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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1134.87853" ID-Pensoft-Pub="1313-2970-1134-129" ID-Pensoft-UUID="0973AEF60C1D54EBA960C4E252F447B0" ID-ZooBank="C79C5E40D9C64E3B816F0201713DBA77" ModsDocID="1313-2970-1134-129" checkinTime="1670590531721" checkinUser="pensoft" docAuthor="Brunke, Adam J." docDate="2022" docId="E88883057D0355F9A33B62C5A65F7029" docLanguage="en" docName="ZooKeys 1134: 129-170" docOrigin="ZooKeys 1134" docPubDate="2022-12-08" docSource="http://dx.doi.org/10.3897/zookeys.1134.87853" docTitle="Quediellus debilis , comb. res." docType="treatment" docVersion="1" id="0973AEF60C1D54EBA960C4E252F447B0" lastPageNumber="129" masterDocId="0973AEF60C1D54EBA960C4E252F447B0" masterDocTitle="A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages" masterLastPageNumber="170" masterPageNumber="129" pageNumber="129" updateTime="1670590531721" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brunke, Adam J.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1158-936X</mods:nameIdentifier>
<mods:affiliation>Agriculture and Agri-Food Canada, Canadian National Collection of Insects, Arachnids and Nematodes, 960 Carling Avenue, Ottawa, Ontario, Canada</mods:affiliation>
<mods:nameIdentifier type="email">adam.j.brunke@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1134</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>129</mods:start>
<mods:end>170</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1134.87853</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1134.87853</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1134-129</mods:identifier>
<mods:identifier type="ZooBank">C79C5E40D9C64E3B816F0201713DBA77</mods:identifier>
<mods:identifier type="Pensoft-UUID">0973AEF60C1D54EBA960C4E252F447B0</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:E88883057D0355F9A33B62C5A65F7029" httpUri="http://treatment.plazi.org/id/E88883057D0355F9A33B62C5A65F7029" lastPageNumber="129" pageId="0" pageNumber="129">
<subSubSection pageId="0" pageNumber="129" type="nomenclature">
<paragraph pageId="0" pageNumber="129">
<taxonomicName LSID="E8888305-7D03-55F9-A33B-62C5A65F7029" authority="(Horn, 1878), comb. res." authorityName=", comb. res." baseAuthorityName="Horn" baseAuthorityYear="1878" class="Insecta" family="Staphylinidae" genus="Quediellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quediellus debilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="debilis">Quediellus debilis (Horn, 1878), comb. res.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="description">
<paragraph pageId="0" pageNumber="129">
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-C dorsal head A, B showing extent of head punctation (white arrow) and color of first antennomere base (black arrow) D-F elytral punctation and microsculpture A Paraquedius puncticeps (Horn) B P. marginicollis sp. nov. C, D Quediellus debilis (Horn) E Quedionuchus longipennis (Mannerheim) F Quedius densiventris (Casey). Abbreviations: af = anterior frontal puncture; b = basal puncture; pf = posterior frontal puncture; sa = supra-antennal puncture. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1134.87853.figure5" httpUri="https://binary.pensoft.net/fig/779763" pageId="0" pageNumber="129">Figs 5C, D</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A aedeagus in ventral view B, C, D, G, H, K, L median lobe of aedeagus B, G, K, L in ventral view, C, D, H, O-Q in lateral view E, F, I, J, M, N underside of paramere showing peg setae A-F Paraquedius marginicollis sp. nov. E atypical specimen from Clallam County, Washington, United States G-J Paraquedius puncticeps (Horn) I lectotype, from Vancouver, British Columbia, Canada J non-type specimen from Vancouver Island K-Q Quediellus debilis (Horn), specimens from east (K, M) and west (L, M) sides of the continental divide. Filled circles = typical ' nanulus' morphotype; open circles = typical ' debilis' morphotype. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure8" httpUri="https://binary.pensoft.net/fig/779766" pageId="0" pageNumber="129">, 8K-Q</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I female tergite X A Iratiquedius amabilis (Smetana) B I. mutator (Smetana) C I. prostans (Horn) D I. seriatus (Horn), evenly convex E I. uncifer sp. nov., discal impression F Paraquedius puncticeps (Horn) G P. marginicollis sp. nov. H-I Quediellus debilis (Horn). Pigmented area on tergite in E not shown for clarity but similar to I. seriatus. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1134.87853.figure9" httpUri="https://binary.pensoft.net/fig/779767" pageId="0" pageNumber="129">, 9H, I</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distributions of A Iratiquedius amabilis (Smetana) (circles), I. mutator (Smetana) (squares) B I. prostans (Horn) (black - specimens examined, white - literature records) C I. seriatus (Horn) (circles), I. uncifer sp. nov. (squares) D Paraquedius puncticeps (Horn) (circles), P. marginicollis sp. nov. (squares) E Quediellus debilis (Horn) (circles) (black - specimens examined, white - literature records)." figureDoi="10.3897/zookeys.1134.87853.figure11" httpUri="https://binary.pensoft.net/fig/779769" pageId="0" pageNumber="129">, 11E (map), 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="reference_group">
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Horn" authorityYear="1878" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius debilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="debilis">Quedius debilis</taxonomicName>
Horn, 1878: 156, 165.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName=", comb. res." baseAuthorityName="Horn" baseAuthorityYear="1878" class="Insecta" family="Staphylinidae" genus="Quediellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quediellus debilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="debilis">Quediellus debilis</taxonomicName>
;
<bibRefCitation author="Casey, TL" journalOrPublisher=", Lancaster, Pennsylvania" pageId="0" pageNumber="129" refId="B5" refString="Casey, TL, 1915. Studies in some staphylinid genera of North America. Memoirs on the Coleoptera VI. The New Era Printing Co., Lancaster, Pennsylvania" title="Studies in some staphylinid genera of North America. Memoirs on the Coleoptera VI. The New Era Printing Co." year="1915">Casey 1915</bibRefCitation>
(redescription).
</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Casey" authorityYear="1915" class="Insecta" family="Staphylinidae" genus="Quediellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quediellus helenae" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="helenae">Quediellus helenae</taxonomicName>
Casey, 1915: 403 syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Casey" authorityYear="1915" class="Insecta" family="Staphylinidae" genus="Quediellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quediellus humilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="humilis">Quediellus humilis</taxonomicName>
Casey, 1915: 403 syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Casey" authorityYear="1915" class="Insecta" family="Staphylinidae" genus="Quediellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quediellus nanulus" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="nanulus">Quediellus nanulus</taxonomicName>
Casey, 1915: 402 syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Horn" authorityYear="1878" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius (Raphirus) debilis" order="Coleoptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="debilis" subGenus="Raphirus">Quedius (Raphirus) debilis</taxonomicName>
;
<bibRefCitation DOI="https://doi.org/10.4039/entm10379fv" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="0" pageNumber="129" pagination="1 - 303" refId="B10" refString="Smetana, A, 1971. Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae). Memoirs of the Entomological Society of Canada 103 (79): 1 - 303, DOI: https://doi.org/10.4039/entm10379fv" title="Revision of the tribe Quediini of North America north of Mexico (Coleoptera: Staphylinidae)." url="https://doi.org/10.4039/entm10379fv" volume="103" year="1971">Smetana 1971</bibRefCitation>
(redescription);
<bibRefCitation DOI="https://doi.org/10.1111/syen.12468" author="Brunke, AJ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="396 - 421" refId="B4" refString="Brunke, AJ, Hansen, AK, Salnitska, M, Kypke, JL, Predeus, AV, Escalona, HE, Chapados, JT, Eyres, J, Richter, R, Smetana, A, Ślipinski, SA, Zwick, A, Hajek, J, Leschen, RAB, Solodovnikov, A, Dettman, JR, 2021. The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics. Systematic Entomology 46 (2): 396 - 421, DOI: https://doi.org/10.1111/syen.12468" title="The limits of Quediini at last (Staphylinidae: Staphylininae): a rove beetle mega-radiation resolved by comprehensive sampling and anchored phylogenomics." url="https://doi.org/10.1111/syen.12468" volume="46" year="2021">Brunke et al. 2021</bibRefCitation>
(phylogeny, as
<taxonomicName family="Staphylinidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Q. nanulus" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="nanulus">Q. nanulus</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="type locality">
<paragraph pageId="0" pageNumber="129">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="129">California (unknown locality), United States.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="type material">
<paragraph pageId="0" pageNumber="129">Type material.</paragraph>
<paragraph pageId="0" pageNumber="129">
<emphasis bold="true" pageId="0" pageNumber="129">
<emphasis italics="true" pageId="0" pageNumber="129">Lectotype</emphasis>
(female, MCZ, examined digitally)
</emphasis>
:
<normalizedToken originalValue="Cala">'Cala'</normalizedToken>
[white typed label] /
<taxonomicName family="Staphylinidae" kingdom="Animalia" lsidName="Q. debilis" order="Coleoptera" pageId="0" pageNumber="129" rank="species" species="debilis">Q. debilis</taxonomicName>
H. [handwritten label] / Type 7270 [red label].
</paragraph>
<paragraph pageId="0" pageNumber="129">
The female lectotype of this species is a classic example of the
<normalizedToken originalValue="debilis">'debilis'</normalizedToken>
morphotype (see below): a larger specimen with elytra at sides much longer than pronotum at midline, elytra at apex wider than pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>