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<document id="6079A6FC6CB742CC98DB58C9D18D713E" ID-CLB-Dataset="20588" ID-DOI="10.11646/zootaxa.5100.2.4" ID-GBIF-Dataset="21987729-d480-4305-8de5-9d38e4399e2d" ID-ISSN="1175-5326" ID-Zenodo-Dep="6145690" ID-ZooBank="A21EE081-4AE7-4129-814D-49605C8DC24D" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645171863462" checkinUser="plazi" docAuthor="Sapieja, Mateusz, Ranocha, Magdalena &amp; Melke, Andrzej" docDate="2022" docId="E8516E6BFFBBDD0F148BF8BB36DD43D7" docLanguage="en" docName="zootaxa.5100.2.4.pdf" docOrigin="Zootaxa 5100 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rugilus Leach 1819" docType="key" docVersion="7" lastPageNumber="256" masterDocId="14681613FFBDDD08141CFFA833414240" masterDocTitle="Distribution of Rugilus mixtus (Lohse, 1956) (Coleoptera: Staphylinidae: Paederinae) in Poland with comments of its morphology and ecological preferences" masterLastPageNumber="258" masterPageNumber="249" pageNumber="255" updateTime="1699314528725" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="358DCFA0704A5FF39C1382A014D45712">Distribution of Rugilus mixtus (Lohse, 1956) (Coleoptera: Staphylinidae: Paederinae) in Poland with comments of its morphology and ecological preferences</mods:title>
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<mods:namePart id="1A22371EF129F649954076A1F47A72AD">Sapieja, Mateusz</mods:namePart>
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<mods:affiliation id="CF5A2DAA1FED7868F19E0836BCFB2224">Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387 Kraków, Poland. sapiejamateusz @ gmail. com, https: // orcid. org / 0000 - 0003 - 1151 - 4245</mods:affiliation>
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<mods:namePart id="D2E5F0E01E15BB203BBD36A641151CEA">Ranocha, Magdalena</mods:namePart>
<mods:affiliation id="80FDBB09A9CD407D9F904D25A23E19FE">Department of Mountain Forests, Forest Research Institute, Fredry 39, 30 - 605 Kraków, Poland.</mods:affiliation>
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<mods:namePart id="41F2AB63C4C1613FBF855157C0E4C3A3">Melke, Andrzej</mods:namePart>
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<mods:affiliation id="1DE80D76AACE1E05EC919695E65F50DF">Department of Zoology, Institute of Biology and Environmental Protection, Pomeranian University, 76 - 200 Słupsk. kusakowaty @ gmail. com, https: // orcid. org / 0000 - 0002 - 2039 - 7979</mods:affiliation>
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<treatment id="E8516E6BFFBBDD0F148BF8BB36DD43D7" ID-DOI="http://doi.org/10.5281/zenodo.6313402" ID-GBIF-Taxon="193399341" ID-Zenodo-Dep="6313402" LSID="urn:lsid:plazi:treatment:E8516E6BFFBBDD0F148BF8BB36DD43D7" httpUri="http://treatment.plazi.org/id/E8516E6BFFBBDD0F148BF8BB36DD43D7" lastPageId="7" lastPageNumber="256" pageId="6" pageNumber="255">
<subSubSection id="28E28CF6FFBBDD0E148BF8BB3600456D" box="[151,1345,1811,1838]" pageId="6" pageNumber="255" type="nomenclature">
<paragraph id="6047DF7DFFBBDD0E148BF8BB3600456D" blockId="6.[151,1345,1811,1838]" box="[151,1345,1811,1838]" pageId="6" pageNumber="255">
The identification key for central European representatives of the genus
<taxonomicName id="A7F8A4FEFFBBDD0E17DDF8BB3756456D" ID-CoL="8VS2Z" authorityName="Leach" authorityYear="1819" box="[961,1047,1811,1837]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="genus">
<emphasis id="528C036FFFBBDD0E17DDF8BB3756456D" box="[961,1047,1811,1837]" italics="true" pageId="6" pageNumber="255">Rugilus</emphasis>
</taxonomicName>
with yellow-tipped elytra:
</paragraph>
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<subSubSection id="28E28CF6FFBBDD0F148BF8FA36DD43D7" lastPageId="7" lastPageNumber="256" pageId="6" pageNumber="255" type="key">
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<paragraph id="6047DF7DFFBBDD0E148BF8FA36DC4527" blockId="6.[151,1437,1874,2042]" box="[151,1437,1874,1896]" pageId="6" pageNumber="255">
<keyLead id="DB097C48FFBBDD0E148BF8FA36DC4527" box="[151,1437,1874,1896]" pageId="6" pageNumber="255">
1. Body length not smaller than 5.0 mm................................................................
<taxonomicName id="A7F8A4FEFFBBDD0E115FF8FB36DC4527" ID-CoL="4TNXQ" baseAuthorityName="Erichson" baseAuthorityYear="1839" box="[1347,1437,1874,1896]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="528C036FFFBBDD0E115FF8FB36DC4527" box="[1347,1437,1874,1896]" italics="true" pageId="6" pageNumber="255">R. similis</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="6047DF7DFFBBDD0E148BF8D836DC45C4" blockId="6.[151,1437,1874,2042]" box="[151,1437,1903,1925]" pageId="6" pageNumber="255">
<keyLead id="DB097C48FFBBDD0E148BF8D836DC45C4" box="[151,1437,1903,1925]" pageId="6" pageNumber="255"> Body length smaller than 5.0 mm..........................................................................2</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB0CC7D8FFBBDD0E148BF82536DD45BA" pageId="6" pageNumber="255">
<paragraph id="6047DF7DFFBBDD0E148BF82536DD459C" blockId="6.[151,1437,1874,2042]" pageId="6" pageNumber="255">
<keyLead id="DB097C48FFBBDD0E148BF82536DD459C" pageId="6" pageNumber="255">
2. Head oval, wider than elytra (HI—1:1.13~1.21, HEI—1: 1.061.10). Eye twice as long as temple (ETI—1.962.14). Elytra black, sometimes with a dark maroon tinge. Legs yellow. Male abdominal sternite VIII with two pairs of long symmetrically set bristles in the middle of a segment and posterior margin (
<figureCitation id="F8C3C3F8FFBBDD0E1734F86F3022459D" box="[808,867,1991,2013]" captionStart="FIGURES 9-14" captionStartId="5.[151,264,1933,1957]" captionTargetBox="[292,1287,364,1897]" captionTargetId="figure-86@5.[291,1296,345,1910]" captionTargetPageId="5" captionText="FIGURES 9-14. VII sternite (9, 10), right metatarsi (11, 12) and right antennae (13, 14); Rugilus erichsoni (9), Rugilus orbiculatus (10, 12, 14), Rugilus mixtus (11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6145700" httpUri="https://zenodo.org/record/6145700/files/figure.png" pageId="6" pageNumber="255">Fig. 9</figureCitation>
). Aedeagus as in
<figureCitation id="F8C3C3F8FFBBDD0E1017F86F3672459D" box="[1035,1331,1991,2013]" captionStart="FIGURES 15-20" captionStartId="6.[151,264,1669,1693]" captionTargetBox="[268,1306,189,1644]" captionTargetId="figure-24@6.[265,1322,181,1645]" captionTargetPageId="6" captionText="FIGURES 15-20. Aedeagus in ventral (15, 16, 17) and lateral (18, 19, 20) view; Rugilus mixtus (15, 18), Rugilus orbiculatus (16, 19), Rugilus erichsoni (17, 20)." figureDoi="http://doi.org/10.5281/zenodo.6145704" httpUri="https://zenodo.org/record/6145704/files/figure.png" pageId="6" pageNumber="255">
Fig. 17, 20...............
<taxonomicName id="A7F8A4FEFFBBDD0E113EF8603672459D" ID-CoL="4TNW9" baseAuthorityName="Fauvel" baseAuthorityYear="1867" box="[1314,1331,1992,2013]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="erichsonii">
<emphasis id="528C036FFFBBDD0E113EF8603672459D" box="[1314,1331,1992,2013]" italics="true" pageId="6" pageNumber="255">R</emphasis>
</taxonomicName>
</figureCitation>
. erichsonii
</keyLead>
</paragraph>
<paragraph id="6047DF7DFFBBDD0E148BF84D36DD45BA" blockId="6.[151,1437,1874,2042]" box="[151,1436,2020,2042]" pageId="6" pageNumber="255">
<keyLead id="DB097C48FFBBDD0E148BF84D36DD45BA" box="[151,1436,2020,2042]" pageId="6" pageNumber="255"> Head width equal to or less than elytra width, eye shorter than two temple lengths. Aedeagus different................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="DB0CC7D8FFBADD0F148BFF3036DD43D7" pageId="7" pageNumber="256">
<paragraph id="6047DF7DFFBADD0F148BFF3036DC4345" blockId="7.[151,1437,151,408]" pageId="7" pageNumber="256">
<keyLead id="DB097C48FFBADD0F148BFF3036DC4345" pageId="7" pageNumber="256">
3. Head oval, wider than long (HI ranges 1.131.14), as wide as or slightly narrower than elytra (HEI—1:0.97). Fourth segment of antennae clavate (4ASI—1:0.830.93) (
<figureCitation id="F8C3C3F8FFBADD0F1665FF1D3181428B" box="[633,704,181,203]" captionStart="FIGURES 9-14" captionStartId="5.[151,264,1933,1957]" captionTargetBox="[292,1287,364,1897]" captionTargetId="figure-86@5.[291,1296,345,1910]" captionTargetPageId="5" captionText="FIGURES 9-14. VII sternite (9, 10), right metatarsi (11, 12) and right antennae (13, 14); Rugilus erichsoni (9), Rugilus orbiculatus (10, 12, 14), Rugilus mixtus (11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6145700" httpUri="https://zenodo.org/record/6145700/files/figure.png" pageId="7" pageNumber="256">Fig. 13</figureCitation>
). Eye almost twice as long as temple (ETI—1.831.87). Elytra dark maroon. Legs maroon. Length index of the first posterior tarsal segment to its total length 1:0.280.29 (
<figureCitation id="F8C3C3F8FFBADD0F1043FF7A37E542A8" box="[1119,1188,210,232]" captionStart="FIGURES 9-14" captionStartId="5.[151,264,1933,1957]" captionTargetBox="[292,1287,364,1897]" captionTargetId="figure-86@5.[291,1296,345,1910]" captionTargetPageId="5" captionText="FIGURES 9-14. VII sternite (9, 10), right metatarsi (11, 12) and right antennae (13, 14); Rugilus erichsoni (9), Rugilus orbiculatus (10, 12, 14), Rugilus mixtus (11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6145700" httpUri="https://zenodo.org/record/6145700/files/figure.png" pageId="7" pageNumber="256">Fig. 11</figureCitation>
). Male abdominal sternite VIII triangularly cut (
<bibRefCitation id="0469A28CFFBADD0F15ADFF4731664345" author="Lohse, G. A." box="[433,551,239,261]" pageId="7" pageNumber="256" pagination="48 - 61" refId="ref6664" refString="Lohse, G. A. (1956) Neuheiten der deutschen Kaferfauna III. Entomologische Blatter fur Biologie und Systematik der Kafer, 52 (1 - 2), 48 - 61." type="journal article" year="1956">Lohse 1956</bibRefCitation>
). Aedeagus as in
<figureCitation id="F8C3C3F8FFBADD0F16D1FF4736154345" box="[717,1364,239,261]" captionStart="FIGURES 15-20" captionStartId="6.[151,264,1669,1693]" captionTargetBox="[268,1306,189,1644]" captionTargetId="figure-24@6.[265,1322,181,1645]" captionTargetPageId="6" captionText="FIGURES 15-20. Aedeagus in ventral (15, 16, 17) and lateral (18, 19, 20) view; Rugilus mixtus (15, 18), Rugilus orbiculatus (16, 19), Rugilus erichsoni (17, 20)." figureDoi="http://doi.org/10.5281/zenodo.6145704" httpUri="https://zenodo.org/record/6145704/files/figure.png" pageId="7" pageNumber="256">
Figs. 15, 18............................................
<taxonomicName id="A7F8A4FEFFBADD0F115FFF5836154345" ID-CoL="4TNWZ" baseAuthorityName="Lohse" baseAuthorityYear="1956" box="[1347,1364,240,261]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="256" phylum="Arthropoda" rank="species" species="mixtus">
<emphasis id="528C036FFFBADD0F115FFF5836154345" box="[1347,1364,240,261]" italics="true" pageId="7" pageNumber="256">R</emphasis>
</taxonomicName>
</figureCitation>
. mixtus
</keyLead>
</paragraph>
<paragraph id="6047DF7DFFBADD0F148BFEA536DD43D7" blockId="7.[151,1437,151,408]" pageId="7" pageNumber="256">
<keyLead id="DB097C48FFBADD0F148BFEA536DD43D7" pageId="7" pageNumber="256">
Head slightly wider than length (HI ranges 1:1.061.09) and much narrower than elytra (HEI—1:0.860.90). Fourth antennal segment barrel-shaped (4ASI—1:0.720.81) (
<figureCitation id="F8C3C3F8FFBADD0F16B9FE8231B04300" box="[677,753,298,320]" captionStart="FIGURES 9-14" captionStartId="5.[151,264,1933,1957]" captionTargetBox="[292,1287,364,1897]" captionTargetId="figure-86@5.[291,1296,345,1910]" captionTargetPageId="5" captionText="FIGURES 9-14. VII sternite (9, 10), right metatarsi (11, 12) and right antennae (13, 14); Rugilus erichsoni (9), Rugilus orbiculatus (10, 12, 14), Rugilus mixtus (11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6145700" httpUri="https://zenodo.org/record/6145700/files/figure.png" pageId="7" pageNumber="256">Fig. 14</figureCitation>
). Eye about 1.5 times longer than temple (ETI ranges 1:1.501.55). Elytra black with markedly contrasting yellow spots at posterior margins. Legs yellow. Length index of the first posterior tarsal segment to the total length 1:0.310.33 (
<figureCitation id="F8C3C3F8FFBADD0F1670FECD31F4433B" box="[620,693,357,379]" captionStart="FIGURES 9-14" captionStartId="5.[151,264,1933,1957]" captionTargetBox="[292,1287,364,1897]" captionTargetId="figure-86@5.[291,1296,345,1910]" captionTargetPageId="5" captionText="FIGURES 9-14. VII sternite (9, 10), right metatarsi (11, 12) and right antennae (13, 14); Rugilus erichsoni (9), Rugilus orbiculatus (10, 12, 14), Rugilus mixtus (11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6145700" httpUri="https://zenodo.org/record/6145700/files/figure.png" pageId="7" pageNumber="256">Fig. 12</figureCitation>
). Male abdominal sternite VIII rounded cut, with one pair symmetrically set long bristles at posterior margin (
<figureCitation id="F8C3C3F8FFBADD0F163DFE2A312843D8" box="[545,617,386,408]" captionStart="FIGURES 9-14" captionStartId="5.[151,264,1933,1957]" captionTargetBox="[292,1287,364,1897]" captionTargetId="figure-86@5.[291,1296,345,1910]" captionTargetPageId="5" captionText="FIGURES 9-14. VII sternite (9, 10), right metatarsi (11, 12) and right antennae (13, 14); Rugilus erichsoni (9), Rugilus orbiculatus (10, 12, 14), Rugilus mixtus (11, 13)." figureDoi="http://doi.org/10.5281/zenodo.6145700" httpUri="https://zenodo.org/record/6145700/files/figure.png" pageId="7" pageNumber="256">Fig. 10</figureCitation>
). Aedeagus as in
<figureCitation id="F8C3C3F8FFBADD0F170CFE2A366443D8" box="[784,1317,386,408]" captionStart="FIGURES 15-20" captionStartId="6.[151,264,1669,1693]" captionTargetBox="[268,1306,189,1644]" captionTargetId="figure-24@6.[265,1322,181,1645]" captionTargetPageId="6" captionText="FIGURES 15-20. Aedeagus in ventral (15, 16, 17) and lateral (18, 19, 20) view; Rugilus mixtus (15, 18), Rugilus orbiculatus (16, 19), Rugilus erichsoni (17, 20)." figureDoi="http://doi.org/10.5281/zenodo.6145704" httpUri="https://zenodo.org/record/6145704/files/figure.png" pageId="7" pageNumber="256">
Figs. 16, 19..................................
<taxonomicName id="A7F8A4FEFFBADD0F1108FE2B366443D8" ID-CoL="4TNX7" baseAuthorityName="Paykull" baseAuthorityYear="1789" box="[1300,1317,387,408]" class="Insecta" family="Staphylinidae" genus="Rugilus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="256" phylum="Arthropoda" rank="species" species="orbiculatus">
<emphasis id="528C036FFFBADD0F1108FE2B366443D8" box="[1300,1317,387,408]" italics="true" pageId="7" pageNumber="256">R</emphasis>
</taxonomicName>
</figureCitation>
. orbiculatus
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
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