310 lines
40 KiB
XML
310 lines
40 KiB
XML
<document id="34912F9403AF8523BCB11555CF1427AC" ID="10.11646/zootaxa.4171.2.11" ID-DOI="10.11646/zootaxa.4171.2.11" ID-GBIF-Dataset="cff8dbcf-c907-4f3e-bd61-0c86dca0f5ab" ID-ISSN="1175-5326" ID-Zenodo-Dep="255165" ID-ZooBank="C3853A62-10CA-4969-BDCB-8032A404EBEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1475144363584" checkinUser="plazi" docAuthor="Choong, Chee Yen" docDate="2016" docId="E849F268FFAE9016EBC74EDE879EADE2" docLanguage="en" docName="zootaxa.4171.2.11.pdf" docOrigin="Zootaxa 4171 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptogomphus tioman Choong, 2016, sp. nov." docType="treatment" docVersion="7" lastPageNumber="385" masterDocId="14708A10FFAE9015EB5048648560AA47" masterDocTitle="Leptogomphus tioman sp. nov. (Odonata: Anisoptera: Gomphidea) from Tioman Island, Peninsular Malaysia" masterLastPageNumber="388" masterPageNumber="382" pageNumber="382" updateTime="1698387786157" updateUser="plazi">
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<mods:mods id="7FB6DA3704F1ED68175D482103AF72F2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0062834D5D521BD28BCBED0F3E0A49E3">
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<mods:title id="E9E37B3A6166BD239FC228AB080503FD">Leptogomphus tioman sp. nov. (Odonata: Anisoptera: Gomphidea) from Tioman Island, Peninsular Malaysia</mods:title>
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</mods:titleInfo>
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<mods:name id="8D5027824CCEBAA8D0836BFE97E7867F" type="personal">
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<mods:role id="D927A41E2E7E1F690764646C23D5D6C3">
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<mods:roleTerm id="2BD929D1517D47E23C383B44C529C3B3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E748A90301BC861D590DD9ADF9D06A5C">Choong, Chee Yen</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3869C4C833BF1A198C2C25A6AD2533A4">text</mods:typeOfResource>
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<mods:titleInfo id="3ADF2B0D035FC24F8D49457CDA007CE5">
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<mods:title id="E2926BA64518A5B9D906D9D9D9204D7D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="2A24A705AA20A817F14BCB6140133352">
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<mods:date id="710383D644DA49583AB83438B309BAC8">2016</mods:date>
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<mods:detail id="00D23803878648E9F23316AEEDC78252" type="volume">
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<mods:number id="E571C1868ED9C9B5035D5628C93E0133">4171</mods:number>
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</mods:detail>
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<mods:detail id="0FF615F9CAD02B91011318B3A8E5297C" type="issue">
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<mods:number id="AF1CBAD9F59DCA461815F05EF68697AD">2</mods:number>
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<mods:start id="3E250F847A721D79E96E04BA10A9207B">382</mods:start>
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<mods:end id="96F99F7685FA6A05B06746A48966D829">388</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="A25F7C288EC448E2DD61E9CD41420831">journal article</mods:classification>
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<mods:identifier id="F2E60674DFA37F2A8E0DF6C47759FB0D" type="DOI">10.11646/zootaxa.4171.2.11</mods:identifier>
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<mods:identifier id="BDB6BBB3A52FB110722478A67580229B" type="GBIF-Dataset">cff8dbcf-c907-4f3e-bd61-0c86dca0f5ab</mods:identifier>
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<mods:identifier id="776896561A22ACC0F0F247D28CA176E0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="1720BFC167459DFFF6CC7DD989E2867E" type="Zenodo-Dep">255165</mods:identifier>
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<mods:identifier id="4999139DB820B2FF55DBA426B066877D" type="ZooBank">C3853A62-10CA-4969-BDCB-8032A404EBEC</mods:identifier>
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</mods:mods>
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<treatment id="E849F268FFAE9016EBC74EDE879EADE2" ID-DOI="http://doi.org/10.5281/zenodo.6087518" ID-GBIF-Taxon="124556825" ID-Zenodo-Dep="6087518" LSID="urn:lsid:plazi:treatment:E849F268FFAE9016EBC74EDE879EADE2" httpUri="http://treatment.plazi.org/id/E849F268FFAE9016EBC74EDE879EADE2" lastPageId="3" lastPageNumber="385" pageId="0" pageNumber="382">
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<subSubSection id="28FA10F5FFAE9015EBC74EDE876BAC92" box="[151,523,1722,1749]" pageId="0" pageNumber="382" type="nomenclature">
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<paragraph id="605F437EFFAE9015EBC74EDE876BAC92" blockId="0.[151,523,1722,1749]" box="[151,523,1722,1749]" pageId="0" pageNumber="382">
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<heading id="3B17F412FFAE9015EBC74EDE876BAC92" bold="true" box="[151,523,1722,1749]" fontSize="11" level="1" pageId="0" pageNumber="382" reason="1">
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<emphasis id="52949F6CFFAE9015EBC74EDE876BAC92" bold="true" box="[151,523,1722,1749]" pageId="0" pageNumber="382">
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<taxonomicName id="A7E038FDFFAE9015EBC74EDE84C6AC93" ID-CoL="9X35C" box="[151,422,1722,1748]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="382" phylum="Arthropoda" rank="species" species="tioman" status="sp. nov.">
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<emphasis id="52949F6CFFAE9015EBC74EDE84C6AC93" bold="true" box="[151,422,1722,1748]" italics="true" pageId="0" pageNumber="382">Leptogomphus tioman</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="49A72217FFAE9015EAFD4EDF876BAC92" box="[429,523,1723,1749]" pageId="0" pageNumber="382" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="28FA10F5FFAE9015EBC74F658181ADB5" pageId="0" pageNumber="382" type="materials_examined">
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<paragraph id="605F437EFFAE9015EBC74F658181ADB5" blockId="0.[151,1436,1793,2034]" pageId="0" pageNumber="382">
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<materialsCitation id="D0884923FFAE9015EBC74F6587B6AD79" ID-GBIF-Occurrence="1315932719" collectingDate="2014-05-18" country="Malaysia" county="Island" location="Si-Mukut" municipality="Mukut" pageId="0" pageNumber="382" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
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<emphasis id="52949F6CFFAE9015EBC74F6584E3AD5E" bold="true" box="[151,387,1793,1818]" pageId="0" pageNumber="382">
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Material.
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<typeStatus id="BF5BFDDCFFAE9015EA5C4F65841DAD5E" box="[268,381,1793,1817]" pageId="0" pageNumber="382" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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Ƌ (TIO3_GOM1), Peninsular
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<collectingCountry id="18F703EEFFAE9015E98B4F658625AD5D" box="[731,837,1793,1818]" name="Malaysia" pageId="0" pageNumber="382">Malaysia</collectingCountry>
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,
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<collectingRegion id="A2248D9CFFAE9015E81F4F6586C4AD5D" box="[847,932,1793,1818]" country="Malaysia" name="Pahang" pageId="0" pageNumber="382">Pahang</collectingRegion>
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,
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<taxonomicName id="A7E038FDFFAE9015E8FF4F658165AD5D" box="[943,1029,1793,1818]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="387" phylum="Arthropoda" rank="species" species="tioman">Tioman</taxonomicName>
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<collectingCounty id="893E3BF2FFAE9015EF5B4F658132AD5D" box="[1035,1106,1793,1818]" pageId="0" pageNumber="382">Island</collectingCounty>
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,
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<collectingMunicipality id="803BD904FFAE9015EF0D4F6581CBAD5D" box="[1117,1195,1793,1818]" pageId="0" pageNumber="382">Mukut</collectingMunicipality>
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, a small rocky stream at the foothill of
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<location id="653F15A5FFAE9015EA344F4184B2AD79" LSID="urn:lsid:plazi:treatment:E849F268FFAE9016EBC74EDE879EADE2:653F15A5FFAE9015EA344F4184B2AD79" box="[356,466,1829,1854]" country="Malaysia" county="Island" municipality="Mukut" name="Si-Mukut" pageId="0" pageNumber="382" stateProvince="Pahang">Si-Mukut</location>
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,
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<date id="145E65BEFFAE9015EAB14F42873DAD79" box="[481,605,1830,1854]" pageId="0" pageNumber="382" value="2014-05-18">
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<collectingDate id="041A9C56FFAE9015EAB14F42873DAD79" box="[481,605,1830,1854]" pageId="0" pageNumber="382" value="2014-05-18">18 v 2014</collectingDate>
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</date>
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, CYC leg
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</materialsCitation>
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.
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<materialsCitation id="D0884923FFAE9015E9B54F418651AD25" ID-GBIF-Occurrence="1315932717" country="Iceland" location="Mukut Village" municipality="Mukut" pageId="0" pageNumber="382" specimenCount="1" typeStatus="paratype">
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<emphasis id="52949F6CFFAE9015E9B54F418607AD7A" bold="true" box="[741,871,1829,1853]" pageId="0" pageNumber="382">
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<typeStatus id="BF5BFDDCFFAE9015E9B54F418602AD7A" box="[741,866,1829,1853]" pageId="0" pageNumber="382" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<docIssue id="24833C91FFAE9015E8224F4286E2AD79" box="[882,898,1830,1854]" pageId="0" pageNumber="382">2</docIssue>
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Ƌ (TIO4_GOM2,3), Sg. Raya,
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<taxonomicName id="A7E038FDFFAE9015EFBF4F418025AD79" box="[1263,1349,1829,1854]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="387" phylum="Arthropoda" rank="species" species="tioman">Tioman</taxonomicName>
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<collectingCountry id="18F703EEFFAE9015EE004F4180F7AD79" box="[1360,1431,1829,1854]" name="Iceland" pageId="0" pageNumber="382">Island</collectingCountry>
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,
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<collectingMunicipality id="803BD904FFAE9015EBC74F2D8585AD25" box="[151,229,1865,1890]" pageId="0" pageNumber="382">Mukut</collectingMunicipality>
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, a small rocky stream at the back of
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<location id="653F15A5FFAE9015E9DA4F2D8651AD25" LSID="urn:lsid:plazi:treatment:E849F268FFAE9016EBC74EDE879EADE2:653F15A5FFAE9015E9DA4F2D8651AD25" box="[650,817,1865,1890]" country="Iceland" municipality="Mukut" name="Mukut Village" pageId="0" pageNumber="382">Mukut Village</location>
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</materialsCitation>
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,
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<materialsCitation id="D0884923FFAE9015E86D4F2E84D9ADC1" ID-GBIF-Occurrence="1315932716" collectingDate="2016-04-18" collectorName="Sg. Asah" country="Iceland" location="Island" pageId="0" pageNumber="382" specimenCount="1" typeStatus="paratype">
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<date id="145E65BEFFAE9015E86D4F2E86DBAD25" box="[829,955,1865,1890]" pageId="0" pageNumber="382" value="2016-04-18">
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<collectingDate id="041A9C56FFAE9015E86D4F2E86DBAD25" box="[829,955,1865,1890]" pageId="0" pageNumber="382" value="2016-04-18">18 iv 2016</collectingDate>
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</date>
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, CYC leg; m (TIO4_GOM1), upstream of
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<collectorName id="CD1526A8FFAE9015EBC74F0A8462ADC1" box="[151,258,1901,1926]" pageId="0" pageNumber="382">Sg. Asah</collectorName>
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,
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<taxonomicName id="A7E038FDFFAE9015EA414F098407ADC1" box="[273,359,1901,1926]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="387" phylum="Arthropoda" rank="species" species="tioman">Tioman</taxonomicName>
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<collectingCountry id="18F703EEFFAE9015EA224F0984D9ADC1" box="[370,441,1901,1926]" name="Iceland" pageId="0" pageNumber="382">Island</collectingCountry>
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||
</materialsCitation>
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||
,
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<materialsCitation id="D0884923FFAE9015EA984F0A85BEADED" ID-GBIF-Occurrence="1315932721" collectingDate="2016-04-17" collectorName="Sg. Asah & Asah Waterfall" country="Iceland" location="Island" pageId="0" pageNumber="382" specimenCount="1" typeStatus="paratype">
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<date id="145E65BEFFAE9015EA984F0A872AADC1" box="[456,586,1901,1926]" pageId="0" pageNumber="382" value="2016-04-17">
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<collectingDate id="041A9C56FFAE9015EA984F0A872AADC1" box="[456,586,1901,1926]" pageId="0" pageNumber="382" value="2016-04-17">17 iv 2016</collectingDate>
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</date>
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, CYC leg; 2Ƌ (TIO4_GOM4,5),
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<collectorName id="CD1526A8FFAE9015E8834F0A815EADC1" box="[979,1086,1901,1926]" pageId="0" pageNumber="382">Sg. Asah</collectorName>
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, near
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<collectorName id="CD1526A8FFAE9015EFD84F098058ADC1" box="[1160,1336,1901,1926]" pageId="0" pageNumber="382">Asah Waterfall</collectorName>
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,
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<taxonomicName id="A7E038FDFFAE9015EE164F0980FCADC1" box="[1350,1436,1901,1926]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="387" phylum="Arthropoda" rank="species" species="tioman">Tioman</taxonomicName>
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<collectingCountry id="18F703EEFFAE9015EBC74FF585BEADED" box="[151,222,1937,1962]" name="Iceland" pageId="0" pageNumber="382">Island</collectingCountry>
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</materialsCitation>
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,
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<materialsCitation id="D0884923FFAE9015EBBC4FF68603AD89" ID-GBIF-Occurrence="1315932720" collectingDate="2016-04-19" collectionCode="DWNP" collectorName="Holotype & Institut Biodiversiti & Lanchang & Department & Wildlife & National Parks" country="Malaysia" location="Malaysia" pageId="0" pageNumber="382" specimenCount="1" typeStatus="paratype">
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<date id="145E65BEFFAE9015EBBC4FF6840EADED" box="[236,366,1937,1962]" pageId="0" pageNumber="382" value="2016-04-19">
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<collectingDate id="041A9C56FFAE9015EBBC4FF6840EADED" box="[236,366,1937,1962]" pageId="0" pageNumber="382" value="2016-04-19">19 iv 2016</collectingDate>
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</date>
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, CYC leg.
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<collectorName id="CD1526A8FFAE9015EAA44FF5873CADED" box="[500,604,1937,1962]" pageId="0" pageNumber="382">Holotype</collectorName>
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to be deposited in
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<collectorName id="CD1526A8FFAE9015E8114FF5817FADED" box="[833,1055,1937,1962]" pageId="0" pageNumber="382">Institut Biodiversiti</collectorName>
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(IBD)
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<collectorName id="CD1526A8FFAE9015EF274FF58188ADED" box="[1143,1256,1937,1962]" pageId="0" pageNumber="382">Lanchang</collectorName>
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,
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<collectorName id="CD1526A8FFAE9015EFA64FF6801AADEE" box="[1270,1402,1938,1961]" pageId="0" pageNumber="382">Department</collectorName>
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||
of
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<collectorName id="CD1526A8FFAE9015EBC74FD18593AD89" box="[151,243,1973,1998]" pageId="0" pageNumber="382">Wildlife</collectorName>
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and
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<collectorName id="CD1526A8FFAE9015EA6D4FD1848CAD89" box="[317,492,1973,1998]" pageId="0" pageNumber="382">National Parks</collectorName>
|
||
(
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<collectionCode id="06F1DBBBFFAE9015E9544FD2873AAD89" box="[516,602,1974,1998]" pageId="0" pageNumber="382">DWNP</collectionCode>
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) Peninsular
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<collectingCountry id="18F703EEFFAE9015E9AA4FD18603AD89" box="[762,867,1973,1998]" name="Malaysia" pageId="0" pageNumber="382">Malaysia</collectingCountry>
|
||
</materialsCitation>
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||
,
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<materialsCitation id="D0884923FFAE9015E8284FD28181ADB5" ID-GBIF-Occurrence="1315932718" country="Malaysia" location="Insect Systematics" municipality="Centre" pageId="0" pageNumber="382" specimenCount="1" typeStatus="paratype">
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<typeStatus id="BF5BFDDCFFAE9015E8284FD28683AD89" box="[888,995,1974,1998]" pageId="0" pageNumber="382" type="paratype">paratypes</typeStatus>
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||
in collections of
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||
<collectingMunicipality id="803BD904FFAE9015EF9D4FD28078AD89" box="[1229,1304,1974,1998]" pageId="0" pageNumber="382">Centre</collectingMunicipality>
|
||
for
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||
<location id="653F15A5FFAE9015EE094FD2847DADB5" LSID="urn:lsid:plazi:treatment:E849F268FFAE9016EBC74EDE879EADE2:653F15A5FFAE9015EE094FD2847DADB5" country="Malaysia" municipality="Centre" name="Insect Systematics" pageId="0" pageNumber="382">Insect Systematics</location>
|
||
(Universiti Kebangsaan
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||
<collectingCountry id="18F703EEFFAE9015E9644FBD87FCADB5" box="[564,668,2009,2034]" name="Malaysia" pageId="0" pageNumber="382">Malaysia</collectingCountry>
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||
) (TIO4_GOM1,2,5) and the IBD (TIO4_GOM3,4).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="349F13F6FFAF9014EBC74C2086A4AE1D" box="[151,964,1092,1114]" httpUri="https://zenodo.org/record/255166/files/figure.png" pageId="1" pageNumber="383" targetBox="[170,1435,209,1068]" targetPageId="1">
|
||
<paragraph id="605F437EFFAF9014EBC74C2086A4AE1D" blockId="1.[151,964,1092,1114]" box="[151,964,1092,1114]" pageId="1" pageNumber="383">
|
||
<emphasis id="52949F6CFFAF9014EBC74C208473AE1E" bold="true" box="[151,275,1092,1114]" pageId="1" pageNumber="383">FIGURE 1.</emphasis>
|
||
<taxonomicName id="A7E038FDFFAF9014EA494C20849AAE1D" box="[281,506,1092,1114]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="383" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="52949F6CFFAF9014EA494C20849AAE1D" box="[281,506,1092,1114]" italics="true" pageId="1" pageNumber="383">Leptogomphus tioman</emphasis>
|
||
</taxonomicName>
|
||
, holotype, lateral view, scale bar = 20.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="28FA10F5FFAF9014EB974CE68032AEDD" box="[199,1362,1153,1178]" pageId="1" pageNumber="383" type="etymology">
|
||
<paragraph id="605F437EFFAF9014EB974CE68032AEDD" blockId="1.[151,1437,1153,2006]" box="[199,1362,1153,1178]" pageId="1" pageNumber="383">
|
||
<emphasis id="52949F6CFFAF9014EB974CE68426AEDD" bold="true" box="[199,326,1154,1178]" pageId="1" pageNumber="383">Etymology</emphasis>
|
||
. The specific name is a noun in apposition and named after
|
||
<taxonomicName id="A7E038FDFFAF9014E8B84CE5815EAEDD" box="[1000,1086,1153,1178]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="387" phylum="Arthropoda" rank="species" species="tioman">Tioman</taxonomicName>
|
||
<collectingCountry id="18F703EEFFAF9014EF164CE581EDAEDD" box="[1094,1165,1153,1178]" name="Iceland" pageId="1" pageNumber="383">Island</collectingCountry>
|
||
, the
|
||
<typeStatus id="BF5BFDDCFFAF9014EF914CE68191AEDD" box="[1217,1265,1154,1178]" pageId="1" pageNumber="383">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="28FA10F5FFAF9017EB974CC184F8AB22" lastPageId="2" lastPageNumber="384" pageId="1" pageNumber="383" type="description">
|
||
<paragraph id="605F437EFFAF9014EB974CC1870CAF09" blockId="1.[151,1437,1153,2006]" pageId="1" pageNumber="383">
|
||
<emphasis id="52949F6CFFAF9014EB974CC1874AAEF9" bold="true" box="[199,554,1189,1214]" pageId="1" pageNumber="383">
|
||
Description of
|
||
<typeStatus id="BF5BFDDCFFAF9014EA2F4CC28486AEF9" box="[383,486,1190,1214]" pageId="1" pageNumber="383" type="holotype">holotype</typeStatus>
|
||
male
|
||
</emphasis>
|
||
. A medium-sized dragonfly with greenish yellow markings (
|
||
<figureCitation id="F8DB5FFBFFAF9014EFA74CC1805DAEF9" box="[1271,1341,1189,1214]" captionStart="FIGURE 1" captionStartId="1.[151,250,1092,1114]" captionTargetBox="[170,1435,209,1068]" captionTargetId="figure@1.[151,1436,195,1068]" captionTargetPageId="1" captionText="FIGURE 1. Leptogomphus tioman, holotype, lateral view, scale bar = 20.0 mm." httpUri="https://zenodo.org/record/255166/files/figure.png" pageId="1" pageNumber="383">Fig. 1</figureCitation>
|
||
). Head: Vertex mainly black, but brown behind tubercles posterior to lateral ocelli; occiput also brown with brown setae. Frons distinctly swollen, mostly greenish yellow. Labrum black with a pair of broad greenish-yellow stripes next to clypeus. Clypeus black with a paired oval depression on postclypeus. Mandibles totally black with greenish-yellow marking at base. Labium greenish yellow.
|
||
</paragraph>
|
||
<paragraph id="605F437EFFAF9014EB974D3D841CACD5" blockId="1.[151,1437,1153,2006]" pageId="1" pageNumber="383">
|
||
Thorax: Prothorax with anterior lobe mainly greenish yellow (
|
||
<figureCitation id="F8DB5FFBFFAF9014E8EF4D3D816CAF35" box="[959,1036,1369,1394]" captionStart="FIGURES 2 – 7" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[151,1412,409,1805]" captionTargetId="figure@2.[151,1436,409,1858]" captionTargetPageId="2" captionText="FIGURES 2 – 7. Leptogomphus tioman, holotype: (2) thorax, dorsal view in live; (3) thorax, lateral view; (4) anal appendages, dorsal view; (5) anal appendages, lateral view; (6) anal appendages, oblique dorsal view; (7) accessary genitalia, lateral view." httpUri="https://zenodo.org/record/255167/files/figure.png" pageId="1" pageNumber="383">Fig. 2</figureCitation>
|
||
); middle lobe with deep median depression anteriorly, a pair of middorsal yellow spots and greenish yellow in lateral edges; posterior lobe dark brown. Synthorax black with greenish-yellow markings; dorsal thoracic stripes broadly connected to sections of mesothoracic collar, which is interrupted medially (
|
||
<figureCitation id="F8DB5FFBFFAF9014E9A34DA1865CAF99" box="[755,828,1477,1502]" captionStart="FIGURES 2 – 7" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[151,1412,409,1805]" captionTargetId="figure@2.[151,1436,409,1858]" captionTargetPageId="2" captionText="FIGURES 2 – 7. Leptogomphus tioman, holotype: (2) thorax, dorsal view in live; (3) thorax, lateral view; (4) anal appendages, dorsal view; (5) anal appendages, lateral view; (6) anal appendages, oblique dorsal view; (7) accessary genitalia, lateral view." httpUri="https://zenodo.org/record/255167/files/figure.png" pageId="1" pageNumber="383">Fig. 2</figureCitation>
|
||
); yellow marking on mesepimeron and pale yellow marking on metepisternum; metepimeron largely yellow with thick black anterior margin (
|
||
<figureCitation id="F8DB5FFBFFAF9014EFB34D8D804FAC45" box="[1251,1327,1513,1538]" captionStart="FIGURES 2 – 7" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[151,1412,409,1805]" captionTargetId="figure@2.[151,1436,409,1858]" captionTargetPageId="2" captionText="FIGURES 2 – 7. Leptogomphus tioman, holotype: (2) thorax, dorsal view in live; (3) thorax, lateral view; (4) anal appendages, dorsal view; (5) anal appendages, lateral view; (6) anal appendages, oblique dorsal view; (7) accessary genitalia, lateral view." httpUri="https://zenodo.org/record/255167/files/figure.png" pageId="1" pageNumber="383">Fig. 3</figureCitation>
|
||
). Wings: Forewing with 16 Ax, 15 Px; hindwing with 13 Ax and 16 Px. Pterostigma dark brown, 3.0 mm. Arculus about 1/3 distance between second and third Ax. Cu crossing proximal to first Ax. Triangle, subtriangle and supratriangle all without crossveins. Legs: stoutly built. All femora with short spines on inner side; all tibiae with two rows of long spines on inner side.
|
||
</paragraph>
|
||
<paragraph id="605F437EFFAF9017EB974EF98067AB5A" blockId="1.[151,1437,1153,2006]" lastBlockId="2.[151,1436,151,357]" lastPageId="2" lastPageNumber="384" pageId="1" pageNumber="383">
|
||
Abdomen: Nearly black with narrow yellow middorsal stripe visible on S1–2. S1 side yellow, S2 with a pair of ventral yellow markings and S3 anterolateral side also yellow. Auricules with yellow marking hardly developed. Abdomen gradually expended from tip of S7, reaching the widest point at tip of S8, then narrowing gradually to tip of S10. Dorsum S10 covered with small denticles (
|
||
<figureCitation id="F8DB5FFBFFAF9014E98D4F6D862BAD65" box="[733,843,1801,1826]" captionStart="FIGURES 2 – 7" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[151,1412,409,1805]" captionTargetId="figure@2.[151,1436,409,1858]" captionTargetPageId="2" captionText="FIGURES 2 – 7. Leptogomphus tioman, holotype: (2) thorax, dorsal view in live; (3) thorax, lateral view; (4) anal appendages, dorsal view; (5) anal appendages, lateral view; (6) anal appendages, oblique dorsal view; (7) accessary genitalia, lateral view." httpUri="https://zenodo.org/record/255167/files/figure.png" pageId="1" pageNumber="383">Figs. 4–6</figureCitation>
|
||
). Anal appendages: Cerci black (
|
||
<figureCitation id="F8DB5FFBFFAF9014EF944F6D8052AD65" box="[1220,1330,1801,1826]" captionStart="FIGURES 2 – 7" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[151,1412,409,1805]" captionTargetId="figure@2.[151,1436,409,1858]" captionTargetPageId="2" captionText="FIGURES 2 – 7. Leptogomphus tioman, holotype: (2) thorax, dorsal view in live; (3) thorax, lateral view; (4) anal appendages, dorsal view; (5) anal appendages, lateral view; (6) anal appendages, oblique dorsal view; (7) accessary genitalia, lateral view." httpUri="https://zenodo.org/record/255167/files/figure.png" pageId="1" pageNumber="383">Figs. 4–6</figureCitation>
|
||
), thick at base and curved slightly inward at about apical 2/3, turned slightly upward at tip; tip of cerci rather sharp in dorsal and lateral views. Lower edge of cercus produced outward and sharply angled inward for apical 1/3; outer margin of angle bearing strong serration (
|
||
<figureCitation id="F8DB5FFBFFAF9014E94C4F11870DADC9" box="[540,621,1909,1934]" captionStart="FIGURES 2 – 7" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[151,1412,409,1805]" captionTargetId="figure@2.[151,1436,409,1858]" captionTargetPageId="2" captionText="FIGURES 2 – 7. Leptogomphus tioman, holotype: (2) thorax, dorsal view in live; (3) thorax, lateral view; (4) anal appendages, dorsal view; (5) anal appendages, lateral view; (6) anal appendages, oblique dorsal view; (7) accessary genitalia, lateral view." httpUri="https://zenodo.org/record/255167/files/figure.png" pageId="1" pageNumber="383">Figs. 4</figureCitation>
|
||
,
|
||
<figureCitation id="F8DB5FFBFFAF9014E92B4F1287EBADC9" box="[635,651,1910,1934]" captionStart="FIGURES 8 – 14" captionStartId="3.[151,264,1485,1508]" captionTargetBox="[164,1422,202,1455]" captionTargetId="figure@3.[150,1436,193,1464]" captionTargetPageId="3" captionText="FIGURES 8 – 14. Anal appendages: (8) L. tioman holotype, dorsal view; (9) L. risi, Malaysia, dorsal view, redrawn from Asahina (1986); (10) L. risi, Thailand, dorsal view, redrawn from Asahina (1986); (11) L. tioman holotype, lateral view; (12) L. risi, Thailand, latral view, redrawn from Asahina (1986); (13) L. tioman holotype, oblique dorsal view; (14) L. risi, Malaysia, oblique dorsal view, redrawn from Asahina (1986)." httpUri="https://zenodo.org/record/255168/files/figure.png" pageId="1" pageNumber="383">8</figureCitation>
|
||
). Small conical, subapical, ventral process on each cercus, visible in oblique dorsal view (
|
||
<figureCitation id="F8DB5FFBFFAF9014EAD34FFD84B2ADF5" box="[387,466,1945,1970]" captionStart="FIGURES 2 – 7" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[151,1412,409,1805]" captionTargetId="figure@2.[151,1436,409,1858]" captionTargetPageId="2" captionText="FIGURES 2 – 7. Leptogomphus tioman, holotype: (2) thorax, dorsal view in live; (3) thorax, lateral view; (4) anal appendages, dorsal view; (5) anal appendages, lateral view; (6) anal appendages, oblique dorsal view; (7) accessary genitalia, lateral view." httpUri="https://zenodo.org/record/255167/files/figure.png" pageId="1" pageNumber="383">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="F8DB5FFBFFAF9014EA8D4FFE849BADF5" box="[477,507,1946,1970]" captionStart="FIGURES 8 – 14" captionStartId="3.[151,264,1485,1508]" captionTargetBox="[164,1422,202,1455]" captionTargetId="figure@3.[150,1436,193,1464]" captionTargetPageId="3" captionText="FIGURES 8 – 14. Anal appendages: (8) L. tioman holotype, dorsal view; (9) L. risi, Malaysia, dorsal view, redrawn from Asahina (1986); (10) L. risi, Thailand, dorsal view, redrawn from Asahina (1986); (11) L. tioman holotype, lateral view; (12) L. risi, Thailand, latral view, redrawn from Asahina (1986); (13) L. tioman holotype, oblique dorsal view; (14) L. risi, Malaysia, oblique dorsal view, redrawn from Asahina (1986)." httpUri="https://zenodo.org/record/255168/files/figure.png" pageId="1" pageNumber="383">13</figureCitation>
|
||
). In lateral view (
|
||
<figureCitation id="F8DB5FFBFFAF9014E9914FFD8670ADF5" box="[705,784,1945,1970]" captionStart="FIGURES 2 – 7" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[151,1412,409,1805]" captionTargetId="figure@2.[151,1436,409,1858]" captionTargetPageId="2" captionText="FIGURES 2 – 7. Leptogomphus tioman, holotype: (2) thorax, dorsal view in live; (3) thorax, lateral view; (4) anal appendages, dorsal view; (5) anal appendages, lateral view; (6) anal appendages, oblique dorsal view; (7) accessary genitalia, lateral view." httpUri="https://zenodo.org/record/255167/files/figure.png" pageId="1" pageNumber="383">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="F8DB5FFBFFAF9014E84C4FFE8659ADF5" box="[796,825,1946,1970]" captionStart="FIGURES 8 – 14" captionStartId="3.[151,264,1485,1508]" captionTargetBox="[164,1422,202,1455]" captionTargetId="figure@3.[150,1436,193,1464]" captionTargetPageId="3" captionText="FIGURES 8 – 14. Anal appendages: (8) L. tioman holotype, dorsal view; (9) L. risi, Malaysia, dorsal view, redrawn from Asahina (1986); (10) L. risi, Thailand, dorsal view, redrawn from Asahina (1986); (11) L. tioman holotype, lateral view; (12) L. risi, Thailand, latral view, redrawn from Asahina (1986); (13) L. tioman holotype, oblique dorsal view; (14) L. risi, Malaysia, oblique dorsal view, redrawn from Asahina (1986)." httpUri="https://zenodo.org/record/255168/files/figure.png" pageId="1" pageNumber="383">11</figureCitation>
|
||
), cercus robust, broad, straight-sided, tapered upwards to a point from apical 2/3; outer surface of cercus covered with small denticles apically. In ventral view, epiproct black, making a wide V-shape before diverging outward to tips; in lateral view bending evenly upward with a dorsal hook at tip. Genitalia (
|
||
<figureCitation id="F8DB5FFBFFAC9017EABE48D88720AA93" box="[494,576,188,213]" captionStart="FIGURES 2 – 7" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[151,1412,409,1805]" captionTargetId="figure@2.[151,1436,409,1858]" captionTargetPageId="2" captionText="FIGURES 2 – 7. Leptogomphus tioman, holotype: (2) thorax, dorsal view in live; (3) thorax, lateral view; (4) anal appendages, dorsal view; (5) anal appendages, lateral view; (6) anal appendages, oblique dorsal view; (7) accessary genitalia, lateral view." httpUri="https://zenodo.org/record/255167/files/figure.png" pageId="2" pageNumber="384">Figs. 7</figureCitation>
|
||
,
|
||
<figureCitation id="F8DB5FFBFFAC9017E90048D88710AA93" box="[592,624,188,212]" captionStart="FIGURES 8 – 14" captionStartId="3.[151,264,1485,1508]" captionTargetBox="[164,1422,202,1455]" captionTargetId="figure@3.[150,1436,193,1464]" captionTargetPageId="3" captionText="FIGURES 8 – 14. Anal appendages: (8) L. tioman holotype, dorsal view; (9) L. risi, Malaysia, dorsal view, redrawn from Asahina (1986); (10) L. risi, Thailand, dorsal view, redrawn from Asahina (1986); (11) L. tioman holotype, lateral view; (12) L. risi, Thailand, latral view, redrawn from Asahina (1986); (13) L. tioman holotype, oblique dorsal view; (14) L. risi, Malaysia, oblique dorsal view, redrawn from Asahina (1986)." httpUri="https://zenodo.org/record/255168/files/figure.png" pageId="2" pageNumber="384">14</figureCitation>
|
||
): Anterior hamule with broad base and pronounced shoulder, tapering posteriorly to a thin layer. Posterior hamule broad throughout, with a pronounced curve point at apical part. Vesica spermalis with rounded posterior lobe on basal segment and two diverging flagella on distal segment.
|
||
</paragraph>
|
||
<paragraph id="605F437EFFAC9017EB97494C8112AB78" blockId="2.[151,1436,151,357]" box="[199,1138,295,320]" pageId="2" pageNumber="384">Measurements of male: Abdomen (incl. appendages) 40.0 mm. Hindwing 31.5 mm.</paragraph>
|
||
<paragraph id="605F437EFFAC9017EB97492884F8AB22" blockId="2.[151,1436,151,357]" box="[199,408,332,357]" pageId="2" pageNumber="384">Female: unknown.</paragraph>
|
||
</subSubSection>
|
||
<caption id="349F13F6FFAC9017EBC74F3280EBADCC" httpUri="https://zenodo.org/record/255167/files/figure.png" pageId="2" pageNumber="384" targetBox="[151,1412,409,1805]" targetPageId="2">
|
||
<paragraph id="605F437EFFAC9017EBC74F3280EBADCC" blockId="2.[151,1436,1878,1931]" pageId="2" pageNumber="384">
|
||
<emphasis id="52949F6CFFAC9017EBC74F32845BAD2B" bold="true" box="[151,315,1878,1901]" pageId="2" pageNumber="384">FIGURES 2–7.</emphasis>
|
||
<taxonomicName id="A7E038FDFFAC9017EA114F338743AD2A" box="[321,547,1879,1901]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="384" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="52949F6CFFAC9017EA114F338743AD2A" box="[321,547,1879,1901]" italics="true" pageId="2" pageNumber="384">Leptogomphus tioman</emphasis>
|
||
</taxonomicName>
|
||
, holotype: (2) thorax, dorsal view in live; (3) thorax, lateral view; (4) anal appendages, dorsal view; (5) anal appendages, lateral view; (6) anal appendages, oblique dorsal view; (7) accessary genitalia, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="349F13F6FFAD9016EBC74DA987FEAC78" httpUri="https://zenodo.org/record/255168/files/figure.png" pageId="3" pageNumber="385" targetBox="[164,1422,202,1455]" targetPageId="3">
|
||
<paragraph id="605F437EFFAD9016EBC74DA987FEAC78" blockId="3.[151,1437,1485,1599]" pageId="3" pageNumber="385">
|
||
<emphasis id="52949F6CFFAD9016EBC74DA9842AAFA4" bold="true" box="[151,330,1485,1508]" pageId="3" pageNumber="385">FIGURES 8–14.</emphasis>
|
||
Anal appendages: (8)
|
||
<taxonomicName id="A7E038FDFFAD9016E96E4DAB87C3AFA3" box="[574,675,1487,1508]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="52949F6CFFAD9016E96E4DAB87C3AFA3" box="[574,675,1487,1508]" italics="true" pageId="3" pageNumber="385">L. tioman</emphasis>
|
||
</taxonomicName>
|
||
holotype, dorsal view; (9)
|
||
<taxonomicName id="A7E038FDFFAD9016E8984DAB8168AFA3" box="[968,1032,1487,1508]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="risi">
|
||
<emphasis id="52949F6CFFAD9016E8984DAB8168AFA3" box="[968,1032,1487,1508]" italics="true" pageId="3" pageNumber="385">L. risi</emphasis>
|
||
</taxonomicName>
|
||
, Malaysia, dorsal view, redrawn from Asahina (1986); (10)
|
||
<taxonomicName id="A7E038FDFFAD9016EA204D8984CDAC45" box="[368,429,1517,1538]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="risi">
|
||
<emphasis id="52949F6CFFAD9016EA204D8984CDAC45" box="[368,429,1517,1538]" italics="true" pageId="3" pageNumber="385">L. risi</emphasis>
|
||
</taxonomicName>
|
||
, Thailand, dorsal view, redrawn from Asahina (1986); (11)
|
||
<taxonomicName id="A7E038FDFFAD9016EF584D898109AC45" box="[1032,1129,1517,1538]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="52949F6CFFAD9016EF584D898109AC45" box="[1032,1129,1517,1538]" italics="true" pageId="3" pageNumber="385">L. tioman</emphasis>
|
||
</taxonomicName>
|
||
holotype, lateral view; (12)
|
||
<taxonomicName id="A7E038FDFFAD9016EED74D8985D9AC66" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="risi">
|
||
<emphasis id="52949F6CFFAD9016EED74D8985D9AC66" italics="true" pageId="3" pageNumber="385">L. risi</emphasis>
|
||
</taxonomicName>
|
||
, Thailand, latral view, redrawn from Asahina (1986); (13)
|
||
<taxonomicName id="A7E038FDFFAD9016E8454E688617AC66" box="[789,887,1548,1569]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="52949F6CFFAD9016E8454E688617AC66" box="[789,887,1548,1569]" italics="true" pageId="3" pageNumber="385">L. tioman</emphasis>
|
||
</taxonomicName>
|
||
holotype, oblique dorsal view; (14)
|
||
<taxonomicName id="A7E038FDFFAD9016EFBE4E68804CAC66" box="[1262,1324,1548,1569]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="risi">
|
||
<emphasis id="52949F6CFFAD9016EFBE4E68804CAC66" box="[1262,1324,1548,1569]" italics="true" pageId="3" pageNumber="385">L. risi</emphasis>
|
||
</taxonomicName>
|
||
, Malaysia, oblique dorsal view, redrawn from Asahina (1986).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="28FA10F5FFAD9016EB974E0F86E8AD7E" pageId="3" pageNumber="385" type="diagnosis">
|
||
<paragraph id="605F437EFFAD9016EB974E0F803BACB7" blockId="3.[151,1436,1643,1957]" pageId="3" pageNumber="385">
|
||
<emphasis id="52949F6CFFAD9016EB974E0F8421ACC3" bold="true" box="[199,321,1643,1668]" pageId="3" pageNumber="385">Diagnosis.</emphasis>
|
||
A greenish-yellow marked
|
||
<taxonomicName id="A7E038FDFFAD9016E9D24E088647ACC3" box="[642,807,1644,1668]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="52949F6CFFAD9016E9D24E088647ACC3" box="[642,807,1644,1668]" italics="true" pageId="3" pageNumber="385">Leptogomphus</emphasis>
|
||
</taxonomicName>
|
||
, readily distinguished from all known regional species except
|
||
<taxonomicName id="A7E038FDFFAD9016EBB84EF5844BACEF" box="[232,299,1681,1704]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="risi">
|
||
<emphasis id="52949F6CFFAD9016EBB84EF5844BACEF" box="[232,299,1681,1704]" italics="true" pageId="3" pageNumber="385">L. risi</emphasis>
|
||
</taxonomicName>
|
||
by its thoracic marking, and from all species by the unique structure of its cerci. The tips of cerci in dorsal view are sharp in
|
||
<taxonomicName id="A7E038FDFFAD9016EAF94ED18774AC8B" box="[425,532,1717,1740]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="52949F6CFFAD9016EAF94ED18774AC8B" box="[425,532,1717,1740]" italics="true" pageId="3" pageNumber="385">L. tioman</emphasis>
|
||
</taxonomicName>
|
||
but round in
|
||
<taxonomicName id="A7E038FDFFAD9016E9FB4ED1878DAC8B" box="[683,749,1717,1740]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="risi">
|
||
<emphasis id="52949F6CFFAD9016E9FB4ED1878DAC8B" box="[683,749,1717,1740]" italics="true" pageId="3" pageNumber="385">L. risi</emphasis>
|
||
</taxonomicName>
|
||
. The tips of the epiproct are closer together in
|
||
<taxonomicName id="A7E038FDFFAD9016EFA94ED18005AC8B" box="[1273,1381,1717,1740]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="52949F6CFFAD9016EFA94ED18005AC8B" box="[1273,1381,1717,1740]" italics="true" pageId="3" pageNumber="385">L. tioman</emphasis>
|
||
</taxonomicName>
|
||
than
|
||
<taxonomicName id="A7E038FDFFAD9016EBC74EBD85B9ACB7" box="[151,217,1753,1776]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="risi">
|
||
<emphasis id="52949F6CFFAD9016EBC74EBD85B9ACB7" box="[151,217,1753,1776]" italics="true" pageId="3" pageNumber="385">L. risi</emphasis>
|
||
</taxonomicName>
|
||
. Compared to
|
||
<taxonomicName id="A7E038FDFFAD9016EAD04EBD84A3ACB7" box="[384,451,1753,1776]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="risi">
|
||
<emphasis id="52949F6CFFAD9016EAD04EBD84A3ACB7" box="[384,451,1753,1776]" italics="true" pageId="3" pageNumber="385">L. risi</emphasis>
|
||
</taxonomicName>
|
||
, the outer margins of the epiproct in
|
||
<taxonomicName id="A7E038FDFFAD9016E8334EBD86AEACB7" box="[867,974,1753,1776]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="385" phylum="Arthropoda" rank="species" species="tioman">
|
||
<emphasis id="52949F6CFFAD9016E8334EBD86AEACB7" box="[867,974,1753,1776]" italics="true" pageId="3" pageNumber="385">L. tioman</emphasis>
|
||
</taxonomicName>
|
||
appear sinuous rather than convex.
|
||
</paragraph>
|
||
<paragraph id="605F437EFFAD9016EB974E9F86E8AD7E" blockId="3.[151,1436,1643,1957]" pageId="3" pageNumber="385">
|
||
<emphasis id="52949F6CFFAD9016EB974E9F844AAD53" bold="true" box="[199,298,1787,1812]" pageId="3" pageNumber="385">Habitat.</emphasis>
|
||
Adults are found on lowland small streams with a sand and gravel substrate in primary as well as secondary forests. The water is clean and flow is slow to moderate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="28FA10F5FFAD9016EB974F27879EADE2" pageId="3" pageNumber="385" type="description">
|
||
<paragraph id="605F437EFFAD9016EB974F27879EADE2" blockId="3.[151,1436,1643,1957]" pageId="3" pageNumber="385">
|
||
<emphasis id="52949F6CFFAD9016EB974F278458AD1B" bold="true" box="[199,312,1859,1884]" pageId="3" pageNumber="385">Behavior.</emphasis>
|
||
Male adults were spotted at noon on sunny days. The adults frequently perched in sunspots on leaves next to the water (
|
||
<figureCitation id="F8DB5FFBFFAD9016EAE34F0C8768ADC6" box="[435,520,1896,1921]" captionStart="FIGURE 17" captionStartId="4.[151,250,1941,1963]" captionTargetBox="[151,1435,948,1918]" captionTargetId="figure@4.[151,1436,948,1920]" captionTargetPageId="4" captionText="FIGURE 17. Leptogomphus tioman male, Sg. Raya, Tioman Island, Peninsular Malaysia, 18 April 2016." httpUri="https://zenodo.org/record/255170/files/figure.png" pageId="3" pageNumber="385">Fig. 17</figureCitation>
|
||
). When alarmed, they flew into the canopy perching on leaves, returning to their former perch after a short interval (roughly 5 minutes).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |