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<document id="12C25CCFCDF689DF93DE613776E4B455" ID-DOI="10.11646/zootaxa.3956.2.5" ID-GBIF-Dataset="693ef71c-2ee2-4da4-a3e1-35f5586899dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="232228" ID-ZooBank="C12EF13A-9C19-4051-80A6-F59B917774AA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461054896861" checkinUser="plazi" docAuthor="Alekseev, Vitalii I. &amp; Grzymala, Traci L." docDate="2015" docId="E7539306FFD4FF99FF0CFDEC4B1AF922" docLanguage="en" docName="zt03956p257.pdf" docOrigin="Zootaxa 3956 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Palaeocnopus densipunctatus Alekseev &amp; Grzymala, 2015, sp. nov." docType="treatment" docUuid="1DBA4CE1-A632-4940-A1E6-5191310511E1" docUuidSource="ZooBank" docVersion="8" lastPageNumber="251" masterDocId="1B6AEB7EFFDFFF95FF9BFFB2484DFFFF" masterDocTitle="New Aderidae (Coleoptera: Tenebrionoidea) from Baltic and Bitterfeld amber" masterLastPageNumber="257" masterPageNumber="239" pageNumber="250" updateTime="1698580836618" updateUser="plazi">
<mods:mods id="1EB7C042E7F06E98D5B88EE7941CB741" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="074C52C3A6679D7BA65F97FE004FF7B2">
<mods:title id="AFB14CC0352B873A1C1C9B0543E9483B">New Aderidae (Coleoptera: Tenebrionoidea) from Baltic and Bitterfeld amber</mods:title>
</mods:titleInfo>
<mods:name id="6DE56EC66C0083D0F4170CAEB00B3A20" type="personal">
<mods:role id="7A6E7342D52E90A41D5D00CC0F9A9258">
<mods:roleTerm id="C378C4DE4788E83CA712A4434019F0DF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EFE3D2D987F339AA9BBD4F202B854C9B">Alekseev, Vitalii I.</mods:namePart>
</mods:name>
<mods:name id="9261D13EA79FC5896282B8D3B1F3F2BF" type="personal">
<mods:role id="A827CD178171EA63CE4656D8531000AA">
<mods:roleTerm id="C1BC1F06CE5D36C03A11F44490EB5F9E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9FF170DCC59AEB44F1361F03169121D8">Grzymala, Traci L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A206BD1C22FDF3A3A9051DC7A65CC0EC">text</mods:typeOfResource>
<mods:relatedItem id="1D91E97FBAE8C13A011BBFDEB5152561" type="host">
<mods:titleInfo id="9BAADFE8788E1EABD6927E8160338FF3">
<mods:title id="025D5360C08107C7B5A0155D1EEA74FB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E48AB0FEB6C9CD8CDC69EE4526C61AE0">
<mods:date id="8D8E16E1D4F4522911157A6AFE07549A">2015</mods:date>
<mods:detail id="E1768D9D91EDF5340473D9821F060193" type="volume">
<mods:number id="D5922AB89A7DDC8119A7FDB7E1441488">3956</mods:number>
</mods:detail>
<mods:detail id="189C351F3EBBA25A32C5D14B1600C32E" type="issue">
<mods:number id="7CD9DEB245EA15FC9149C143E76628B4">2</mods:number>
</mods:detail>
<mods:extent id="D9E5A8494F77A78DF0C8768D92B61571" unit="page">
<mods:start id="67090FD3450E4D06C15975778701FE50">239</mods:start>
<mods:end id="024EBA79270E29BECE8DD15D4B48FED0">257</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="73B3E6D8A9C8067CAEF020EF502CA8A7">journal article</mods:classification>
<mods:identifier id="7F3095C2AF45B45D0F6465737A72EC54" type="DOI">10.11646/zootaxa.3956.2.5</mods:identifier>
<mods:identifier id="83748223A69EC8E24CECA5E2C50BBE43" type="GBIF-Dataset">693ef71c-2ee2-4da4-a3e1-35f5586899dc</mods:identifier>
<mods:identifier id="767EB2E53EFEBDF0D69794C5D2633C3D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4C8C2C07A9EF64A280AFB631A62AFF50" type="Zenodo-Dep">232228</mods:identifier>
<mods:identifier id="38F50630DD90C62F4964214909DDF668" type="ZooBank">C12EF13A-9C19-4051-80A6-F59B917774AA</mods:identifier>
</mods:mods>
<treatment id="E7539306FFD4FF99FF0CFDEC4B1AF922" ID-DOI="http://doi.org/10.5281/zenodo.5658595" ID-GBIF-Taxon="119612821" ID-Zenodo-Dep="5658595" LSID="urn:lsid:zoobank.org:act:1DBA4CE1-A632-4940-A1E6-5191310511E1" httpUri="http://treatment.plazi.org/id/E7539306FFD4FF99FF0CFDEC4B1AF922" lastPageId="12" lastPageNumber="251" pageId="11" pageNumber="250">
<subSubSection id="27E0719BFFD4FF9EFF0CFDEC4915FD65" pageId="11" pageNumber="250" type="nomenclature">
<paragraph id="6F452210FFD4FF9EFF0CFDEC4B31FD87" blockId="11.[151,892,606,666]" box="[151,892,606,632]" pageId="11" pageNumber="250">
<heading id="340D957CFFD4FF9EFF0CFDEC4B31FD87" bold="true" box="[151,892,606,632]" fontSize="11" level="1" pageId="11" pageNumber="250" reason="1">
<emphasis id="5D8EFE02FFD4FF9EFF0CFDEC4B31FD87" bold="true" box="[151,892,606,632]" pageId="11" pageNumber="250">
<taxonomicName id="A8FA5993FFD4FF9EFF0CFDEC49B0FD87" box="[151,509,606,632]" class="Insecta" family="Aderidae" genus="Palaeocnopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="250" phylum="Arthropoda" rank="species" species="densipunctatus" status="sp. nov.">
<emphasis id="5D8EFE02FFD4FF9EFF0CFDEC49B0FD87" bold="true" box="[151,509,606,632]" italics="true" pageId="11" pageNumber="250">Palaeocnopus densipunctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="46BD4379FFD4FF9EFD9FFDEC4A2FFD87" box="[516,610,606,632]" pageId="11" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
Alekseev &amp; Grzymala
</emphasis>
</heading>
</paragraph>
<paragraph id="6F452210FFD4FF9EFF0CFD334915FD65" blockId="11.[151,892,606,666]" box="[151,344,641,666]" pageId="11" pageNumber="250">
(
<figureCitation id="F7C13E95FFD4FF9EFF04FD3348B1FD65" box="[159,252,641,666]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1394,1417]" captionTargetBox="[239,1347,193,1372]" captionTargetId="figure@12.[239,1347,193,1373]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Palaeocnopus spp. illustrations. 21) Palaeocnopus densipunctatus sp. nov. A. forebody, B. antenna; 22) Palaeocnopus saeticornis sp. nov. A. forebody, B. antenna; 23) Palaeocnopus glabricornis sp. nov. A. forebody, B. antenna; 24) Palaeocnopus mara sp. nov. A. forebody, B. antenna." httpUri="https://zenodo.org/record/232237/files/figure.png" pageId="11" pageNumber="250">Figs. 21</figureCitation>
,
<figureCitation id="F7C13E95FFD4FF9EFE93FD33491DFD65" box="[264,336,641,666]" captionStart="FIGURES 25 – 27" captionStartId="13.[151,264,1354,1377]" captionTargetBox="[164,1416,769,1330]" captionTargetId="figure@13.[164,1422,769,1333]" captionTargetPageId="13" captionText="FIGURES 25 – 27. Palaeocnopus densipunctatus sp. nov. habitus photographs. 25) Holotype, lateral view; 26) Paratype (No. 217 - 1), dorsal view; 27) Paratype (No. 217 - 1), ventral view. Scale bars = 0.5 mm" httpUri="https://zenodo.org/record/232238/files/figure.png" pageId="11" pageNumber="250">25–27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="27E0719BFFD4FF9EFF0CFD7B4A63FB25" pageId="11" pageNumber="250" type="materials_examined">
<paragraph id="6F452210FFD4FF9EFF0CFD7B4A63FB25" blockId="11.[151,1437,713,2034]" pageId="11" pageNumber="250">
<emphasis id="5D8EFE02FFD4FF9EFF0CFD7B49CFFD1D" bold="true" box="[151,386,713,738]" pageId="11" pageNumber="250">Material examined.</emphasis>
<typeStatus id="B0419CB2FFD4FF9EFE10FD7B49BCFD1D" box="[395,497,713,738]" pageId="11" pageNumber="250" type="holotype">
<emphasis id="5D8EFE02FFD4FF9EFE10FD7B49BCFD1D" box="[395,497,713,738]" italics="true" pageId="11" pageNumber="250">Holotype</emphasis>
</typeStatus>
No. 420-6 [
<collectionCode id="09EBBAD5FFD4FF9EFDE4FD784A98FD1D" box="[639,725,714,738]" pageId="11" pageNumber="250">CCHH</collectionCode>
], possible female (due to small eyes and short antennae) (
<figureCitation id="F7C13E95FFD4FF9EFAF4FD7B48FAFCF9" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1394,1417]" captionTargetBox="[239,1347,193,1372]" captionTargetId="figure@12.[239,1347,193,1373]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Palaeocnopus spp. illustrations. 21) Palaeocnopus densipunctatus sp. nov. A. forebody, B. antenna; 22) Palaeocnopus saeticornis sp. nov. A. forebody, B. antenna; 23) Palaeocnopus glabricornis sp. nov. A. forebody, B. antenna; 24) Palaeocnopus mara sp. nov. A. forebody, B. antenna." httpUri="https://zenodo.org/record/232237/files/figure.png" pageId="11" pageNumber="250">Fig. 22</figureCitation>
). The beetle inclusion is preserved in a polished piece of transparent amber with a yellowish shade and the amber piece is embedded in polyester resin. The syninclusions are represented by two trichomes and one phorid fly (Diptera) of length
<quantity id="A8028FF5FFD4FF9EFEEEFC84499DFCB2" box="[373,464,822,846]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="250" unit="mm" value="1.2">1.2 mm</quantity>
.
<typeStatus id="B0419CB2FFD4FF9EFE47FC844A0EFCB2" box="[476,579,822,845]" pageId="11" pageNumber="250" type="paratype">
<emphasis id="5D8EFE02FFD4FF9EFE47FC844A0EFCB2" box="[476,579,822,845]" italics="true" pageId="11" pageNumber="250">Paratype</emphasis>
</typeStatus>
No. 217-1 [
<collectionCode id="09EBBAD5FFD4FF9EFD55FC844B69FCB1" box="[718,804,822,846]" pageId="11" pageNumber="250">CCHH</collectionCode>
], possible female (
<figureCitation id="F7C13E95FFD4FF9EFC61FC874CC8FCB1" box="[1018,1157,821,846]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1394,1417]" captionTargetBox="[239,1347,193,1372]" captionTargetId="figure@12.[239,1347,193,1373]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Palaeocnopus spp. illustrations. 21) Palaeocnopus densipunctatus sp. nov. A. forebody, B. antenna; 22) Palaeocnopus saeticornis sp. nov. A. forebody, B. antenna; 23) Palaeocnopus glabricornis sp. nov. A. forebody, B. antenna; 24) Palaeocnopus mara sp. nov. A. forebody, B. antenna." httpUri="https://zenodo.org/record/232237/files/figure.png" pageId="11" pageNumber="250">Figs. 23–24</figureCitation>
). The beetle inclusion is preserved in a polished piece of transparent amber, yellow in color, without any further fixation. The amber piece is parallelepiped, with maximum length
<quantity id="A8028FF5FFD4FF9EFDDBFCCF4AC2FC6A" box="[576,655,893,918]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="250" unit="mm" value="16.0">16 mm</quantity>
and maximum width
<quantity id="A8028FF5FFD4FF9EFC19FCCC4B8BFC6A" box="[898,966,894,918]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="250" unit="mm" value="9.0">9 mm</quantity>
. The syninclusions are represented by one well-preserved rove beetle (
<taxonomicName id="A8FA5993FFD4FF9EFE7AFC134A2FFC45" box="[481,610,929,954]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="250" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="A8FA5993FFD4FF9EFDE9FC134B5DFC45" box="[626,784,929,954]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="250" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
:
<taxonomicName id="A8FA5993FFD4FF9EFCBAFC134BD4FC45" box="[801,921,929,954]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="250" phylum="Arthropoda" rank="subFamily" subFamily="Omaliinae">Omaliinae</taxonomicName>
, aff.
<taxonomicName id="A8FA5993FFD4FF9EFC47FC134D4BFC45" authority="Heer, 1839" authorityName="Heer" authorityYear="1839" box="[988,1286,929,954]" class="Insecta" family="Staphylinidae" genus="Phloeonomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="250" phylum="Arthropoda" rank="genus">
<emphasis id="5D8EFE02FFD4FF9EFC47FC134C38FC45" box="[988,1141,929,954]" italics="true" pageId="11" pageNumber="250">Phloeonomus</emphasis>
Heer, 1839
</taxonomicName>
, possibly an undescribed species, personal observation) of length
<quantity id="A8028FF5FFD4FF9EFD61FC744B1BFC22" box="[762,854,966,990]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="250" unit="mm" value="1.7">1.7 mm</quantity>
.
<typeStatus id="B0419CB2FFD4FF9EFCFFFC744B86FC22" box="[868,971,966,989]" pageId="11" pageNumber="250" type="paratype">
<emphasis id="5D8EFE02FFD4FF9EFCFFFC744B86FC22" box="[868,971,966,989]" italics="true" pageId="11" pageNumber="250">Paratype</emphasis>
</typeStatus>
No. AWI-107 [
<collectionCode id="09EBBAD5FFD4FF9EFB19FC744C83FC21" box="[1154,1230,966,990]" pageId="11" pageNumber="250">CVIA</collectionCode>
], possible female. The beetle is included in a polished piece of transparent yellow amber, without any further fixation. The piece is flat, rounded, with maximum length
<quantity id="A8028FF5FFD4FF9EFDABFBBC4A32FBDA" box="[560,639,1038,1062]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="250" unit="mm" value="19.0">19 mm</quantity>
and maximum width
<quantity id="A8028FF5FFD4FF9EFCE9FBBC4B89FBDA" box="[882,964,1038,1062]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="250" unit="mm" value="14.0">14 mm</quantity>
. The syninclusions are represented by two trichomes.
<typeStatus id="B0419CB2FFD4FF9EFE8EFB804931FBB6" box="[277,380,1074,1097]" pageId="11" pageNumber="250" type="paratype">
<emphasis id="5D8EFE02FFD4FF9EFE8EFB804931FBB6" box="[277,380,1074,1097]" italics="true" pageId="11" pageNumber="250">Paratype</emphasis>
</typeStatus>
No. 25-4 [
<collectionCode id="09EBBAD5FFD4FF9EFE62FB804A02FBB5" box="[505,591,1074,1098]" pageId="11" pageNumber="250">CCHH</collectionCode>
], possible female. The beetle inclusion is preserved in a polished piece of transparent amber with an orange shade. The amber piece is embedded in polyester resin.
<typeStatus id="B0419CB2FFD4FF9EFB08FBE44CB7FB92" box="[1171,1274,1110,1133]" pageId="11" pageNumber="250" type="paratype">
<emphasis id="5D8EFE02FFD4FF9EFB08FBE44CB7FB92" box="[1171,1274,1110,1133]" italics="true" pageId="11" pageNumber="250">Paratype</emphasis>
</typeStatus>
No. AWI-121 [
<collectionCode id="09EBBAD5FFD4FF9EFF06FBC848A4FB6D" box="[157,233,1146,1170]" pageId="11" pageNumber="250">CVIA</collectionCode>
], possible female. The beetle is included in a polished piece of transparent yellow amber, without any further fixation. The piece is flat, with maximum length
<quantity id="A8028FF5FFD4FF9EFC90FB2C4B17FB4A" box="[779,858,1182,1206]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="11" pageNumber="250" unit="mm" value="23.0">23 mm</quantity>
and maximum width
<quantity id="A8028FF5FFD4FF9EFBD6FB2C4CD2FB4A" box="[1101,1183,1182,1206]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="250" unit="mm" value="17.0">17 mm</quantity>
. The syninclusions are represented by numerous trichomes.
</paragraph>
</subSubSection>
<subSubSection id="27E0719BFFD4FF9EFF5CFB574BD6FADE" pageId="11" pageNumber="250" type="etymology">
<paragraph id="6F452210FFD4FF9EFF5CFB574BD6FADE" blockId="11.[151,1437,713,2034]" pageId="11" pageNumber="250">
<emphasis id="5D8EFE02FFD4FF9EFF5CFB57490BFB01" bold="true" box="[199,326,1253,1278]" pageId="11" pageNumber="250">Etymology</emphasis>
. This species is named from a combination of Latin “
<emphasis id="5D8EFE02FFD4FF9EFC31FB574BBBFB01" box="[938,1014,1253,1278]" italics="true" pageId="11" pageNumber="250">densus</emphasis>
”[dense] and Latin “
<emphasis id="5D8EFE02FFD4FF9EFB46FB544D70FB02" box="[1245,1341,1254,1277]" italics="true" pageId="11" pageNumber="250">punctum</emphasis>
” [point, puncture], referring to the dense and coarse punctation on the vertex.
</paragraph>
</subSubSection>
<subSubSection id="27E0719BFFD4FF9EFF5CFA9F49D3FA71" pageId="11" pageNumber="250" type="materials_examined">
<paragraph id="6F452210FFD4FF9EFF5CFA9F4A16FABA" blockId="11.[151,1437,713,2034]" box="[199,603,1325,1350]" pageId="11" pageNumber="250">
<emphasis id="5D8EFE02FFD4FF9EFF5CFA9F4903FAB9" bold="true" box="[199,334,1325,1350]" pageId="11" pageNumber="250">
<typeStatus id="B0419CB2FFD4FF9EFF5CFA9F494FFAB9" box="[199,258,1325,1350]" pageId="11" pageNumber="250">Type</typeStatus>
strata
</emphasis>
. Baltic Amber. Eocene.
</paragraph>
<paragraph id="6F452210FFD4FF9EFF5CFAE349D3FA71" blockId="11.[151,1437,713,2034]" pageId="11" pageNumber="250">
<emphasis id="5D8EFE02FFD4FF9EFF5CFAE3492AFA95" bold="true" box="[199,359,1361,1386]" pageId="11" pageNumber="250">
<typeStatus id="B0419CB2FFD4FF9EFF5CFAE3494FFA95" box="[199,258,1361,1386]" pageId="11" pageNumber="250">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="17ED6280FFD4FF9EFEE5FAE34986FA95" box="[382,459,1361,1386]" name="Russia" pageId="11" pageNumber="250">Russia</collectingCountry>
, the Kaliningrad region, the Sambian [Samland] peninsula, Yantarny settlement [formerly Palmnicken].
</paragraph>
</subSubSection>
<subSubSection id="27E0719BFFD4FF99FF5CFA2B4CE7F9F9" lastPageId="12" lastPageNumber="251" pageId="11" pageNumber="250" type="description">
<paragraph id="6F452210FFD4FF9EFF5CFA2B4904F9E1" blockId="11.[151,1437,713,2034]" pageId="11" pageNumber="250">
<emphasis id="5D8EFE02FFD4FF9EFF5CFA2B491CFA4D" bold="true" box="[199,337,1433,1458]" pageId="11" pageNumber="250">Description</emphasis>
. Length approximately
<quantity id="A8028FF5FFD4FF9EFDF3FA284ABFFA4E" box="[616,754,1434,1458]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="11" pageNumber="250" unit="mm" value="1.6" valueMax="1.8" valueMin="1.4">1.4–1.8 mm</quantity>
(No. 420-6—
<quantity id="A8028FF5FFD4FF9EFC03FA2B4C4CFA4E" box="[920,1025,1433,1458]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="11" pageNumber="250" unit="mm" value="1.75">1.75 mm</quantity>
; No. 217—
<quantity id="A8028FF5FFD4FF9EFB10FA284CA8FA4E" box="[1163,1253,1434,1458]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="250" unit="mm" value="1.7">1.7 mm</quantity>
; AWI-107—
<quantity id="A8028FF5FFD4FF9EFAE7FA28488BFA2A" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="250" unit="mm" value="1.8">1.8 mm</quantity>
; No. 25-4—
<quantity id="A8028FF5FFD4FF9EFEC7FA0F49F5FA2A" box="[348,440,1469,1494]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="250" unit="mm" value="1.5">1.5 mm</quantity>
; No. AWI-121—
<quantity id="A8028FF5FFD4FF9EFD1FFA0C4AACFA29" box="[644,737,1470,1494]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="250" unit="mm" value="1.4">1.4 mm</quantity>
); moderately convex, elongate; uniformly dark gray. Upper surface uniseriate, clothed with short pubescence. Body length 2.7× maximum body width. Elytral length 4.1× pronotal length.
</paragraph>
<paragraph id="6F452210FFD4FF9EFF5CF99B4DDBF951" blockId="11.[151,1437,713,2034]" pageId="11" pageNumber="250">
<emphasis id="5D8EFE02FFD4FF9EFF5CF99B4949F9BD" box="[199,260,1577,1602]" italics="true" pageId="11" pageNumber="250">Head</emphasis>
. Eyes large, oval, hemispherical, coarsely faceted, adjacent to hind margin of head; interocular space wider than one ocular diameter; temples narrow; apical maxillary palpomere acute, triangular; apical labial palpomere oval. Vertex densely and coarsely punctate. Antenna (
<figureCitation id="F7C13E95FFD4FF9EFC38F9C34BB6F975" box="[931,1019,1649,1674]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1394,1417]" captionTargetBox="[239,1347,193,1372]" captionTargetId="figure@12.[239,1347,193,1373]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Palaeocnopus spp. illustrations. 21) Palaeocnopus densipunctatus sp. nov. A. forebody, B. antenna; 22) Palaeocnopus saeticornis sp. nov. A. forebody, B. antenna; 23) Palaeocnopus glabricornis sp. nov. A. forebody, B. antenna; 24) Palaeocnopus mara sp. nov. A. forebody, B. antenna." httpUri="https://zenodo.org/record/232237/files/figure.png" pageId="11" pageNumber="250">Fig. 21</figureCitation>
B) filiform, 11-segmented, sparsely pubescent; reaching basal third of elytra when folded backward; antennomere length ratios: 5-4-4-4-5-5-5-4-4-4-6.
</paragraph>
<paragraph id="6F452210FFD4FF9EFF5CF90B4999F87A" blockId="11.[151,1437,713,2034]" pageId="11" pageNumber="250">
<emphasis id="5D8EFE02FFD4FF9EFF5CF90B4955F92D" box="[199,280,1721,1746]" italics="true" pageId="11" pageNumber="250">Thorax</emphasis>
. Pronotum transverse (
<figureCitation id="F7C13E95FFD4FF9EFDB2F90B4A33F92D" box="[553,638,1721,1746]" captionStart="FIGURES 21 – 24" captionStartId="12.[151,264,1394,1417]" captionTargetBox="[239,1347,193,1372]" captionTargetId="figure@12.[239,1347,193,1373]" captionTargetPageId="12" captionText="FIGURES 21 – 24. Palaeocnopus spp. illustrations. 21) Palaeocnopus densipunctatus sp. nov. A. forebody, B. antenna; 22) Palaeocnopus saeticornis sp. nov. A. forebody, B. antenna; 23) Palaeocnopus glabricornis sp. nov. A. forebody, B. antenna; 24) Palaeocnopus mara sp. nov. A. forebody, B. antenna." httpUri="https://zenodo.org/record/232237/files/figure.png" pageId="11" pageNumber="250">Fig. 21</figureCitation>
A), widest basally; glabrous; coarsely and densely punctate; without visible pubescence; with two deep, well-defined sub-basal impressions, almost fused medially, and with two deep sub-triangular lateral impressions at anterior angles. Posterior angles rounded. Scutellum subquadrate, with posterior angles rounded. Elytron subparallel, moderately convex, with slight depression on disc; width 0.52× length; punctation irregular, coarse, dense. Elytral pubescence short, more visible laterally and apically. Metathoracic wings present.
</paragraph>
<paragraph id="6F452210FFD4FF9EFF5CF82348B2F80D" blockId="11.[151,1437,713,2034]" pageId="11" pageNumber="250">
<emphasis id="5D8EFE02FFD4FF9EFF5CF823497CF855" box="[199,305,1937,1962]" italics="true" pageId="11" pageNumber="250">Abdomen</emphasis>
. Each abdominal ventrite with large deep punctures, punctures subequal to those on pro-, meso- and metathorax. Separation of abdominal ventrite I and abdominal ventrite II fine, but well-defined laterally, obsolete medially.
</paragraph>
<caption id="3B857298FFD3FF99FF0CFAC04AADFA3A" httpUri="https://zenodo.org/record/232237/files/figure.png" pageId="12" pageNumber="251" targetBox="[239,1347,193,1372]" targetPageId="12">
<paragraph id="6F452210FFD3FF99FF0CFAC04AADFA3A" blockId="12.[151,1436,1394,1478]" pageId="12" pageNumber="251">
<emphasis id="5D8EFE02FFD3FF99FF0CFAC0491BFA78" bold="true" box="[151,342,1394,1417]" pageId="12" pageNumber="251">FIGURES 21–24.</emphasis>
<taxonomicName id="A8FA5993FFD3FF99FEC4FAC049A2FA77" box="[351,495,1394,1416]" class="Insecta" family="Aderidae" genus="Palaeocnopus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="249" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="5D8EFE02FFD3FF99FEC4FAC049A2FA77" box="[351,495,1394,1416]" italics="true" pageId="12" pageNumber="251">Palaeocnopus</emphasis>
</taxonomicName>
spp. illustrations.
<emphasis id="5D8EFE02FFD3FF99FD2BFAC04A9FFA76" bold="true" box="[688,722,1394,1417]" pageId="12" pageNumber="251">21)</emphasis>
<taxonomicName id="A8FA5993FFD3FF99FD40FAC04C46FA77" box="[731,1035,1394,1416]" class="Insecta" family="Aderidae" genus="Palaeocnopus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="251" phylum="Arthropoda" rank="species" species="densipunctatus" status="sp. nov.">
<emphasis id="5D8EFE02FFD3FF99FD40FAC04C46FA77" box="[731,1035,1394,1416]" italics="true" pageId="12" pageNumber="251">Palaeocnopus densipunctatus</emphasis>
</taxonomicName>
<emphasis id="5D8EFE02FFD3FF99FB8FFAC14CC4FA78" bold="true" box="[1044,1161,1394,1416]" pageId="12" pageNumber="251">
<taxonomicNameLabel id="46BD4379FFD3FF99FB8FFAC14C25FA77" box="[1044,1128,1395,1416]" pageId="12" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
forebody,
<emphasis id="5D8EFE02FFD3FF99FB60FAC14D5FFA77" bold="true" box="[1275,1298,1395,1416]" pageId="12" pageNumber="251">B.</emphasis>
antenna;
<emphasis id="5D8EFE02FFD3FF99FAE1FAC04DD6FA76" bold="true" box="[1402,1435,1394,1417]" pageId="12" pageNumber="251">22)</emphasis>
<taxonomicName id="A8FA5993FFD3FF99FF0CFA2349D0FA58" box="[151,413,1425,1447]" class="Insecta" family="Aderidae" genus="Palaeocnopus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="251" phylum="Arthropoda" rank="species" species="saeticornis" status="sp. nov.">
<emphasis id="5D8EFE02FFD3FF99FF0CFA2349D0FA58" box="[151,413,1425,1447]" italics="true" pageId="12" pageNumber="251">Palaeocnopus saeticornis</emphasis>
</taxonomicName>
<emphasis id="5D8EFE02FFD3FF99FE3FFA234A5BFA59" bold="true" box="[420,534,1425,1446]" pageId="12" pageNumber="251">
<taxonomicNameLabel id="46BD4379FFD3FF99FE3FFA2349BBFA59" box="[420,502,1425,1446]" pageId="12" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
forebody,
<emphasis id="5D8EFE02FFD3FF99FD1EFA234AD0FA59" bold="true" box="[645,669,1425,1446]" pageId="12" pageNumber="251">B.</emphasis>
antenna;
<emphasis id="5D8EFE02FFD3FF99FC9AFA224B6EFA58" bold="true" box="[769,803,1424,1447]" pageId="12" pageNumber="251">23)</emphasis>
<taxonomicName id="A8FA5993FFD3FF99FCB0FA234C72FA58" box="[811,1087,1425,1447]" class="Insecta" family="Aderidae" genus="Palaeocnopus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="251" phylum="Arthropoda" rank="species" species="glabricornis" status="sp. nov.">
<emphasis id="5D8EFE02FFD3FF99FCB0FA234C72FA58" box="[811,1087,1425,1447]" italics="true" pageId="12" pageNumber="251">Palaeocnopus glabricornis</emphasis>
</taxonomicName>
<emphasis id="5D8EFE02FFD3FF99FBDDFA234CF5FA59" bold="true" box="[1094,1208,1425,1446]" pageId="12" pageNumber="251">
<taxonomicNameLabel id="46BD4379FFD3FF99FBDDFA234CD5FA59" box="[1094,1176,1425,1446]" pageId="12" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
forebody,
<emphasis id="5D8EFE02FFD3FF99FABCFA234D72FA59" bold="true" box="[1319,1343,1425,1446]" pageId="12" pageNumber="251">B.</emphasis>
antenna;
<emphasis id="5D8EFE02FFD3FF99FF0CFA1D48F5FA39" bold="true" box="[151,184,1455,1478]" pageId="12" pageNumber="251">24)</emphasis>
<taxonomicName id="A8FA5993FFD3FF99FF24FA1D49C4FA39" box="[191,393,1455,1478]" class="Insecta" family="Aderidae" genus="Palaeocnopus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="251" phylum="Arthropoda" rank="species" species="mara" status="sp. nov.">
<emphasis id="5D8EFE02FFD3FF99FF24FA1D49C4FA39" box="[191,393,1455,1478]" italics="true" pageId="12" pageNumber="251">Palaeocnopus mara</emphasis>
</taxonomicName>
<emphasis id="5D8EFE02FFD3FF99FE0BFA024A4DFA3B" bold="true" box="[400,512,1455,1477]" pageId="12" pageNumber="251">
<taxonomicNameLabel id="46BD4379FFD3FF99FE0BFA0249ACFA3A" box="[400,481,1456,1477]" pageId="12" pageNumber="251" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
forebody,
<emphasis id="5D8EFE02FFD3FF99FDF6FA024AC9FA3A" bold="true" box="[621,644,1456,1477]" pageId="12" pageNumber="251">B.</emphasis>
antenna.
</paragraph>
</caption>
<paragraph id="6F452210FFD3FF99FF5CFA5C4CE7F9F9" blockId="12.[151,1436,1517,1758]" box="[199,1194,1517,1542]" pageId="12" pageNumber="251">
<emphasis id="5D8EFE02FFD3FF99FF5CFA5C48B1F9FA" box="[199,252,1518,1541]" italics="true" pageId="12" pageNumber="251">Legs</emphasis>
. Metatarsomere I approximately equal in length to metatarsomeres II-IV combined.
</paragraph>
</subSubSection>
<subSubSection id="27E0719BFFD3FF99FF5CF9A34B1AF922" pageId="12" pageNumber="251" type="diagnosis">
<paragraph id="6F452210FFD3FF99FF5CF9A34B1AF922" blockId="12.[151,1436,1517,1758]" pageId="12" pageNumber="251">
<emphasis id="5D8EFE02FFD3FF99FF5CF9A34977F9D5" bold="true" box="[199,314,1553,1578]" pageId="12" pageNumber="251">Diagnosis</emphasis>
.
<taxonomicName id="A8FA5993FFD3FF99FED0F9A34AD6F9D5" box="[331,667,1553,1578]" class="Insecta" family="Aderidae" genus="Palaeocnopus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="251" phylum="Arthropoda" rank="species" species="densipunctatus" status="sp. nov.">
<emphasis id="5D8EFE02FFD3FF99FED0F9A34AD6F9D5" box="[331,667,1553,1578]" italics="true" pageId="12" pageNumber="251">Palaeocnopus densipunctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="46BD4379FFD3FF99FD3FF9A34B4DF9D5" box="[676,768,1553,1578]" pageId="12" pageNumber="251" rank="species">
<emphasis id="5D8EFE02FFD3FF99FD3FF9A34B4DF9D5" bold="true" box="[676,768,1553,1578]" pageId="12" pageNumber="251">sp. nov.</emphasis>
</taxonomicNameLabel>
is characterized by the antennal ratios, the two deep subbasal pronotal impressions, and the densely, coarsely punctate vertex.
<taxonomicName id="A8FA5993FFD3FF99FC47F9874D7DF9B1" box="[988,1328,1589,1614]" class="Insecta" family="Aderidae" genus="Palaeocnopus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="251" phylum="Arthropoda" rank="species" species="densipunctatus" status="sp. nov.">
<emphasis id="5D8EFE02FFD3FF99FC47F9874D7DF9B1" box="[988,1328,1589,1614]" italics="true" pageId="12" pageNumber="251">Palaeocnopus densipunctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="46BD4379FFD3FF99FAA7F9874DD1F9B1" box="[1340,1436,1589,1614]" pageId="12" pageNumber="251" rank="species">
<emphasis id="5D8EFE02FFD3FF99FAA7F9874DD1F9B1" bold="true" box="[1340,1436,1589,1614]" pageId="12" pageNumber="251">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="A8FA5993FFD3FF99FEB8F9E84985F98D" box="[291,456,1625,1650]" class="Insecta" family="Aderidae" genus="Palaeocnopus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="251" phylum="Arthropoda" rank="species" species="glabricornis">
<emphasis id="5D8EFE02FFD3FF99FEB8F9E84985F98D" box="[291,456,1625,1650]" italics="true" pageId="12" pageNumber="251">P. glabricornis</emphasis>
</taxonomicName>
and
<emphasis id="5D8EFE02FFD3FF99FE65F9E84A1CF98E" box="[510,593,1626,1649]" italics="true" pageId="12" pageNumber="251">P. m ar a</emphasis>
by the punctation of the head and pronotum, and from
<emphasis id="5D8EFE02FFD3FF99FB20F9E84D00F98E" box="[1211,1357,1626,1649]" italics="true" pageId="12" pageNumber="251">P. s ae t i c or n i s</emphasis>
by the two separate pronotal impressions and absence of long pubescence on the antennae and elytra. Antennal ratios of this new species are similar to
<emphasis id="5D8EFE02FFD3FF99FE75F9104A0FF946" box="[494,578,1698,1721]" italics="true" pageId="12" pageNumber="251">P. m a r a</emphasis>
, but the presence of the two closed sub-basal pronotal impressions makes
<taxonomicName id="A8FA5993FFD3FF99FA1CF9104972F921" class="Insecta" family="Aderidae" genus="Palaeocnopus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="251" phylum="Arthropoda" rank="species" species="densipunctatus" status="sp. nov.">
<emphasis id="5D8EFE02FFD3FF99FA1CF9104972F921" italics="true" pageId="12" pageNumber="251">P. densipunctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="46BD4379FFD3FF99FEDDF97749D2F922" box="[326,415,1733,1758]" pageId="12" pageNumber="251" rank="species">
<emphasis id="5D8EFE02FFD3FF99FEDDF97749D5F921" bold="true" box="[326,408,1733,1758]" pageId="12" pageNumber="251">sp. nov</emphasis>
.
</taxonomicNameLabel>
appear more similar to
<taxonomicName id="A8FA5993FFD3FF99FD31F9744B1DF921" box="[682,848,1733,1758]" class="Insecta" family="Aderidae" genus="Palaeocnopus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="251" phylum="Arthropoda" rank="species" species="glabricornis">
<emphasis id="5D8EFE02FFD3FF99FD31F9744B1DF921" box="[682,848,1733,1758]" italics="true" pageId="12" pageNumber="251">P. glabricornis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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