443 lines
74 KiB
XML
443 lines
74 KiB
XML
<document id="A2D51F0A5BF2FF4548F920497B582D77" ID-CLB-Dataset="187955" ID-DOI="10.11646/zootaxa.5205.6.3" ID-GBIF-Dataset="9f91a850-2805-4dcf-84f1-4e4910792215" ID-ISSN="1175-5326" ID-Zenodo-Dep="7318720" ID-ZooBank="62DD9430-0B11-46A9-A590-B960F859FE5A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1668422342881" checkinUser="plazi" docAuthor="Hongsuwong, Thitipong, Sanguansub, Sunisa & Jaitrong, Weeyawat" docDate="2022" docId="E853E5051379FFEEFF03DF9AFA24FBFA" docLanguage="en" docName="zootaxa.5205.6.3.pdf" docOrigin="Zootaxa 5205 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parastasia Westwood 1841" docType="key" docVersion="7" lastPageNumber="560" masterDocId="146A9D7D1370FFE3FF94D90FFFB8FF8E" masterDocTitle="A new species and a new synonym in the scarab genus Parastasia Westwood, 1841 (Coleoptera: Scarabaeidae: Rutelinae), with a key to species from Thailand" masterLastPageNumber="562" masterPageNumber="547" pageNumber="556" updateTime="1699411326001" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="A76D992822DEC78C01821271E4F51D53">
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<mods:title id="913D1C3495482E03E944CE89940102DA">A new species and a new synonym in the scarab genus Parastasia Westwood, 1841 (Coleoptera: Scarabaeidae: Rutelinae), with a key to species from Thailand</mods:title>
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<mods:name id="BE81CC10E8C09A3FAE8D788E6581D8AF" type="personal">
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<mods:role id="2C385FD7FCC207E9FF40AD090DEBD88B">
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<mods:roleTerm id="D4F76D75642FB4421D7945D1093602CA">Author</mods:roleTerm>
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<mods:namePart id="0A4F6D6755D88649CA3736CB9487CA3D">Hongsuwong, Thitipong</mods:namePart>
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<mods:affiliation id="C89E417E52C1A799A03D11242DA5672A">Department of Entomology, Faculty of Agriculture at Kamphaeng Saen, Kasetsart University, Nakhon Pathom, 73140, Thailand. Office of Natural Science Research, National Science Museum, 39 Moo 3, Khlong 5, Khlong Luang, Pathum Thani 12120, Thailand. polyrhachis @ yahoo. com; https: // orcid. org / 0000 - 0003 - 1362 - 0754</mods:affiliation>
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<mods:name id="D58536D7812014BC31A34CAB364118C7" type="personal">
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<mods:roleTerm id="63AA57CEAB4D2BBDBE67382F0F4AA371">Author</mods:roleTerm>
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<mods:namePart id="B3CFFF4127749D36858FB3F6C9A3D334">Sanguansub, Sunisa</mods:namePart>
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<mods:affiliation id="178798A2F5BDEA7BE6C9EF856BD08B87">Department of Entomology, Faculty of Agriculture at Kamphaeng Saen, Kasetsart University, Nakhon Pathom, 73140, Thailand. Office of Natural Science Research, National Science Museum, 39 Moo 3, Khlong 5, Khlong Luang, Pathum Thani 12120, Thailand. polyrhachis @ yahoo. com; https: // orcid. org / 0000 - 0003 - 1362 - 0754</mods:affiliation>
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<mods:roleTerm id="0A0914E85EC255D9AA97F50430EC1684">Author</mods:roleTerm>
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<mods:namePart id="0A06375B2C08BE1340BA9CFB50214D3B">Jaitrong, Weeyawat</mods:namePart>
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<mods:nameIdentifier id="B7B3D532ECCBF0AE9511DB795EB3D6AC" type="ORCID">0000-0003-1362-0754</mods:nameIdentifier>
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<mods:affiliation id="4AB8A9D5B43B9A8CD6434D93DDABF0B1">Biology Divisions, Faculty of Science and Technology, Rajamangala University of Technology Thanyaburi, Pathum Thani 12120 Thailand.</mods:affiliation>
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<mods:nameIdentifier id="FC9379284E7958736A85360E314C91FC" type="email">polyrhachis@yahoo.com</mods:nameIdentifier>
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<mods:title id="9B4ADCEFE86471C85908514E3C1C57F2">Zootaxa</mods:title>
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<mods:date id="17E812DE4796119A008E0E03261B0C0C">2022</mods:date>
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<mods:number id="D4E0138669576B1BC4B12933B474637B">2022-11-11</mods:number>
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<mods:number id="F47F8854C5DE12C3DB1407BDB4EFBF12">5205</mods:number>
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<treatment id="E853E5051379FFEEFF03DF9AFA24FBFA" ID-DOI="http://doi.org/10.5281/zenodo.7327245" ID-GBIF-Taxon="205076399" ID-Zenodo-Dep="7327245" LSID="urn:lsid:plazi:treatment:E853E5051379FFEEFF03DF9AFA24FBFA" httpUri="http://treatment.plazi.org/id/E853E5051379FFEEFF03DF9AFA24FBFA" lastPageId="13" lastPageNumber="560" pageId="9" pageNumber="556">
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<subSubSection id="28E007981379FFEAFF03DF9AFCBFF920" box="[151,775,1684,1711]" pageId="9" pageNumber="556" type="nomenclature">
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<paragraph id="604554131379FFEAFF03DF9AFCBFF920" blockId="9.[151,775,1684,1711]" box="[151,775,1684,1711]" pageId="9" pageNumber="556">
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<heading id="3B0DE37F1379FFEAFF03DF9AFCBFF920" bold="true" box="[151,775,1684,1711]" fontSize="11" level="1" pageId="9" pageNumber="556" reason="1">
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<emphasis id="528E88011379FFEAFF03DF9AFCBFF920" bold="true" box="[151,775,1684,1711]" pageId="9" pageNumber="556">
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Key to species of
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<taxonomicName id="A7FA2F901379FFEAFEFADF9BFE55F920" authorityName="Westwood" authorityYear="1841" box="[366,493,1684,1710]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="556" phylum="Arthropoda" rank="genus">
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<emphasis id="528E88011379FFEAFEFADF9BFE55F920" bold="true" box="[366,493,1684,1710]" italics="true" pageId="9" pageNumber="556">Parastasia</emphasis>
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</taxonomicName>
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known from
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<collectingCountry id="18ED14831379FFEAFD01DF9BFCBFF920" box="[661,775,1684,1710]" name="Thailand" pageId="9" pageNumber="556">Thailand</collectingCountry>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="28E007981379FFEEFF03DFDCFA24FBFA" lastPageId="13" lastPageNumber="560" pageId="9" pageNumber="556" type="key">
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<keyStep id="DB0E4CB61379FFEAFF03DFDCFA24F8CE" pageId="9" pageNumber="556">
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<paragraph id="604554131379FFEAFF03DFDCFA24F88B" blockId="9.[151,1437,1745,2003]" pageId="9" pageNumber="556">
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<keyLead id="DB0BF7261379FFEAFF03DFDCFA24F88B" pageId="9" pageNumber="556">
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1 Small to medium in size, total length ˂
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<quantity id="A702F9F61379FFEAFDC9DFDCFD1BF966" box="[605,675,1747,1768]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="556" unit="mm" value="20.0">20 mm</quantity>
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; elytra with distinct rows of punctures; in lateral view, parameres not bilobed. ...................................................................................................
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<emphasis id="528E88011379FFEAFA04DFFFFA24F88B" bold="true" box="[1424,1436,1776,1797]" pageId="9" pageNumber="556">2</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="604554131379FFEAFF03DE01FA24F8CE" blockId="9.[151,1437,1745,2003]" pageId="9" pageNumber="556">
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<keyLead id="DB0BF7261379FFEAFF03DE01FA24F8CE" pageId="9" pageNumber="556">
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- Large in size, TL ≥
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<quantity id="A702F9F61379FFEAFE31DE01FE54F8AD" box="[421,492,1806,1827]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="556" unit="mm" value="20.0">20 mm</quantity>
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; rows of punctures on elytra indistinct (
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<figureCitation id="F8C148961379FFEAFCE7DE02FC04F8AD" box="[883,956,1805,1827]" captionStart="FIGURES 51–60" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[159,1424,192,1495]" captionTargetId="figure-18@11.[151,1436,181,1495]" captionTargetPageId="11" captionText="FIGURES 51–60. Characteristics of Parastasia species, non-type specimens. 51, Parastasia birmana Arrow, 1899, male (THNHM-I-09992); 52, 56, 59, Parastasia anomala Arrow, 1899, male (THNHM-I-09977); 53, Parastasia westwoodii Westwood, 1841, female (CTH); 54, Parastasia masumotoi Wada & Muramoto, 1999, male (CTH); 55, Parastasia sulcipennis Gestro, 1888, male (CTH); 57, 60, Parastasia bigibbosa Nonfried, 1891, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus." figureDoi="http://doi.org/10.5281/zenodo.7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="9" pageNumber="556">Fig. 51</figureCitation>
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); parameres slightly asymmetric, apex bilobed in lateral view (
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<figureCitation id="F8C148961379FFEAFEF4DE24FE6BF8CF" box="[352,467,1835,1857]" captionStart="FIGURES 61–70" captionStartId="12.[151,264,1853,1877]" captionTargetBox="[157,1420,190,1829]" captionTargetId="figure-18@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURES 61–70.Male genitalia of Parastasia species, non-type specimens.61–62, Parastasia birmanaArrow, 1899(THNHM- I-09992); 63–64, 69 Parastasia masumotoi Wada & Muramoto, 1999 (CTH); 65–66, 70, Parastasia sulcipennis Gestro, 1888 (CTH); 67, Parastasia anomala Arrow, 1899 (THNHM-I-09977); 68, Parastasia bigibbosa Nonfried, 1891 (CUT); 61, 63, 65, parameres in dorsal view; 62, 64, 66, aedeagus in lateral view; 67–70, endophallus." figureDoi="http://doi.org/10.5281/zenodo.7318740" httpUri="https://zenodo.org/record/7318740/files/figure.png" pageId="9" pageNumber="556">Figs. 61–62</figureCitation>
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)............................................................
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<taxonomicName id="A7FA2F901379FFEAFB3ADE24FA24F8CE" authority="Arrow, 1899" authorityName="Arrow" authorityYear="1899" box="[1198,1436,1834,1856]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="556" phylum="Arthropoda" rank="species" species="birmana">
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<emphasis id="528E88011379FFEAFB3ADE24FA24F8CE" bold="true" box="[1198,1436,1834,1856]" pageId="9" pageNumber="556">
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<emphasis id="528E88011379FFEAFB3ADE24FAA1F8CE" bold="true" box="[1198,1305,1834,1856]" italics="true" pageId="9" pageNumber="556">P. birmana</emphasis>
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<bibRefCitation id="046B29E21379FFEAFA8ADE24FA24F8CE" author="Arrow, G. J." box="[1310,1436,1835,1856]" pageId="9" pageNumber="556" pagination="479 - 499" refId="ref7039" refString="Arrow, G. J. (1899) On sexual dimorphism in the rutelid genus Parastasia, with descriptions of new species. Transactions of the Entomological Society London, 4, 479 - 499. https: // doi. org / 10.1111 / j. 1365 - 2311.1899. tb 00990. x" type="journal article" year="1899">Arrow, 1899</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="DB0E4CB61379FFEAFF03DE47FA24F816" pageId="9" pageNumber="556">
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<paragraph id="604554131379FFEAFF03DE47FA25F8F5" blockId="9.[151,1437,1745,2003]" pageId="9" pageNumber="556">
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<keyLead id="DB0BF7261379FFEAFF03DE47FA25F8F5" pageId="9" pageNumber="556">
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2 All claws simple, sickle-shaped without branching (
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<figureCitation id="F8C148961379FFEAFD7DDE47FC8DF8D0" box="[745,821,1864,1886]" captionStart="FIGURES 51–60" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[159,1424,192,1495]" captionTargetId="figure-18@11.[151,1436,181,1495]" captionTargetPageId="11" captionText="FIGURES 51–60. Characteristics of Parastasia species, non-type specimens. 51, Parastasia birmana Arrow, 1899, male (THNHM-I-09992); 52, 56, 59, Parastasia anomala Arrow, 1899, male (THNHM-I-09977); 53, Parastasia westwoodii Westwood, 1841, female (CTH); 54, Parastasia masumotoi Wada & Muramoto, 1999, male (CTH); 55, Parastasia sulcipennis Gestro, 1888, male (CTH); 57, 60, Parastasia bigibbosa Nonfried, 1891, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus." figureDoi="http://doi.org/10.5281/zenodo.7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="9" pageNumber="556">Fig. 58</figureCitation>
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); male aedeagus asymmetric, parameres fused together (
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<figureCitation id="F8C148961379FFEAFAF8DE47FEA7F8F5" captionStart="FIGURES 40–50" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[203,1381,190,1907]" captionTargetId="figure-18@10.[191,1396,181,1919]" captionTargetPageId="10" captionText="FIGURES 40–50. Parastasia bimaculata (Guérin-Méneville, 1843), non-type specimens. 40–42, 46, male (CTH); 43–45, 47, female (CTH); 48–50, aedeagus of male; 40, 43, 48, dorsal view; 41, 44, 49, ventral view; 42, 45, 50, lateral view; 46–47, caudal view." figureDoi="http://doi.org/10.5281/zenodo.7318736" httpUri="https://zenodo.org/record/7318736/files/figure.png" pageId="9" pageNumber="556">Figs. 48–50</figureCitation>
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).............................................................
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<taxonomicName id="A7FA2F901379FFEAFB9BDE69FA25F8F5" authority="(Guerin-Meneville, 1843)" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1843" box="[1039,1437,1893,1915]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="556" phylum="Arthropoda" rank="species" species="bimaculata">
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<emphasis id="528E88011379FFEAFB9BDE69FA25F8F5" bold="true" box="[1039,1437,1893,1915]" pageId="9" pageNumber="556">
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<emphasis id="528E88011379FFEAFB9BDE69FB2DF8F5" bold="true" box="[1039,1173,1893,1915]" italics="true" pageId="9" pageNumber="556">P. bimaculata</emphasis>
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(
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<bibRefCitation id="046B29E21379FFEAFB35DE6AFA2DF8F5" author="Guerin-Meneville, F. E." box="[1185,1429,1893,1915]" pageId="9" pageNumber="556" pagination="1 - 98" refId="ref7235" refString="Guerin-Meneville, F. E. (1843) Animaux articulees. In: Delessert A. (Eds.), Souvenirs d'un voyage dans l'Inde 2. Fortin, Paris, pp. 1 - 98." type="book chapter" year="1843">Guérin-Méneville, 1843</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="604554131379FFEAFF03DE8CFA24F816" blockId="9.[151,1437,1745,2003]" box="[151,1436,1923,1945]" pageId="9" pageNumber="556">
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<keyLead id="DB0BF7261379FFEAFF03DE8CFA24F816" box="[151,1436,1923,1945]" pageId="9" pageNumber="556">
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- External claw of meso- and metatarsi bifurcate; aedeagus symmetric or asymmetric, but parameres not fused together.....
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<emphasis id="528E88011379FFEAFA04DE8CFA24F816" bold="true" box="[1424,1436,1923,1944]" pageId="9" pageNumber="556">3</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="DB0E4CB61379FFEAFF03DEAFFA24F85D" pageId="9" pageNumber="556">
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<paragraph id="604554131379FFEAFF03DEAFFA24F83B" blockId="9.[151,1437,1745,2003]" box="[151,1436,1952,1974]" pageId="9" pageNumber="556">
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<keyLead id="DB0BF7261379FFEAFF03DEAFFA24F83B" box="[151,1436,1952,1974]" pageId="9" pageNumber="556">
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3 Propygidium and pygidium covered with decumbent setae (
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<figureCitation id="F8C148961379FFEAFC8BDEAFFC2AF838" box="[799,914,1952,1974]" captionStart="FIGURES 29–39" captionStartId="8.[151,264,1943,1967]" captionTargetBox="[186,1415,192,1919]" captionTargetId="figure-18@8.[172,1415,181,1919]" captionTargetPageId="8" captionText="FIGURES 29–39. Parastasia bigibbosa Nonfried, 1981, non-type specimens. 29–31, 35, male (THNHM-I-09982); 32–34, 36, female (THNHM-I-09981); 37–39, aedeagus of male; 29, 32, 37, dorsal view; 30, 33, 38, ventral view; 31, 34, 39, habitus in lateral view; 35–36, caudal view." figureDoi="http://doi.org/10.5281/zenodo.7318732" httpUri="https://zenodo.org/record/7318732/files/figure.png" pageId="9" pageNumber="556">Figs. 35–36</figureCitation>
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).........................................
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<emphasis id="528E88011379FFEAFA04DEAFFA24F83B" bold="true" box="[1424,1436,1952,1973]" pageId="9" pageNumber="556">4</emphasis>
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</keyLead>
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</paragraph>
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<paragraph id="604554131379FFEAFF03DEB1FA24F85D" blockId="9.[151,1437,1745,2003]" box="[151,1436,1981,2003]" pageId="9" pageNumber="556">
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<keyLead id="DB0BF7261379FFEAFF03DEB1FA24F85D" box="[151,1436,1981,2003]" pageId="9" pageNumber="556">
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- Propygidium and pygidium without long setae (
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<figureCitation id="F8C148961379FFEAFD39DEB2FD50F85D" box="[685,744,1981,2003]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1864,1888]" captionTargetBox="[277,1315,249,1829]" captionTargetId="figure-34@3.[255,1332,234,1837]" captionTargetPageId="3" captionText="FIGURES 1–5. Parastasia spinosa Hongsuwong, Sanguansub & Jaitrong, new species (holotype, male, THNHM-I-09998). 1, habitus in dorsal view; 2, habitus in ventral view; 3, habitus in lateral view; 4, habitus in caudal view; 5, inner side of right protarsus." figureDoi="http://doi.org/10.5281/zenodo.7318722" httpUri="https://zenodo.org/record/7318722/files/figure.png" pageId="9" pageNumber="556">Fig. 4</figureCitation>
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)........................................................
|
||
<emphasis id="528E88011379FFEAFA04DEB1FA24F85D" bold="true" box="[1424,1436,1982,2003]" pageId="9" pageNumber="556">7</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<caption id="3485049B137AFFE9FF03DE98FF76F876" ID-DOI="http://doi.org/10.5281/zenodo.7318736" ID-Zenodo-Dep="7318736" httpUri="https://zenodo.org/record/7318736/files/figure.png" pageId="10" pageNumber="557" startId="10.[151,264,1943,1967]" targetBox="[203,1381,190,1907]" targetPageId="10">
|
||
<paragraph id="60455413137AFFE9FF03DE98FF76F876" blockId="10.[151,1436,1943,2040]" pageId="10" pageNumber="557">
|
||
<emphasis id="528E8801137AFFE9FF03DE98FEEDF821" bold="true" box="[151,341,1943,1967]" pageId="10" pageNumber="557">FIGURES 40–50.</emphasis>
|
||
<taxonomicName id="A7FA2F90137AFFE9FECFDE97FCE9F83E" authority="(Guerin-Meneville, 1843)" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1843" box="[347,849,1944,1968]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="557" phylum="Arthropoda" rank="species" species="bimaculata">
|
||
<emphasis id="528E8801137AFFE9FECFDE97FDFAF83E" box="[347,578,1944,1968]" italics="true" pageId="10" pageNumber="557">Parastasia bimaculata</emphasis>
|
||
(
|
||
<bibRefCitation id="046B29E2137AFFE9FDDBDE97FCF2F83E" author="Guerin-Meneville, F. E." box="[591,842,1944,1968]" pageId="10" pageNumber="557" pagination="1 - 98" refId="ref7235" refString="Guerin-Meneville, F. E. (1843) Animaux articulees. In: Delessert A. (Eds.), Souvenirs d'un voyage dans l'Inde 2. Fortin, Paris, pp. 1 - 98." type="book chapter" year="1843">Guérin-Méneville, 1843</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, non-type specimens. 40–42, 46, male (CTH); 43–45, 47, female (CTH); 48–50, aedeagus of male; 40, 43, 48, dorsal view; 41, 44, 49, ventral view; 42, 45, 50, lateral view; 46–47, caudal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3485049B137BFFE8FF03DCE1FD01F918" ID-DOI="http://doi.org/10.5281/zenodo.7318738" ID-Zenodo-Dep="7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="11" pageNumber="558" startId="11.[151,264,1518,1542]" targetBox="[159,1424,192,1495]" targetPageId="11">
|
||
<paragraph id="60455413137BFFE8FF03DCE1FD01F918" blockId="11.[151,1436,1518,1687]" pageId="11" pageNumber="558">
|
||
<emphasis id="528E8801137BFFE8FF03DCE1FEE0F988" bold="true" box="[151,344,1518,1542]" pageId="11" pageNumber="558">FIGURES 51–60.</emphasis>
|
||
Characteristics of
|
||
<taxonomicName id="A7FA2F90137BFFE8FDB7DCE0FD5FF989" box="[547,743,1519,1543]" pageId="11" pageNumber="558">
|
||
<emphasis id="528E8801137BFFE8FDB7DCE0FD29F988" box="[547,657,1519,1542]" italics="true" pageId="11" pageNumber="558">Parastasia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, non-type specimens. 51,
|
||
<taxonomicName id="A7FA2F90137BFFE8FC6ADCE0FAE5F989" authority="Arrow, 1899" authorityName="Arrow" authorityYear="1899" box="[1022,1373,1519,1543]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="birmana">
|
||
<emphasis id="528E8801137BFFE8FC6ADCE0FB74F989" box="[1022,1228,1519,1543]" italics="true" pageId="11" pageNumber="558">Parastasia birmana</emphasis>
|
||
<bibRefCitation id="046B29E2137BFFE8FB42DCE0FAE5F989" author="Arrow, G. J." box="[1238,1373,1519,1543]" pageId="11" pageNumber="558" pagination="479 - 499" refId="ref7039" refString="Arrow, G. J. (1899) On sexual dimorphism in the rutelid genus Parastasia, with descriptions of new species. Transactions of the Entomological Society London, 4, 479 - 499. https: // doi. org / 10.1111 / j. 1365 - 2311.1899. tb 00990. x" type="journal article" year="1899">Arrow, 1899</bibRefCitation>
|
||
</taxonomicName>
|
||
, male (THNHM-I-09992); 52, 56, 59,
|
||
<taxonomicName id="A7FA2F90137BFFE8FE61DF1CFCE4F9A5" authority="Arrow, 1899" authorityName="Arrow" authorityYear="1899" box="[501,860,1555,1579]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="anomala">
|
||
<emphasis id="528E8801137BFFE8FE61DF1CFD71F9A5" box="[501,713,1555,1579]" italics="true" pageId="11" pageNumber="558">Parastasia anomala</emphasis>
|
||
<bibRefCitation id="046B29E2137BFFE8FD47DF1CFCE4F9A5" author="Arrow, G. J." box="[723,860,1555,1579]" pageId="11" pageNumber="558" pagination="479 - 499" refId="ref7039" refString="Arrow, G. J. (1899) On sexual dimorphism in the rutelid genus Parastasia, with descriptions of new species. Transactions of the Entomological Society London, 4, 479 - 499. https: // doi. org / 10.1111 / j. 1365 - 2311.1899. tb 00990. x" type="journal article" year="1899">Arrow, 1899</bibRefCitation>
|
||
</taxonomicName>
|
||
, male (THNHM-I-09977); 53,
|
||
<taxonomicName id="A7FA2F90137BFFE8FB24DF1CFEFBF9C1" authority="Westwood, 1841" authorityName="Westwood" authorityYear="1841" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="westwoodii">
|
||
<emphasis id="528E8801137BFFE8FB24DF1CFA24F9A5" box="[1200,1436,1555,1579]" italics="true" pageId="11" pageNumber="558">Parastasia westwoodii</emphasis>
|
||
<bibRefCitation id="046B29E2137BFFE8FF03DF38FEFBF9C1" author="Westwood, J. O." box="[151,323,1591,1615]" pageId="11" pageNumber="558" pagination="34 - 35" refId="ref8510" refString="Westwood, J. O. (1841) On the Australian genus Cryptodu s, and upon Parastasia, the Asiatic representative of the Rutelidae. Journal of Proceedings of the Entomological Society of London, 2: 34 - 35." type="journal article" year="1841">Westwood, 1841</bibRefCitation>
|
||
</taxonomicName>
|
||
, female (CTH); 54,
|
||
<taxonomicName id="A7FA2F90137BFFE8FD9BDF38FC46F9C1" authority="Wada & Muramoto, 1999" authorityName="Wada & Muramoto" authorityYear="1999" box="[527,1022,1591,1615]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="masumotoi">
|
||
<emphasis id="528E8801137BFFE8FD9BDF38FD4AF9C0" box="[527,754,1591,1614]" italics="true" pageId="11" pageNumber="558">Parastasia masumotoi</emphasis>
|
||
<bibRefCitation id="046B29E2137BFFE8FD6CDF38FC46F9C1" author="Wada K. & Muramoto R." box="[760,1022,1591,1615]" pageId="11" pageNumber="558" pagination="167 - 175" refId="ref8478" refString="Wada K. & Muramoto R. (1999) Three New Species of the genus Parastasia from Southeast Asia. Elytra, 27 (1): 167 - 175." type="journal article" year="1999">Wada & Muramoto, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
, male (CTH); 55,
|
||
<taxonomicName id="A7FA2F90137BFFE8FB22DF38FEA6F9FD" authority="Gestro, 1888" authorityName="Gestro" authorityYear="1888" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="sulcipennis">
|
||
<emphasis id="528E8801137BFFE8FB22DF38FA24F9C1" box="[1206,1436,1591,1615]" italics="true" pageId="11" pageNumber="558">Parastasia sulcipennis</emphasis>
|
||
Gestro, 1888
|
||
</taxonomicName>
|
||
, male (CTH); 57, 60,
|
||
<taxonomicName id="A7FA2F90137BFFE8FD93DF54FC30F9FD" authority="Nonfried, 1891" authorityName="Nonfried" authorityYear="1891" box="[519,904,1627,1651]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="bigibbosa">
|
||
<emphasis id="528E8801137BFFE8FD93DF54FD59F9FD" box="[519,737,1627,1651]" italics="true" pageId="11" pageNumber="558">Parastasia bigibbosa</emphasis>
|
||
<bibRefCitation id="046B29E2137BFFE8FD7DDF54FC30F9FD" author="Nonfried, A. F." box="[745,904,1627,1651]" pageId="11" pageNumber="558" pagination="221 - 240" refId="ref7630" refString="Nonfried, A. F. (1891) Neue afrikanische, central-amerikanische und ost-asiatische Melolonthiden und Ruteliden. Berliner Entomologische Zeitschrift, 36 (2), 221 - 240." type="journal article" year="1891">Nonfried, 1891</bibRefCitation>
|
||
</taxonomicName>
|
||
, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3485049B137CFFEFFF03DE32FC5CF84C" ID-DOI="http://doi.org/10.5281/zenodo.7318740" ID-Zenodo-Dep="7318740" httpUri="https://zenodo.org/record/7318740/files/figure.png" pageId="12" pageNumber="559" startId="12.[151,264,1853,1877]" targetBox="[157,1420,190,1829]" targetPageId="12">
|
||
<paragraph id="60455413137CFFEFFF03DE32FC5CF84C" blockId="12.[151,1437,1853,1986]" pageId="12" pageNumber="559">
|
||
<emphasis id="528E8801137CFFEFFF03DE32FEEAF8DB" bold="true" box="[151,338,1853,1877]" pageId="12" pageNumber="559">FIGURES 61–70.</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="A7FA2F90137CFFEFFD96DE31FD78F8D8" box="[514,704,1854,1878]" pageId="12" pageNumber="559">
|
||
<emphasis id="528E8801137CFFEFFD96DE31FDC8F8DB" box="[514,624,1854,1877]" italics="true" pageId="12" pageNumber="559">Parastasia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, non-type specimens. 61–62,
|
||
<taxonomicName id="A7FA2F90137CFFEFFC77DE31FA93F8D8" authority="Arrow, 1899" authorityName="Arrow" authorityYear="1899" box="[995,1323,1854,1878]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="birmana">
|
||
<emphasis id="528E8801137CFFEFFC77DE31FB13F8D8" box="[995,1195,1854,1878]" italics="true" pageId="12" pageNumber="559">Parastasia birmana</emphasis>
|
||
<bibRefCitation id="046B29E2137CFFEFFB39DE31FA93F8D8" author="Arrow, G. J." box="[1197,1323,1854,1878]" pageId="12" pageNumber="559" pagination="479 - 499" refId="ref7039" refString="Arrow, G. J. (1899) On sexual dimorphism in the rutelid genus Parastasia, with descriptions of new species. Transactions of the Entomological Society London, 4, 479 - 499. https: // doi. org / 10.1111 / j. 1365 - 2311.1899. tb 00990. x" type="journal article" year="1899">Arrow, 1899</bibRefCitation>
|
||
</taxonomicName>
|
||
(THNHM- I-09992); 63–64, 69
|
||
<taxonomicName id="A7FA2F90137CFFEFFEF8DE6DFCE5F8F4" authority="Wada & Muramoto, 1999" authorityName="Wada & Muramoto" authorityYear="1999" box="[364,861,1890,1914]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="masumotoi">
|
||
<emphasis id="528E8801137CFFEFFEF8DE6DFDE8F8F7" box="[364,592,1890,1913]" italics="true" pageId="12" pageNumber="559">Parastasia masumotoi</emphasis>
|
||
<bibRefCitation id="046B29E2137CFFEFFDC3DE6DFCE5F8F4" author="Wada K. & Muramoto R." box="[599,861,1890,1914]" pageId="12" pageNumber="559" pagination="167 - 175" refId="ref8478" refString="Wada K. & Muramoto R. (1999) Three New Species of the genus Parastasia from Southeast Asia. Elytra, 27 (1): 167 - 175." type="journal article" year="1999">Wada & Muramoto, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
(CTH); 65–66, 70,
|
||
<taxonomicName id="A7FA2F90137CFFEFFBBEDE6DFA25F8F4" authority="Gestro, 1888" authorityName="Gestro" authorityYear="1888" box="[1066,1437,1890,1914]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="sulcipennis">
|
||
<emphasis id="528E8801137CFFEFFBBEDE6DFAA9F8F4" box="[1066,1297,1890,1914]" italics="true" pageId="12" pageNumber="559">Parastasia sulcipennis</emphasis>
|
||
Gestro, 1888
|
||
</taxonomicName>
|
||
(CTH); 67,
|
||
<taxonomicName id="A7FA2F90137CFFEFFE9ADE89FDD8F810" authority="Arrow, 1899" authorityName="Arrow" authorityYear="1899" box="[270,608,1926,1950]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="anomala">
|
||
<emphasis id="528E8801137CFFEFFE9ADE89FE63F810" box="[270,475,1926,1950]" italics="true" pageId="12" pageNumber="559">Parastasia anomala</emphasis>
|
||
<bibRefCitation id="046B29E2137CFFEFFE74DE89FDD8F810" author="Arrow, G. J." box="[480,608,1926,1950]" pageId="12" pageNumber="559" pagination="479 - 499" refId="ref7039" refString="Arrow, G. J. (1899) On sexual dimorphism in the rutelid genus Parastasia, with descriptions of new species. Transactions of the Entomological Society London, 4, 479 - 499. https: // doi. org / 10.1111 / j. 1365 - 2311.1899. tb 00990. x" type="journal article" year="1899">Arrow, 1899</bibRefCitation>
|
||
</taxonomicName>
|
||
(THNHM-I-09977); 68,
|
||
<taxonomicName id="A7FA2F90137CFFEFFCF4DE89FB62F810" authority="Nonfried, 1891" authorityName="Nonfried" authorityYear="1891" box="[864,1242,1926,1950]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="bigibbosa">
|
||
<emphasis id="528E8801137CFFEFFCF4DE89FB80F810" box="[864,1080,1926,1950]" italics="true" pageId="12" pageNumber="559">Parastasia bigibbosa</emphasis>
|
||
<bibRefCitation id="046B29E2137CFFEFFBAADE89FB62F810" author="Nonfried, A. F." box="[1086,1242,1926,1950]" pageId="12" pageNumber="559" pagination="221 - 240" refId="ref7630" refString="Nonfried, A. F. (1891) Neue afrikanische, central-amerikanische und ost-asiatische Melolonthiden und Ruteliden. Berliner Entomologische Zeitschrift, 36 (2), 221 - 240." type="journal article" year="1891">Nonfried, 1891</bibRefCitation>
|
||
</taxonomicName>
|
||
(CUT); 61, 63, 65, parameres in dorsal view; 62, 64, 66, aedeagus in lateral view; 67–70, endophallus.
|
||
</paragraph>
|
||
</caption>
|
||
<keyStep id="DB0E4CB6137DFFEEFF03D997FA24FEB1" pageId="13" pageNumber="560">
|
||
<paragraph id="60455413137DFFEEFF03D997FA24FF69" blockId="13.[151,1437,150,1141]" pageId="13" pageNumber="560">
|
||
<keyLead id="DB0BF726137DFFEEFF03D997FA24FF69" pageId="13" pageNumber="560">
|
||
4 Eyes narrow, ratio of IOD and HW ˃ 0.67 (
|
||
<figureCitation id="F8C14896137DFFEEFD04D998FD61FF23" box="[656,729,151,173]" captionStart="FIGURES 51–60" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[159,1424,192,1495]" captionTargetId="figure-18@11.[151,1436,181,1495]" captionTargetPageId="11" captionText="FIGURES 51–60. Characteristics of Parastasia species, non-type specimens. 51, Parastasia birmana Arrow, 1899, male (THNHM-I-09992); 52, 56, 59, Parastasia anomala Arrow, 1899, male (THNHM-I-09977); 53, Parastasia westwoodii Westwood, 1841, female (CTH); 54, Parastasia masumotoi Wada & Muramoto, 1999, male (CTH); 55, Parastasia sulcipennis Gestro, 1888, male (CTH); 57, 60, Parastasia bigibbosa Nonfried, 1891, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus." figureDoi="http://doi.org/10.5281/zenodo.7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="13" pageNumber="560">Fig. 56</figureCitation>
|
||
); antennal clubs distinctly shorter than antennal segment 1–7 combined; mesotarsi of male somewhat enlarged, segment 5 as long as or longer than segment 1–4 combined (
|
||
<figureCitation id="F8C14896137DFFEEFB23D9BAFAB8FF45" box="[1207,1280,181,203]" captionStart="FIGURES 51–60" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[159,1424,192,1495]" captionTargetId="figure-18@11.[151,1436,181,1495]" captionTargetPageId="11" captionText="FIGURES 51–60. Characteristics of Parastasia species, non-type specimens. 51, Parastasia birmana Arrow, 1899, male (THNHM-I-09992); 52, 56, 59, Parastasia anomala Arrow, 1899, male (THNHM-I-09977); 53, Parastasia westwoodii Westwood, 1841, female (CTH); 54, Parastasia masumotoi Wada & Muramoto, 1999, male (CTH); 55, Parastasia sulcipennis Gestro, 1888, male (CTH); 57, 60, Parastasia bigibbosa Nonfried, 1891, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus." figureDoi="http://doi.org/10.5281/zenodo.7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="13" pageNumber="560">Fig. 56</figureCitation>
|
||
); median sac of endophallus membranous without any appendages or spines, but sometimes distinctly with sclerotic surface (
|
||
<figureCitation id="F8C14896137DFFEEFB6DD9DDFAF9FF66" box="[1273,1345,210,232]" captionStart="FIGURES 61–70" captionStartId="12.[151,264,1853,1877]" captionTargetBox="[157,1420,190,1829]" captionTargetId="figure-18@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURES 61–70.Male genitalia of Parastasia species, non-type specimens.61–62, Parastasia birmanaArrow, 1899(THNHM- I-09992); 63–64, 69 Parastasia masumotoi Wada & Muramoto, 1999 (CTH); 65–66, 70, Parastasia sulcipennis Gestro, 1888 (CTH); 67, Parastasia anomala Arrow, 1899 (THNHM-I-09977); 68, Parastasia bigibbosa Nonfried, 1891 (CUT); 61, 63, 65, parameres in dorsal view; 62, 64, 66, aedeagus in lateral view; 67–70, endophallus." figureDoi="http://doi.org/10.5281/zenodo.7318740" httpUri="https://zenodo.org/record/7318740/files/figure.png" pageId="13" pageNumber="560">Fig. 67</figureCitation>
|
||
)......
|
||
<emphasis id="528E8801137DFFEEFA04D9DDFA24FF69" bold="true" box="[1424,1436,210,231]" pageId="13" pageNumber="560">5</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="60455413137DFFEEFF03D9FFFA24FEB1" blockId="13.[151,1437,150,1141]" pageId="13" pageNumber="560">
|
||
<keyLead id="DB0BF726137DFFEEFF03D9FFFA24FEB1" pageId="13" pageNumber="560">
|
||
- Eyes broad, ratio of IOD and HW ˂ 0.60 (
|
||
<figureCitation id="F8C14896137DFFEEFDFED9E0FD08FE8B" box="[618,688,239,261]" captionStart="FIGURES 51–60" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[159,1424,192,1495]" captionTargetId="figure-18@11.[151,1436,181,1495]" captionTargetPageId="11" captionText="FIGURES 51–60. Characteristics of Parastasia species, non-type specimens. 51, Parastasia birmana Arrow, 1899, male (THNHM-I-09992); 52, 56, 59, Parastasia anomala Arrow, 1899, male (THNHM-I-09977); 53, Parastasia westwoodii Westwood, 1841, female (CTH); 54, Parastasia masumotoi Wada & Muramoto, 1999, male (CTH); 55, Parastasia sulcipennis Gestro, 1888, male (CTH); 57, 60, Parastasia bigibbosa Nonfried, 1891, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus." figureDoi="http://doi.org/10.5281/zenodo.7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="13" pageNumber="560">Fig. 57</figureCitation>
|
||
); antennal clubs clearly longer than antennal segment 1–7 combined; mesotarsi simple in both sexes, tarsal segment 5 shorter than segment 1–4 combined (
|
||
<figureCitation id="F8C14896137DFFEEFC25D802FC41FEAD" box="[945,1017,269,291]" captionStart="FIGURES 51–60" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[159,1424,192,1495]" captionTargetId="figure-18@11.[151,1436,181,1495]" captionTargetPageId="11" captionText="FIGURES 51–60. Characteristics of Parastasia species, non-type specimens. 51, Parastasia birmana Arrow, 1899, male (THNHM-I-09992); 52, 56, 59, Parastasia anomala Arrow, 1899, male (THNHM-I-09977); 53, Parastasia westwoodii Westwood, 1841, female (CTH); 54, Parastasia masumotoi Wada & Muramoto, 1999, male (CTH); 55, Parastasia sulcipennis Gestro, 1888, male (CTH); 57, 60, Parastasia bigibbosa Nonfried, 1891, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus." figureDoi="http://doi.org/10.5281/zenodo.7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="13" pageNumber="560">Fig. 60</figureCitation>
|
||
); median sac of endophallus with numerous spines (
|
||
<figureCitation id="F8C14896137DFFEEFEBFD825FECBFECE" box="[299,371,298,320]" captionStart="FIGURES 61–70" captionStartId="12.[151,264,1853,1877]" captionTargetBox="[157,1420,190,1829]" captionTargetId="figure-18@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURES 61–70.Male genitalia of Parastasia species, non-type specimens.61–62, Parastasia birmanaArrow, 1899(THNHM- I-09992); 63–64, 69 Parastasia masumotoi Wada & Muramoto, 1999 (CTH); 65–66, 70, Parastasia sulcipennis Gestro, 1888 (CTH); 67, Parastasia anomala Arrow, 1899 (THNHM-I-09977); 68, Parastasia bigibbosa Nonfried, 1891 (CUT); 61, 63, 65, parameres in dorsal view; 62, 64, 66, aedeagus in lateral view; 67–70, endophallus." figureDoi="http://doi.org/10.5281/zenodo.7318740" httpUri="https://zenodo.org/record/7318740/files/figure.png" pageId="13" pageNumber="560">Fig. 68</figureCitation>
|
||
).................................................................
|
||
<taxonomicName id="A7FA2F90137DFFEEFB1CD825FA24FEB1" authority="Nonfried, 1891" authorityName="Nonfried" authorityYear="1891" box="[1160,1436,298,320]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="560" phylum="Arthropoda" rank="species" species="bigibbosa">
|
||
<emphasis id="528E8801137DFFEEFB1CD825FA24FEB1" bold="true" box="[1160,1436,298,320]" pageId="13" pageNumber="560">
|
||
<emphasis id="528E8801137DFFEEFB1CD825FB47FECE" bold="true" box="[1160,1279,298,320]" italics="true" pageId="13" pageNumber="560">P. bigibbosa</emphasis>
|
||
<bibRefCitation id="046B29E2137DFFEEFA90D825FA24FEB1" author="Nonfried, A. F." box="[1284,1436,298,320]" pageId="13" pageNumber="560" pagination="221 - 240" refId="ref7630" refString="Nonfried, A. F. (1891) Neue afrikanische, central-amerikanische und ost-asiatische Melolonthiden und Ruteliden. Berliner Entomologische Zeitschrift, 36 (2), 221 - 240." type="journal article" year="1891">Nonfried, 1891</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="DB0E4CB6137DFFEEFF03D848FA24FE3A" pageId="13" pageNumber="560">
|
||
<paragraph id="60455413137DFFEEFF03D848FA24FEF4" blockId="13.[151,1437,150,1141]" pageId="13" pageNumber="560">
|
||
<keyLead id="DB0BF726137DFFEEFF03D848FA24FEF4" pageId="13" pageNumber="560">
|
||
5 Dorsal surface mostly brownish (
|
||
<figureCitation id="F8C14896137DFFEEFDB9D848FDC0FED3" box="[557,632,327,349]" captionStart="FIGURES 51–60" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[159,1424,192,1495]" captionTargetId="figure-18@11.[151,1436,181,1495]" captionTargetPageId="11" captionText="FIGURES 51–60. Characteristics of Parastasia species, non-type specimens. 51, Parastasia birmana Arrow, 1899, male (THNHM-I-09992); 52, 56, 59, Parastasia anomala Arrow, 1899, male (THNHM-I-09977); 53, Parastasia westwoodii Westwood, 1841, female (CTH); 54, Parastasia masumotoi Wada & Muramoto, 1999, male (CTH); 55, Parastasia sulcipennis Gestro, 1888, male (CTH); 57, 60, Parastasia bigibbosa Nonfried, 1891, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus." figureDoi="http://doi.org/10.5281/zenodo.7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="13" pageNumber="560">Fig. 53</figureCitation>
|
||
); mesotarsi clearly enlarged in male, segment 5 much longer than segments 1–4 combined............................................................................................
|
||
<emphasis id="528E8801137DFFEEFA04D86AFA24FEF4" bold="true" box="[1424,1436,357,378]" pageId="13" pageNumber="560">6</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="60455413137DFFEEFF03D88CFA24FE3A" blockId="13.[151,1437,150,1141]" pageId="13" pageNumber="560">
|
||
<keyLead id="DB0BF726137DFFEEFF03D88CFA24FE3A" pageId="13" pageNumber="560">
|
||
- Dorsal surface mostly orange-reddish (
|
||
<figureCitation id="F8C14896137DFFEEFDCCD88DFD27FE16" box="[600,671,386,408]" captionStart="FIGURES 51–60" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[159,1424,192,1495]" captionTargetId="figure-18@11.[151,1436,181,1495]" captionTargetPageId="11" captionText="FIGURES 51–60. Characteristics of Parastasia species, non-type specimens. 51, Parastasia birmana Arrow, 1899, male (THNHM-I-09992); 52, 56, 59, Parastasia anomala Arrow, 1899, male (THNHM-I-09977); 53, Parastasia westwoodii Westwood, 1841, female (CTH); 54, Parastasia masumotoi Wada & Muramoto, 1999, male (CTH); 55, Parastasia sulcipennis Gestro, 1888, male (CTH); 57, 60, Parastasia bigibbosa Nonfried, 1891, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus." figureDoi="http://doi.org/10.5281/zenodo.7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="13" pageNumber="560">Fig. 52</figureCitation>
|
||
); segment 5 of mesotarsi as long as segments 1–4 combined in male.........................................................................................
|
||
<taxonomicName id="A7FA2F90137DFFEEFB3FD8AFFA24FE3A" authority="Arrow, 1899" authorityName="Arrow" authorityYear="1899" box="[1195,1436,415,437]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="560" phylum="Arthropoda" rank="species" species="anomala">
|
||
<emphasis id="528E8801137DFFEEFB3FD8AFFA24FE3A" bold="true" box="[1195,1436,415,437]" pageId="13" pageNumber="560">
|
||
<emphasis id="528E8801137DFFEEFB3FD8AFFAA2FE3B" bold="true" box="[1195,1306,415,437]" italics="true" pageId="13" pageNumber="560">P. anomala</emphasis>
|
||
<bibRefCitation id="046B29E2137DFFEEFA8AD890FA24FE3A" author="Arrow, G. J." box="[1310,1436,415,436]" pageId="13" pageNumber="560" pagination="479 - 499" refId="ref7039" refString="Arrow, G. J. (1899) On sexual dimorphism in the rutelid genus Parastasia, with descriptions of new species. Transactions of the Entomological Society London, 4, 479 - 499. https: // doi. org / 10.1111 / j. 1365 - 2311.1899. tb 00990. x" type="journal article" year="1899">Arrow, 1899</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="DB0E4CB6137DFFEEFF03D8B2FA24FDC9" pageId="13" pageNumber="560">
|
||
<paragraph id="60455413137DFFEEFF03D8B2FA24FD82" blockId="13.[151,1437,150,1141]" pageId="13" pageNumber="560">
|
||
<keyLead id="DB0BF726137DFFEEFF03D8B2FA24FD82" pageId="13" pageNumber="560">
|
||
6 Ground color of dorsal surface reddish brown to dark brown, posterolateral portions of pronotum with a yellow to testaceous patch; in dorsal view, male aedeagus symmetric.
|
||
<collectingCountry id="18ED1483137DFFEEFD2DD8D5FD55FE7E" box="[697,749,474,496]" name="India" pageId="13" pageNumber="560">India</collectingCountry>
|
||
,
|
||
<collectingCountry id="18ED1483137DFFEEFD62D8D5FC86FE7E" box="[758,830,474,496]" name="Bhutan" pageId="13" pageNumber="560">Bhutan</collectingCountry>
|
||
,
|
||
<collectingCountry id="18ED1483137DFFEEFCDCD8D5FC3CFE7E" box="[840,900,474,496]" name="China" pageId="13" pageNumber="560">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="18ED1483137DFFEEFC18D8D5FC58FE7E" box="[908,992,474,496]" name="Vietnam" pageId="13" pageNumber="560">Vietnam</collectingCountry>
|
||
,
|
||
<collectingCountry id="18ED1483137DFFEEFC7ED8D4FBA3FE7E" box="[1002,1051,475,496]" name="Laos" pageId="13" pageNumber="560">Laos</collectingCountry>
|
||
, and northern
|
||
<collectingCountry id="18ED1483137DFFEEFB30D8D5FB44FE7E" box="[1188,1276,474,496]" name="Thailand" pageId="13" pageNumber="560">Thailand</collectingCountry>
|
||
..............................................................................................
|
||
<taxonomicName id="A7FA2F90137DFFEEFB57D8F7FA24FD82" authority="Ohaus, 1898" authorityName="Ohaus" authorityYear="1898" box="[1219,1436,503,525]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="560" phylum="Arthropoda" rank="species" species="indica">
|
||
<emphasis id="528E8801137DFFEEFB57D8F7FA24FD82" bold="true" box="[1219,1436,503,525]" pageId="13" pageNumber="560">
|
||
<emphasis id="528E8801137DFFEEFB57D8F7FAAFFD83" bold="true" box="[1219,1303,503,525]" italics="true" pageId="13" pageNumber="560">P. indica</emphasis>
|
||
<bibRefCitation id="046B29E2137DFFEEFA8AD8F8FA24FD82" author="Ohaus, F." box="[1310,1436,503,525]" pageId="13" pageNumber="560" pagination="3 - 41" refId="ref7662" refString="Ohaus, F. (1898) Phaenomeridae. Stettiner Entomologische Zeitung, 59 (1 - 2), 3 - 41." type="journal article" year="1898">Ohaus, 1898</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="60455413137DFFEEFF03DB1AFA24FDC9" blockId="13.[151,1437,150,1141]" pageId="13" pageNumber="560">
|
||
<keyLead id="DB0BF726137DFFEEFF03DB1AFA24FDC9" pageId="13" pageNumber="560">
|
||
- Ground color of dorsal surface brownish with dark brown brands on pronotum and elytra (
|
||
<figureCitation id="F8C14896137DFFEEFBECDB1AFB7CFDA5" box="[1144,1220,533,555]" captionStart="FIGURES 51–60" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[159,1424,192,1495]" captionTargetId="figure-18@11.[151,1436,181,1495]" captionTargetPageId="11" captionText="FIGURES 51–60. Characteristics of Parastasia species, non-type specimens. 51, Parastasia birmana Arrow, 1899, male (THNHM-I-09992); 52, 56, 59, Parastasia anomala Arrow, 1899, male (THNHM-I-09977); 53, Parastasia westwoodii Westwood, 1841, female (CTH); 54, Parastasia masumotoi Wada & Muramoto, 1999, male (CTH); 55, Parastasia sulcipennis Gestro, 1888, male (CTH); 57, 60, Parastasia bigibbosa Nonfried, 1891, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus." figureDoi="http://doi.org/10.5281/zenodo.7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="13" pageNumber="560">Fig. 53</figureCitation>
|
||
); aedeagus curve and asymmetric. Southern
|
||
<collectingCountry id="18ED1483137DFFEEFE2EDB3DFDAAFDC6" box="[442,530,562,584]" name="Thailand" pageId="13" pageNumber="560">Thailand</collectingCountry>
|
||
,
|
||
<collectingCountry id="18ED1483137DFFEEFD8FDB3DFDCDFDC6" box="[539,629,562,584]" name="Malaysia" pageId="13" pageNumber="560">Malaysia</collectingCountry>
|
||
,
|
||
<collectingCountry id="18ED1483137DFFEEFDEBDB3DFD67FDC6" box="[639,735,562,584]" name="Indonesia" pageId="13" pageNumber="560">Indonesia</collectingCountry>
|
||
, and
|
||
<collectingCountry id="18ED1483137DFFEEFC85DB3DFC38FDC6" box="[785,896,562,584]" name="Philippines" pageId="13" pageNumber="560">Philippines</collectingCountry>
|
||
...................
|
||
<taxonomicName id="A7FA2F90137DFFEEFBFADB3DFA24FDC9" authority="Westwood, 1841" authorityName="Westwood" authorityYear="1841" box="[1134,1436,562,584]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="560" phylum="Arthropoda" rank="species" species="westwoodii">
|
||
<emphasis id="528E8801137DFFEEFBFADB3DFA24FDC9" bold="true" box="[1134,1436,562,584]" pageId="13" pageNumber="560">
|
||
<emphasis id="528E8801137DFFEEFBFADB3DFB49FDC6" bold="true" box="[1134,1265,562,584]" italics="true" pageId="13" pageNumber="560">P. westwoodii</emphasis>
|
||
<bibRefCitation id="046B29E2137DFFEEFB63DB3DFA24FDC9" author="Westwood, J. O." box="[1271,1436,562,583]" pageId="13" pageNumber="560" pagination="34 - 35" refId="ref8510" refString="Westwood, J. O. (1841) On the Australian genus Cryptodu s, and upon Parastasia, the Asiatic representative of the Rutelidae. Journal of Proceedings of the Entomological Society of London, 2: 34 - 35." type="journal article" year="1841">Westwood, 1841</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="DB0E4CB6137DFFEEFF03DB5FFA24FC9A" pageId="13" pageNumber="560">
|
||
<paragraph id="60455413137DFFEEFF03DB5FFA24FD11" blockId="13.[151,1437,150,1141]" pageId="13" pageNumber="560">
|
||
<keyLead id="DB0BF726137DFFEEFF03DB5FFA24FD11" pageId="13" pageNumber="560">
|
||
7 Ground color of dorsal surface reddish, elytra unicolor (
|
||
<figureCitation id="F8C14896137DFFEEFD63DB40FC81FDEB" box="[759,825,591,613]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1864,1888]" captionTargetBox="[277,1315,249,1829]" captionTargetId="figure-34@3.[255,1332,234,1837]" captionTargetPageId="3" captionText="FIGURES 1–5. Parastasia spinosa Hongsuwong, Sanguansub & Jaitrong, new species (holotype, male, THNHM-I-09998). 1, habitus in dorsal view; 2, habitus in ventral view; 3, habitus in lateral view; 4, habitus in caudal view; 5, inner side of right protarsus." figureDoi="http://doi.org/10.5281/zenodo.7318722" httpUri="https://zenodo.org/record/7318722/files/figure.png" pageId="13" pageNumber="560">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="F8C14896137DFFEEFCD1DB40FCD8FDEB" box="[837,864,591,613]" captionStart="FIGURES 51–60" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[159,1424,192,1495]" captionTargetId="figure-18@11.[151,1436,181,1495]" captionTargetPageId="11" captionText="FIGURES 51–60. Characteristics of Parastasia species, non-type specimens. 51, Parastasia birmana Arrow, 1899, male (THNHM-I-09992); 52, 56, 59, Parastasia anomala Arrow, 1899, male (THNHM-I-09977); 53, Parastasia westwoodii Westwood, 1841, female (CTH); 54, Parastasia masumotoi Wada & Muramoto, 1999, male (CTH); 55, Parastasia sulcipennis Gestro, 1888, male (CTH); 57, 60, Parastasia bigibbosa Nonfried, 1891, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus." figureDoi="http://doi.org/10.5281/zenodo.7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="13" pageNumber="560">55</figureCitation>
|
||
); external margin of parameres sinuate in dorsal view (
|
||
<figureCitation id="F8C14896137DFFEEFAF8DB40FF44FD0C" captionStart="FIGURES 6–16" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1385,193,1911]" captionTargetId="figure-18@4.[202,1385,181,1919]" captionTargetPageId="4" captionText="FIGURES 6–16. Mouthparts and male genitalia of Parastasia spinosa Hongsuwong, Sanguansub & Jaitrong, new species (holotype, male, THNHM-I-09998). 6, 7, labrum; 8, 9, left mandible; 10, 11 maxillae; 12, mentum; 13, 14, 15, male parameres; 16, endophallus; 6, 8, 10, 13, dorsal view; 7, 9, 11, 12, 14, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7318726" httpUri="https://zenodo.org/record/7318726/files/figure.png" pageId="13" pageNumber="560">Figs. 13</figureCitation>
|
||
,
|
||
<figureCitation id="F8C14896137DFFEEFE92DB62FE99FD0D" box="[262,289,621,643]" captionStart="FIGURES 61–70" captionStartId="12.[151,264,1853,1877]" captionTargetBox="[157,1420,190,1829]" captionTargetId="figure-18@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURES 61–70.Male genitalia of Parastasia species, non-type specimens.61–62, Parastasia birmanaArrow, 1899(THNHM- I-09992); 63–64, 69 Parastasia masumotoi Wada & Muramoto, 1999 (CTH); 65–66, 70, Parastasia sulcipennis Gestro, 1888 (CTH); 67, Parastasia anomala Arrow, 1899 (THNHM-I-09977); 68, Parastasia bigibbosa Nonfried, 1891 (CUT); 61, 63, 65, parameres in dorsal view; 62, 64, 66, aedeagus in lateral view; 67–70, endophallus." figureDoi="http://doi.org/10.5281/zenodo.7318740" httpUri="https://zenodo.org/record/7318740/files/figure.png" pageId="13" pageNumber="560">65</figureCitation>
|
||
); in lateral view, ventral margin of parameres plain without distinct concavity (
|
||
<figureCitation id="F8C14896137DFFEEFB8CDB62FBD1FD0D" box="[1048,1129,621,643]" captionStart="FIGURES 6–16" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1385,193,1911]" captionTargetId="figure-18@4.[202,1385,181,1919]" captionTargetPageId="4" captionText="FIGURES 6–16. Mouthparts and male genitalia of Parastasia spinosa Hongsuwong, Sanguansub & Jaitrong, new species (holotype, male, THNHM-I-09998). 6, 7, labrum; 8, 9, left mandible; 10, 11 maxillae; 12, mentum; 13, 14, 15, male parameres; 16, endophallus; 6, 8, 10, 13, dorsal view; 7, 9, 11, 12, 14, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7318726" httpUri="https://zenodo.org/record/7318726/files/figure.png" pageId="13" pageNumber="560">Figs. 15</figureCitation>
|
||
,
|
||
<figureCitation id="F8C14896137DFFEEFBE7DB62FB36FD0D" box="[1139,1166,621,643]" captionStart="FIGURES 61–70" captionStartId="12.[151,264,1853,1877]" captionTargetBox="[157,1420,190,1829]" captionTargetId="figure-18@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURES 61–70.Male genitalia of Parastasia species, non-type specimens.61–62, Parastasia birmanaArrow, 1899(THNHM- I-09992); 63–64, 69 Parastasia masumotoi Wada & Muramoto, 1999 (CTH); 65–66, 70, Parastasia sulcipennis Gestro, 1888 (CTH); 67, Parastasia anomala Arrow, 1899 (THNHM-I-09977); 68, Parastasia bigibbosa Nonfried, 1891 (CUT); 61, 63, 65, parameres in dorsal view; 62, 64, 66, aedeagus in lateral view; 67–70, endophallus." figureDoi="http://doi.org/10.5281/zenodo.7318740" httpUri="https://zenodo.org/record/7318740/files/figure.png" pageId="13" pageNumber="560">66</figureCitation>
|
||
); anterior portion of median sac of endophallus with small spines, remaining followed by large spines (
|
||
<figureCitation id="F8C14896137DFFEEFC38DB85FC45FD2E" box="[940,1021,650,672]" captionStart="FIGURES 6–16" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1385,193,1911]" captionTargetId="figure-18@4.[202,1385,181,1919]" captionTargetPageId="4" captionText="FIGURES 6–16. Mouthparts and male genitalia of Parastasia spinosa Hongsuwong, Sanguansub & Jaitrong, new species (holotype, male, THNHM-I-09998). 6, 7, labrum; 8, 9, left mandible; 10, 11 maxillae; 12, mentum; 13, 14, 15, male parameres; 16, endophallus; 6, 8, 10, 13, dorsal view; 7, 9, 11, 12, 14, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7318726" httpUri="https://zenodo.org/record/7318726/files/figure.png" pageId="13" pageNumber="560">Figs. 16</figureCitation>
|
||
,
|
||
<figureCitation id="F8C14896137DFFEEFB93DB85FB9AFD11" box="[1031,1058,650,671]" captionStart="FIGURES 61–70" captionStartId="12.[151,264,1853,1877]" captionTargetBox="[157,1420,190,1829]" captionTargetId="figure-18@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURES 61–70.Male genitalia of Parastasia species, non-type specimens.61–62, Parastasia birmanaArrow, 1899(THNHM- I-09992); 63–64, 69 Parastasia masumotoi Wada & Muramoto, 1999 (CTH); 65–66, 70, Parastasia sulcipennis Gestro, 1888 (CTH); 67, Parastasia anomala Arrow, 1899 (THNHM-I-09977); 68, Parastasia bigibbosa Nonfried, 1891 (CUT); 61, 63, 65, parameres in dorsal view; 62, 64, 66, aedeagus in lateral view; 67–70, endophallus." figureDoi="http://doi.org/10.5281/zenodo.7318740" httpUri="https://zenodo.org/record/7318740/files/figure.png" pageId="13" pageNumber="560">70</figureCitation>
|
||
)..............................
|
||
<emphasis id="528E8801137DFFEEFA04DB85FA24FD11" bold="true" box="[1424,1436,650,671]" pageId="13" pageNumber="560">8</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="60455413137DFFEEFF03DBA7FA24FC9A" blockId="13.[151,1437,150,1141]" pageId="13" pageNumber="560">
|
||
<keyLead id="DB0BF726137DFFEEFF03DBA7FA24FC9A" pageId="13" pageNumber="560">
|
||
- Ground color of dorsal surface orange, external and posterior portions of elytra black (
|
||
<figureCitation id="F8C14896137DFFEEFBD5DBA8FB33FD33" box="[1089,1163,679,701]" captionStart="FIGURES 51–60" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[159,1424,192,1495]" captionTargetId="figure-18@11.[151,1436,181,1495]" captionTargetPageId="11" captionText="FIGURES 51–60. Characteristics of Parastasia species, non-type specimens. 51, Parastasia birmana Arrow, 1899, male (THNHM-I-09992); 52, 56, 59, Parastasia anomala Arrow, 1899, male (THNHM-I-09977); 53, Parastasia westwoodii Westwood, 1841, female (CTH); 54, Parastasia masumotoi Wada & Muramoto, 1999, male (CTH); 55, Parastasia sulcipennis Gestro, 1888, male (CTH); 57, 60, Parastasia bigibbosa Nonfried, 1891, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus." figureDoi="http://doi.org/10.5281/zenodo.7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="13" pageNumber="560">Fig. 54</figureCitation>
|
||
); in dorsal view, parameres without concavity (
|
||
<figureCitation id="F8C14896137DFFEEFE0FDBCAFE5AFD55" box="[411,482,709,731]" captionStart="FIGURES 61–70" captionStartId="12.[151,264,1853,1877]" captionTargetBox="[157,1420,190,1829]" captionTargetId="figure-18@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURES 61–70.Male genitalia of Parastasia species, non-type specimens.61–62, Parastasia birmanaArrow, 1899(THNHM- I-09992); 63–64, 69 Parastasia masumotoi Wada & Muramoto, 1999 (CTH); 65–66, 70, Parastasia sulcipennis Gestro, 1888 (CTH); 67, Parastasia anomala Arrow, 1899 (THNHM-I-09977); 68, Parastasia bigibbosa Nonfried, 1891 (CUT); 61, 63, 65, parameres in dorsal view; 62, 64, 66, aedeagus in lateral view; 67–70, endophallus." figureDoi="http://doi.org/10.5281/zenodo.7318740" httpUri="https://zenodo.org/record/7318740/files/figure.png" pageId="13" pageNumber="560">Fig. 63</figureCitation>
|
||
); in lateral view, ventral margin of parameres thickly marginated and deeply concave one-fifth from base (
|
||
<figureCitation id="F8C14896137DFFEEFE8DDBEDFED8FD76" box="[281,352,738,760]" captionStart="FIGURES 61–70" captionStartId="12.[151,264,1853,1877]" captionTargetBox="[157,1420,190,1829]" captionTargetId="figure-18@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURES 61–70.Male genitalia of Parastasia species, non-type specimens.61–62, Parastasia birmanaArrow, 1899(THNHM- I-09992); 63–64, 69 Parastasia masumotoi Wada & Muramoto, 1999 (CTH); 65–66, 70, Parastasia sulcipennis Gestro, 1888 (CTH); 67, Parastasia anomala Arrow, 1899 (THNHM-I-09977); 68, Parastasia bigibbosa Nonfried, 1891 (CUT); 61, 63, 65, parameres in dorsal view; 62, 64, 66, aedeagus in lateral view; 67–70, endophallus." figureDoi="http://doi.org/10.5281/zenodo.7318740" httpUri="https://zenodo.org/record/7318740/files/figure.png" pageId="13" pageNumber="560">Fig. 64</figureCitation>
|
||
); anterior portion of median sac of endophallus with few large spines, remaining followed by medium spines (
|
||
<figureCitation id="F8C14896137DFFEEFAE1DBEDFF45FC9B" captionStart="FIGURES 61–70" captionStartId="12.[151,264,1853,1877]" captionTargetBox="[157,1420,190,1829]" captionTargetId="figure-18@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURES 61–70.Male genitalia of Parastasia species, non-type specimens.61–62, Parastasia birmanaArrow, 1899(THNHM- I-09992); 63–64, 69 Parastasia masumotoi Wada & Muramoto, 1999 (CTH); 65–66, 70, Parastasia sulcipennis Gestro, 1888 (CTH); 67, Parastasia anomala Arrow, 1899 (THNHM-I-09977); 68, Parastasia bigibbosa Nonfried, 1891 (CUT); 61, 63, 65, parameres in dorsal view; 62, 64, 66, aedeagus in lateral view; 67–70, endophallus." figureDoi="http://doi.org/10.5281/zenodo.7318740" httpUri="https://zenodo.org/record/7318740/files/figure.png" pageId="13" pageNumber="560">Fig. 69</figureCitation>
|
||
).................................................................
|
||
<taxonomicName id="A7FA2F90137DFFEEFB9FDA0FFA24FC9A" authority="Wada & Muramoto, 1999" authorityName="Wada & Muramoto" authorityYear="1999" box="[1035,1436,767,789]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="560" phylum="Arthropoda" rank="species" species="masumotoi">
|
||
<emphasis id="528E8801137DFFEEFB9FDA0FFA24FC9A" bold="true" box="[1035,1436,767,789]" pageId="13" pageNumber="560">
|
||
<emphasis id="528E8801137DFFEEFB9FDA0FFB37FC9A" bold="true" box="[1035,1167,767,789]" italics="true" pageId="13" pageNumber="560">P. masumotoi</emphasis>
|
||
<bibRefCitation id="046B29E2137DFFEEFB02DA0FFA24FC9A" author="Wada K. & Muramoto R." box="[1174,1436,767,789]" pageId="13" pageNumber="560" pagination="167 - 175" refId="ref8478" refString="Wada K. & Muramoto R. (1999) Three New Species of the genus Parastasia from Southeast Asia. Elytra, 27 (1): 167 - 175." type="journal article" year="1999">Wada & Muramoto, 1999</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="DB0E4CB6137DFFEEFF03DA12FA24FC04" pageId="13" pageNumber="560">
|
||
<paragraph id="60455413137DFFEEFF03DA12FA24FCC1" blockId="13.[151,1437,150,1141]" pageId="13" pageNumber="560">
|
||
<keyLead id="DB0BF726137DFFEEFF03DA12FA24FCC1" pageId="13" pageNumber="560">
|
||
8 Scutellum and ventral surface reddish to dark reddish brown (
|
||
<figureCitation id="F8C14896137DFFEEFCB2DA12FC38FCBC" box="[806,896,797,819]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1864,1888]" captionTargetBox="[277,1315,249,1829]" captionTargetId="figure-34@3.[255,1332,234,1837]" captionTargetPageId="3" captionText="FIGURES 1–5. Parastasia spinosa Hongsuwong, Sanguansub & Jaitrong, new species (holotype, male, THNHM-I-09998). 1, habitus in dorsal view; 2, habitus in ventral view; 3, habitus in lateral view; 4, habitus in caudal view; 5, inner side of right protarsus." figureDoi="http://doi.org/10.5281/zenodo.7318722" httpUri="https://zenodo.org/record/7318722/files/figure.png" pageId="13" pageNumber="560">Figs. 1–2</figureCitation>
|
||
,
|
||
<figureCitation id="F8C14896137DFFEEFC1DDA12FC1CFCBD" box="[905,932,797,819]" captionStart="FIGURES 51–60" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[159,1424,192,1495]" captionTargetId="figure-18@11.[151,1436,181,1495]" captionTargetPageId="11" captionText="FIGURES 51–60. Characteristics of Parastasia species, non-type specimens. 51, Parastasia birmana Arrow, 1899, male (THNHM-I-09992); 52, 56, 59, Parastasia anomala Arrow, 1899, male (THNHM-I-09977); 53, Parastasia westwoodii Westwood, 1841, female (CTH); 54, Parastasia masumotoi Wada & Muramoto, 1999, male (CTH); 55, Parastasia sulcipennis Gestro, 1888, male (CTH); 57, 60, Parastasia bigibbosa Nonfried, 1891, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus." figureDoi="http://doi.org/10.5281/zenodo.7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="13" pageNumber="560">55</figureCitation>
|
||
); in lateral view, basal of ventral margin of parameres without protrusion.................................................................................... 9
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="60455413137DFFEEFF03DA57FA24FC04" blockId="13.[151,1437,150,1141]" pageId="13" pageNumber="560">
|
||
<keyLead id="DB0BF726137DFFEEFF03DA57FA24FC04" pageId="13" pageNumber="560">
|
||
- Scutellum and ventral surface nearly black; in lateral view, basal of ventral margin of parameres clearly protruding backward, process sharp and elongate as spinose formed.................................
|
||
<taxonomicName id="A7FA2F90137DFFEEFBBFDA7AFA24FC04" authority="Kuijten, 1992" authorityName="Kuijten" authorityYear="1992" box="[1067,1436,884,907]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="560" phylum="Arthropoda" rank="species" species="selangorica">
|
||
<emphasis id="528E8801137DFFEEFBBFDA7AFA24FC04" bold="true" box="[1067,1436,884,907]" pageId="13" pageNumber="560">
|
||
<emphasis id="528E8801137DFFEEFBBFDA7AFAB4FC04" bold="true" box="[1067,1292,884,906]" italics="true" pageId="13" pageNumber="560">Parastasia selangorica</emphasis>
|
||
<bibRefCitation id="046B29E2137DFFEEFA86DA7AFA24FC04" author="Kuijten, P. J." box="[1298,1436,885,907]" pageId="13" pageNumber="560" pagination="1 - 207" refId="ref7465" refString="Kuijten, P. J. (1992) A revision of the genus Parastasia in the Indo - Australian region (Coleoptera: Scarabaeidae, Rutelinae). Zoologische Verhandelingen, 275, 1 - 207." type="journal article" year="1992">Kuijten, 1992</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="DB0E4CB6137DFFEEFF03DA9DFA24FBFA" pageId="13" pageNumber="560">
|
||
<paragraph id="60455413137DFFEEFF03DA9DFA25FB8E" blockId="13.[151,1437,150,1141]" pageId="13" pageNumber="560">
|
||
<keyLead id="DB0BF726137DFFEEFF03DA9DFA25FB8E" pageId="13" pageNumber="560">
|
||
9 Rather small in size (
|
||
<figureCitation id="F8C14896137DFFEEFE33DA9DFE65FC29" box="[423,477,914,936]" captionStart="FIGURES 1–5" captionStartId="3.[151,264,1864,1888]" captionTargetBox="[277,1315,249,1829]" captionTargetId="figure-34@3.[255,1332,234,1837]" captionTargetPageId="3" captionText="FIGURES 1–5. Parastasia spinosa Hongsuwong, Sanguansub & Jaitrong, new species (holotype, male, THNHM-I-09998). 1, habitus in dorsal view; 2, habitus in ventral view; 3, habitus in lateral view; 4, habitus in caudal view; 5, inner side of right protarsus." figureDoi="http://doi.org/10.5281/zenodo.7318722" httpUri="https://zenodo.org/record/7318722/files/figure.png" pageId="13" pageNumber="560">Fig. 1</figureCitation>
|
||
); first row of punctures on elytra sinuate; metasternal process stout; in dorsal view, apex of parameres roundly convex (
|
||
<figureCitation id="F8C14896137DFFEEFE11DAA0FE74FC4B" box="[389,460,943,965]" captionStart="FIGURES 6–16" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1385,193,1911]" captionTargetId="figure-18@4.[202,1385,181,1919]" captionTargetPageId="4" captionText="FIGURES 6–16. Mouthparts and male genitalia of Parastasia spinosa Hongsuwong, Sanguansub & Jaitrong, new species (holotype, male, THNHM-I-09998). 6, 7, labrum; 8, 9, left mandible; 10, 11 maxillae; 12, mentum; 13, 14, 15, male parameres; 16, endophallus; 6, 8, 10, 13, dorsal view; 7, 9, 11, 12, 14, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7318726" httpUri="https://zenodo.org/record/7318726/files/figure.png" pageId="13" pageNumber="560">Fig. 13</figureCitation>
|
||
); in lateral view, dorsal and ventral margins almost straight (
|
||
<figureCitation id="F8C14896137DFFEEFB87DAA0FBE1FC4B" box="[1043,1113,943,965]" captionStart="FIGURES 6–16" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1385,193,1911]" captionTargetId="figure-18@4.[202,1385,181,1919]" captionTargetPageId="4" captionText="FIGURES 6–16. Mouthparts and male genitalia of Parastasia spinosa Hongsuwong, Sanguansub & Jaitrong, new species (holotype, male, THNHM-I-09998). 6, 7, labrum; 8, 9, left mandible; 10, 11 maxillae; 12, mentum; 13, 14, 15, male parameres; 16, endophallus; 6, 8, 10, 13, dorsal view; 7, 9, 11, 12, 14, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7318726" httpUri="https://zenodo.org/record/7318726/files/figure.png" pageId="13" pageNumber="560">Fig. 15</figureCitation>
|
||
); median sac of endophallus with a cluster of dense small spines anteriorly, posterior part of median sac with five dark large spines ventrally (
|
||
<figureCitation id="F8C14896137DFFEEFB7FDAC2FA8BFC6D" box="[1259,1331,973,995]" captionStart="FIGURES 6–16" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[214,1385,193,1911]" captionTargetId="figure-18@4.[202,1385,181,1919]" captionTargetPageId="4" captionText="FIGURES 6–16. Mouthparts and male genitalia of Parastasia spinosa Hongsuwong, Sanguansub & Jaitrong, new species (holotype, male, THNHM-I-09998). 6, 7, labrum; 8, 9, left mandible; 10, 11 maxillae; 12, mentum; 13, 14, 15, male parameres; 16, endophallus; 6, 8, 10, 13, dorsal view; 7, 9, 11, 12, 14, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7318726" httpUri="https://zenodo.org/record/7318726/files/figure.png" pageId="13" pageNumber="560">Fig. 16</figureCitation>
|
||
).......................................................
|
||
<emphasis id="528E8801137DFFEEFCB9DAE5FA25FB8E" bold="true" box="[813,1437,1002,1024]" pageId="13" pageNumber="560">
|
||
<taxonomicName id="A7FA2F90137DFFEEFCB9DAE5FAA4FB8E" authority="Hongsuwong, Sanguansub & Jaitrong" authorityName="Hongsuwong, Sanguansub & Jaitrong" authorityYear="2022" box="[813,1308,1002,1024]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="560" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
|
||
<emphasis id="528E8801137DFFEEFCB9DAE5FC28FC71" bold="true" box="[813,912,1002,1023]" italics="true" pageId="13" pageNumber="560">P. spinosa</emphasis>
|
||
Hongsuwong, Sanguansub & Jaitrong
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="49BD357A137DFFEEFAB2DAE5FA25FB8E" box="[1318,1437,1002,1024]" pageId="13" pageNumber="560" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="60455413137DFFEEFF03DD07FA24FBFA" blockId="13.[151,1437,150,1141]" pageId="13" pageNumber="560">
|
||
<keyLead id="DB0BF726137DFFEEFF03DD07FA24FBFA" pageId="13" pageNumber="560">
|
||
- Medium in size (
|
||
<figureCitation id="F8C14896137DFFEEFE16DD08FE71FB93" box="[386,457,1031,1053]" captionStart="FIGURES 51–60" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[159,1424,192,1495]" captionTargetId="figure-18@11.[151,1436,181,1495]" captionTargetPageId="11" captionText="FIGURES 51–60. Characteristics of Parastasia species, non-type specimens. 51, Parastasia birmana Arrow, 1899, male (THNHM-I-09992); 52, 56, 59, Parastasia anomala Arrow, 1899, male (THNHM-I-09977); 53, Parastasia westwoodii Westwood, 1841, female (CTH); 54, Parastasia masumotoi Wada & Muramoto, 1999, male (CTH); 55, Parastasia sulcipennis Gestro, 1888, male (CTH); 57, 60, Parastasia bigibbosa Nonfried, 1891, male (THNHM-I-09982); 51–55, habitus in dorsal view; 56–57, head in dorsal view; 58–60, mesotarsus." figureDoi="http://doi.org/10.5281/zenodo.7318738" httpUri="https://zenodo.org/record/7318738/files/figure.png" pageId="13" pageNumber="560">Fig. 55</figureCitation>
|
||
); first row of punctures on elytra mostly straight; metasternal process sharply produced; in dorsal view, apex of parameres almost truncate with a small carina near internal margin (
|
||
<figureCitation id="F8C14896137DFFEEFC46DD2AFBA4FBB5" box="[978,1052,1061,1083]" captionStart="FIGURES 61–70" captionStartId="12.[151,264,1853,1877]" captionTargetBox="[157,1420,190,1829]" captionTargetId="figure-18@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURES 61–70.Male genitalia of Parastasia species, non-type specimens.61–62, Parastasia birmanaArrow, 1899(THNHM- I-09992); 63–64, 69 Parastasia masumotoi Wada & Muramoto, 1999 (CTH); 65–66, 70, Parastasia sulcipennis Gestro, 1888 (CTH); 67, Parastasia anomala Arrow, 1899 (THNHM-I-09977); 68, Parastasia bigibbosa Nonfried, 1891 (CUT); 61, 63, 65, parameres in dorsal view; 62, 64, 66, aedeagus in lateral view; 67–70, endophallus." figureDoi="http://doi.org/10.5281/zenodo.7318740" httpUri="https://zenodo.org/record/7318740/files/figure.png" pageId="13" pageNumber="560">Fig. 65</figureCitation>
|
||
); in lateral view, dorsal margin weakly concave, while ventral margin convex (
|
||
<figureCitation id="F8C14896137DFFEEFDF9DD4DFD17FBD6" box="[621,687,1090,1112]" captionStart="FIGURES 61–70" captionStartId="12.[151,264,1853,1877]" captionTargetBox="[157,1420,190,1829]" captionTargetId="figure-18@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURES 61–70.Male genitalia of Parastasia species, non-type specimens.61–62, Parastasia birmanaArrow, 1899(THNHM- I-09992); 63–64, 69 Parastasia masumotoi Wada & Muramoto, 1999 (CTH); 65–66, 70, Parastasia sulcipennis Gestro, 1888 (CTH); 67, Parastasia anomala Arrow, 1899 (THNHM-I-09977); 68, Parastasia bigibbosa Nonfried, 1891 (CUT); 61, 63, 65, parameres in dorsal view; 62, 64, 66, aedeagus in lateral view; 67–70, endophallus." figureDoi="http://doi.org/10.5281/zenodo.7318740" httpUri="https://zenodo.org/record/7318740/files/figure.png" pageId="13" pageNumber="560">Fig.66</figureCitation>
|
||
); median sac of endophallus with sparse small spines anteriorly, ventral of posterior part without dominantly large spines (
|
||
<figureCitation id="F8C14896137DFFEEFD3EDD50FD4AFBFB" box="[682,754,1119,1141]" captionStart="FIGURES 61–70" captionStartId="12.[151,264,1853,1877]" captionTargetBox="[157,1420,190,1829]" captionTargetId="figure-18@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURES 61–70.Male genitalia of Parastasia species, non-type specimens.61–62, Parastasia birmanaArrow, 1899(THNHM- I-09992); 63–64, 69 Parastasia masumotoi Wada & Muramoto, 1999 (CTH); 65–66, 70, Parastasia sulcipennis Gestro, 1888 (CTH); 67, Parastasia anomala Arrow, 1899 (THNHM-I-09977); 68, Parastasia bigibbosa Nonfried, 1891 (CUT); 61, 63, 65, parameres in dorsal view; 62, 64, 66, aedeagus in lateral view; 67–70, endophallus." figureDoi="http://doi.org/10.5281/zenodo.7318740" httpUri="https://zenodo.org/record/7318740/files/figure.png" pageId="13" pageNumber="560">Fig. 70</figureCitation>
|
||
)....................................
|
||
<taxonomicName id="A7FA2F90137DFFEEFB31DD50FA24FBFA" authority="Gestro, 1888" authorityName="Gestro" authorityYear="1888" box="[1189,1436,1119,1141]" class="Insecta" family="Rutelidae" genus="Parastasia" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="560" phylum="Arthropoda" rank="species" species="sulcipennis">
|
||
<emphasis id="528E8801137DFFEEFB31DD50FA24FBFA" bold="true" box="[1189,1436,1119,1141]" pageId="13" pageNumber="560">
|
||
<emphasis id="528E8801137DFFEEFB31DD50FAADFBFB" bold="true" box="[1189,1301,1119,1141]" italics="true" pageId="13" pageNumber="560">sulcipennis</emphasis>
|
||
Gestro, 1888
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |