168" masterPageNumber="117" pageNumber="143" updateTime="1693793045367" updateUser="admin">
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<mods:title id="513668D5A72C3CF9C42A71EC6C254764">Systematic studies on Pseudomyrmex acacia-ants (Hymenoptera: Formicidae: Pseudomyrmecinae)</mods:title>
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<mods:namePart id="A18E5941A1F49045EBA9212700670756">Ward, Philip S.</mods:namePart>
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<mods:title id="8C772B4019662D3C291C383D6822FBDF">Journal of Hymenoptera Research</mods:title>
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<mods:date id="D7E93FF4BB95F37B5E902E612146B1F8">1993</mods:date>
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<treatment id="E9140597FBACEA19CBBCB792E0942BFB" ID-DOI="http://doi.org/10.5281/zenodo.3512904" ID-GBIF-Taxon="125136789" ID-Zenodo-Dep="3512904" LSID="urn:lsid:plazi:treatment:E9140597FBACEA19CBBCB792E0942BFB" httpUri="http://treatment.plazi.org/id/E9140597FBACEA19CBBCB792E0942BFB" lastPageId="28" lastPageNumber="143" pageId="27" pageNumber="143">
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<taxonomicName id="9F8EB22FC43A4706EC5002CDF2BE2F3B" ID-CoL="4P4F5" ID-ENA="600724" authority="Ward" authorityName="Ward" authorityYear="1993" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="143" phylum="Arthropoda" rank="species" species="janzeni" status="sp. nov.">Pseudomyrmex janzeni Ward</taxonomicName>
,
<taxonomicNameLabel id="6EB82A29CB32DB638D9CFE718CF46590" pageId="27" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="6A2780FDB9D1F00C1D6619FCF9B34551" pageId="27" pageNumber="143" type="description">
<paragraph id="197B9D0663551F6C3F2499D7D2D2D1B7" pageId="27" pageNumber="143" type="mainText">(Figs. 19, 29, 64, 70)</paragraph>
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. -
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,
<collectingMunicipality id="654D8A19C8E465BA165466AD1BCDAEFE" pageId="27" pageNumber="143">Nayarit</collectingMunicipality>
:
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<quantity id="C9CBA686E1B37EEA1B6FE9A185FE9807" metricMagnitude="4" metricUnit="m" metricValue="9.656064" pageId="27" pageNumber="143" unit="mi" value="60.0">60 mi</quantity>
. SE Acaponeta, Hwy.
<date id="FA186BBD22D83A51D4052E88070B4A4A" pageId="27" pageNumber="143" value="1915-09-15">
15,
<collectingDate id="9C96A2414BFFFD2750DA0376D348CC29" pageId="27" pageNumber="143">15.ix. 1963</collectingDate>
</date>
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,
<collectorName id="76206FB1382D366410C9BAC79CFEED4A" pageId="27" pageNumber="143">D. H. Janzen</collectorName>
(
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). HW 1.00, HL 1.08, EL 0.51, PL 0.55, PH 0.36.
</materialsCitation>
</paragraph>
<paragraph id="89681C15B24A16FA08249C7B95FC3303" pageId="27" pageNumber="143" type="mainText">
<materialsCitation id="62ECFC10E1C3782AA0E21F3EF91F496D" ID-GBIF-Occurrence="1416640631" collectingDate="1963-09-15" collectorName="D. H. Janzen" country="Mexico" location="60 mi. SE Acaponeta, Hwy. 15" pageId="27" pageNumber="143" specimenCount="273" specimenCount-male="45" specimenCount-queen="62" specimenCount-worker="166" stateProvince="Nayarit" typeStatus="paratype">
<typeStatus id="584AE200A9333F99AD7649155565011A" pageId="27" pageNumber="143">Paratypes</typeStatus>
. - Same data as holotype: series of
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,
<specimenCount id="D8D5CB2EDC204AF4BFA9A8AB1690E386" pageId="27" pageNumber="143" type="queen">62 queens</specimenCount>
and
<specimenCount id="7FCA7791CBCFA241EDD70AB58DAA6CFF" pageId="27" pageNumber="143" type="male">45 males</specimenCount>
</materialsCitation>
<materialsCitation id="9C0846CE94599A54BFB0F1FD90E15F2F" ID-GBIF-Occurrence="1416640639" collectingDate="1963-09-15" collectionCode="AMNH" collectorName="D. H. Janzen" country="Mexico" location="60 mi. SE Acaponeta, Hwy. 15" pageId="27" pageNumber="143" stateProvince="Nayarit" typeStatus="paratype">
(
<collectionCode id="1C1B27C16002DB87078435800404450B" collectionName="USA, New York, New York, American Museum of Natural History" pageId="27" pageNumber="143">AMNH</collectionCode>
,
</materialsCitation>
<materialsCitation id="BC0A234AFD12E1FFABFB5380B0E8E1A4" ID-GBIF-Occurrence="1416640671" collectingDate="1963-09-15" collectionCode="BMNH" collectorName="D. H. Janzen" country="Mexico" location="60 mi. SE Acaponeta, Hwy. 15" pageId="27" pageNumber="143" stateProvince="Nayarit" typeStatus="paratype">
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,
</materialsCitation>
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<collectionCode id="4B69861AB97090E2BC860D98240BF7D4" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="27" pageNumber="143">CASC</collectionCode>
,
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<materialsCitation id="C1462A613886FFC27E7D80C087BA2544" ID-GBIF-Occurrence="1416640677" collectingDate="1963-09-15" collectionCode="EBCC" collectorName="D. H. Janzen" country="Mexico" location="60 mi. SE Acaponeta, Hwy. 15" pageId="27" pageNumber="143" stateProvince="Nayarit" typeStatus="paratype">
<collectionCode id="720506F487918AD540376EE73E1C7DD4" collectionName="Mexico, Jalisco, San Patricio, Universidad Nacional Autonoma de Mexico, Estacion de Biologia &quot;Chamela&quot;" pageId="27" pageNumber="143">EBCC</collectionCode>
,
</materialsCitation>
<materialsCitation id="EEA70C1347E7A31B325FA751C9594FA7" ID-GBIF-Occurrence="1416640672" collectingDate="1963-09-15" collectionCode="GBFM" collectorName="D. H. Janzen" country="Mexico" location="60 mi. SE Acaponeta, Hwy. 15" pageId="27" pageNumber="143" stateProvince="Nayarit" typeStatus="paratype">
<collectionCode id="451461A33C47F4F7EA071E3E540E1406" collectionName="Panama, Panama, Universidad de Panama" pageId="27" pageNumber="143">GBFM</collectionCode>
,
</materialsCitation>
<materialsCitation id="E89CEB6363C9DD1347D188D1CB897D9C" ID-GBIF-Occurrence="1416640660" collectingDate="1963-09-15" collectionCode="INBC" collectorName="D. H. Janzen" country="Mexico" location="60 mi. SE Acaponeta, Hwy. 15" pageId="27" pageNumber="143" stateProvince="Nayarit" typeStatus="paratype">
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,
</materialsCitation>
<materialsCitation id="7B813CBB45BAD21E1BE818D93C2E29CB" ID-GBIF-Occurrence="1416640664" collectingDate="1963-09-15" collectionCode="LACM" collectorName="D. H. Janzen" country="Mexico" location="60 mi. SE Acaponeta, Hwy. 15" pageId="27" pageNumber="143" stateProvince="Nayarit" typeStatus="paratype">
<collectionCode id="3F504C962E247B2D5F173B2234101949" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History" pageId="27" pageNumber="143">LACM</collectionCode>
,
</materialsCitation>
<materialsCitation id="9B375F793151052F171709F70DEC3679" ID-GBIF-Occurrence="1416640668" collectingDate="1963-09-15" collectionCode="MCZC" collectorName="D. H. Janzen" country="Mexico" location="60 mi. SE Acaponeta, Hwy. 15" pageId="27" pageNumber="143" stateProvince="Nayarit" typeStatus="paratype">
<collectionCode id="B090FECF9B2BF0FDE8DBB4C97C745192" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology" pageId="27" pageNumber="143">MCZC</collectionCode>
,
</materialsCitation>
<materialsCitation id="545FD2C17F01CF1ACBA411B1C44C2833" ID-GBIF-Occurrence="1416640640" collectingDate="1963-09-15" collectionCode="MCZC" collectorName="D. H. Janzen" country="Mexico" location="60 mi. SE Acaponeta, Hwy. 15" pageId="27" pageNumber="143" stateProvince="Nayarit" typeStatus="paratype">
<collectionCode id="E4DBD857043E1946F2B6BCC1F59641CB" collectionName="Brazil, Sao Paulo, Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="27" pageNumber="143">MZSP</collectionCode>
,
</materialsCitation>
<materialsCitation id="E29497409BFDD302D929F995D4CBFD78" ID-GBIF-Occurrence="1416640674" collectingDate="1963-09-15" collectionCode="PSWC" collectorName="D. H. Janzen" country="Mexico" location="60 mi. SE Acaponeta, Hwy. 15" pageId="27" pageNumber="143" stateProvince="Nayarit" typeStatus="paratype">
<collectionCode id="5A2DAC5EFC6D960B1CC74C0EEFEA165B" collectionName="PSWC" pageId="27" pageNumber="143">PSWC</collectionCode>
,
</materialsCitation>
<materialsCitation id="6BA43DF9FF09EFC61506FD439CFB2361" ID-GBIF-Occurrence="1416640675" collectingDate="1963-09-15" collectionCode="UCDC" collectorName="D. H. Janzen" country="Mexico" location="60 mi. SE Acaponeta, Hwy. 15" pageId="27" pageNumber="143" stateProvince="Nayarit" typeStatus="paratype">
<collectionCode id="7C342EAEBEA7349E531001AADEE6D77A" collectionName="USA, California, Davis, University of California, R.M. Bohart Museum of Entomology" pageId="27" pageNumber="143">UCDC</collectionCode>
,
</materialsCitation>
<materialsCitation id="DFC9240F92582E6377E1F5716D9D074C" ID-GBIF-Occurrence="1416640646" collectingDate="1963-09-15" collectionCode="USNM" collectorName="D. H. Janzen" country="Mexico" location="60 mi. SE Acaponeta, Hwy. 15" pageId="27" pageNumber="143" stateProvince="Nayarit" typeStatus="paratype">
<collectionCode id="0707B5F5C21A4A870D77240C5E9AB956" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="27" pageNumber="143">USNM</collectionCode>
). Additional non-type material is listed below
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7EE65B8D12662D65501DCFFB2AB5C587" pageId="27" pageNumber="143" type="description">
<paragraph id="4EE1EB0F8388CF6F14BD2FB80E49C947" pageId="27" pageNumber="143" type="mainText">Worker measurements (n = 12). - HL 1.00-1.18, HW 0.93-1.03, MFC 0.063-0.096, Cl 0.88-0.94, REL 0.45-0.47, REL2 0.49-0.52, OOI 1.45-2.29, VI 0.58-0.68, FCI 0.068-0.098, SI 0.42-0.45, SI2 0.84-0.91, NI 0.62-0.69, PLI 0.59-0.71, PWI 0.63- 0.73, PPWI 1.37-1.73.</paragraph>
</subSubSection>
<subSubSection id="82BEC132E1D289513955D4EB00AD74F9" pageId="27" pageNumber="143" type="diagnosis">
<paragraph id="ECDEE55003A2B40D4C7572C1C00577EF" pageId="27" pageNumber="143" type="mainText">
Worker diagnosis. - Very similar to
<taxonomicName id="6E30BF3BD238C20858D73882AE658FCD" baseAuthorityName="F. Smith" baseAuthorityYear="1877" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="143" phylum="Arthropoda" rank="species" species="ferrugineus">P. ferrugineus</taxonomicName>
(q.v.) except as follows. Size smaller, on average. In lateral view mesonotum less steeply inclined; basal and declivitous faces of propodeum forming a less obtuse angle (compare Figs. 27 and 29). Petiole shorter and higher, on average (Fig. 48). Weak rugulo-punctate sculpture on propodeum even less evident than in
<taxonomicName id="50AE33D07A64F60B020435587D468069" baseAuthorityName="F. Smith" baseAuthorityYear="1877" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="143" phylum="Arthropoda" rank="species" species="ferrugineus">P. ferrugineus</taxonomicName>
. Pubescence denser, becoming decumbent to suberect on parts of body, most notably the gula (Fig. 19). Head and mesosoma rather light orange-brown, gaster the same or a slightly darker brown.
</paragraph>
</subSubSection>
<subSubSection id="81713D2E62081D395F1A7A45E455F919" pageId="27" pageNumber="143" type="discussion">
<paragraph id="B75853B2BF6DF0F64095EF57E08BAC74" pageId="27" pageNumber="143" type="mainText">
Comments. - Within the
<taxonomicName id="D3453C9EEF6605C7CE3D110338A51F8C" baseAuthorityName="F. Smith" baseAuthorityYear="1877" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="143" phylum="Arthropoda" rank="species" species="ferrugineus">P. ferrugineus</taxonomicName>
group
<taxonomicName id="06EC1371E7586C22AA190AC543E9F598" authorityName="Ward" authorityYear="1993" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="143" phylum="Arthropoda" rank="species" species="janzeni">P. janzeni</taxonomicName>
can be characterized by its relatively small size (worker and queen
<collectionCode id="8F38A2FE5FFF6834057FE3E23043BD15" pageId="27" pageNumber="143">HW</collectionCode>
&lt;1.04), broad head (worker Cl&gt; 0.86), laterally rounded median clypeal lobe, and uniform orange-brown color.
<taxonomicName id="0F416328961AE598553AB36944AF8357" authorityName="Ward" authorityYear="1993" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="143" phylum="Arthropoda" rank="species" species="janzeni">P. janzeni</taxonomicName>
is evidently closely related to
<taxonomicName id="D8DAB063BDE38C0991B81066DD02B391" baseAuthorityName="F. Smith" baseAuthorityYear="1877" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="143" phylum="Arthropoda" rank="species" species="ferrugineus">P. ferrugineus</taxonomicName>
(as surmised by
<bibRefCitation id="6EFB3C0EDF15AC6BB5B31DB02296DAA9" author="Janzen, D. H." journalOrPublisher="Journal of Animal Ecology" pagination="727 - 750" part="42" refId="ref40688" refString="Janzen, D. H. 1973. Evolution of polygynous obligate acacia-ants in western Mexico. Journal of Animal Ecology 42: 727 - 750." title="Evolution of polygynous obligate acacia-ants in western Mexico" type="journal article" year="1973">Janzen 1973</bibRefCitation>
); all of the metric measurements and indices of these two species overlap, although there is a tendency for
<taxonomicName id="AFAF11DB865999FE305C334A1BFC0651" authorityName="Ward" authorityYear="1993" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="143" phylum="Arthropoda" rank="species" species="janzeni">P. janzeni</taxonomicName>
workers to have shorter, higher petioles (Fig. 48). Workers and queens of
<taxonomicName id="D3F9A3554963EA4342D47D8090B3866F" authorityName="Ward" authorityYear="1993" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="143" phylum="Arthropoda" rank="species" species="janzeni">P. janzeni</taxonomicName>
are perhaps best distinguished from those of
<taxonomicName id="61C8F7C4C5C5C98269925C414C56F06F" baseAuthorityName="F. Smith" baseAuthorityYear="1877" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="143" phylum="Arthropoda" rank="species" species="ferrugineus">P. ferrugineus</taxonomicName>
by the combination of lighter orange-brown color, suberect gular pubescence (best seen in a backlit lateral view of the head), and the flatter profile of the worker mesosoma (see description above and Fig. 29). While some individuals of the highly variable
<taxonomicName id="51DB09927FE880BD864CBB93253CF15D" baseAuthorityName="F. Smith" baseAuthorityYear="1877" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="143" phylum="Arthropoda" rank="species" species="ferrugineus">P. ferrugineus</taxonomicName>
approach these conditions there is no indication of a convergence towards this morphology in western Mexico (Guerrero) where populations of
<taxonomicName id="A9130D9B2676B9C45B9B7E7F244B0A12" baseAuthorityName="F. Smith" baseAuthorityYear="1877" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="143" phylum="Arthropoda" rank="species" species="ferrugineus">P. ferrugineus</taxonomicName>
come closest to those of
<taxonomicName id="A3753DDD97CD229686A3B17F59B51BF6" authorityName="Ward" authorityYear="1993" class="Insecta" family="Formicidae" genus="Pseudomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="143" phylum="Arthropoda" rank="species" species="janzeni">P. janzeni</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E7C4F0E06542F4BE7FDA6C199605E435" lastPageId="28" lastPageNumber="144" pageId="27" pageNumber="143" type="distribution">
<paragraph id="10A992C8F13A459F8FDC96B0C481EA64" lastPageId="28" lastPageNumber="144" pageId="27" pageNumber="143" type="mainText">
Distribution and biology. - First recognized by
<bibRefCitation id="C87D48E540646685F7CB99145CD94736" author="Janzen, D. H." journalOrPublisher="Ecology" pagination="26 - 35" part="48" refId="ref40577" refString="Janzen, D. H. 1967 a. Fire, vegetation structure, and the ant x acacia interaction in Central America. Ecology 48: 26 - 35." title="Fire, vegetation structure, and the ant x acacia interaction in Central America" type="journal article" year="1967">Janzen (1967a</bibRefCitation>
,
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,
<bibRefCitation id="4C907D6648B3C37BB49CDF196E36987F" author="Janzen, D. H." journalOrPublisher="Journal of Animal Ecology" pagination="727 - 750" part="42" refId="ref40688" refString="Janzen, D. H. 1973. Evolution of polygynous obligate acacia-ants in western Mexico. Journal of Animal Ecology 42: 727 - 750." title="Evolution of polygynous obligate acacia-ants in western Mexico" type="journal article" year="1973">1973</bibRefCitation>
) as a distinct but undescribed species,
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is confined to a limited area in western Mexico (Fig. 70) where it is sympatric with the much darker colored and commoner
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. Colonies occupy
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and are polygynous; additional details of the life history can be found in
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