237 lines
27 KiB
XML
237 lines
27 KiB
XML
<document id="DE5B764FB661017C99B316C85C7B906B" ID-DOI="10.11646/zootaxa.5405.2.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="10603393" ID-ZooBank="DD57E713-2FD1-4A05-8E5B-13A0D27A0B5D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1706773369097" checkinUser="plazi" docAuthor="Hong, Yong, Inkhavilay, Khamla & James, Samuel W." docDate="2024" docId="E73187B3813C6F6D799C2BA7E49625B0" docLanguage="en" docName="zootaxa.5405.2.7.pdf" docOrigin="Zootaxa 5405 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5405.2.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amynthas piyakarnae Hong and James 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="278" masterDocId="1B08FFCB81396F6B790B2B30E07A2565" masterDocTitle="Three new earthworms of the Amynthas corticis-group (Clitellata: Megascolecidae) from the Nam Et-Phouleoi, Laos" masterLastPageNumber="280" masterPageNumber="273" pageNumber="278" updateTime="1706773890757" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9C6A4A8E86AC014972BC9B3A10FBC475">Three new earthworms of the Amynthas corticis-group (Clitellata: Megascolecidae) from the Nam Et-Phouleoi, Laos</mods:title>
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<mods:name id="D5EC1544FB51AF307D9453D3875EC98C" type="personal">
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<mods:role id="FC6D86C9CA3FD088DEC87230BDFD403C">
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<mods:roleTerm id="9DED718235A4CB998ABE5B0906AE53B1">Author</mods:roleTerm>
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<mods:namePart id="E8FE2532CC67212CEE192444B1E45CFE">Hong, Yong</mods:namePart>
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<mods:affiliation id="923D15C3A7329743453FE6F0ACC535C4">Department of Agricultural Biology, College of Agriculture & Life Sciences, Jeonbuk National University, Jeonju 54896, Korea;</mods:affiliation>
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<mods:name id="CA8BD8E777565BAB2C46EDA37C34CB89" type="personal">
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<mods:roleTerm id="8EFA12A10E96F8B2FAFC0979F430960E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B599E0BECD324CA6B123BB953CC11109">Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation id="CAEEB7AFD50A4A90F1861D349E418BCC">Center of Excellence in Biodiversity, Research Academic and Service Office, National University of Laos;</mods:affiliation>
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<mods:roleTerm id="5A859F5E7A700A99C7596970707C03D2">Author</mods:roleTerm>
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<mods:namePart id="D45F256A4C60FFF85D04A32C148753AC">James, Samuel W.</mods:namePart>
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<mods:affiliation id="9C564ABA957AECACC4C9454413818005">Department of Biology, University of Iowa, Iowa City, Iowa, USA 52242.</mods:affiliation>
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<mods:title id="7F28EFDF8F6F6CCF4C59724B11BEAC33">Zootaxa</mods:title>
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<mods:part id="8DFFC4216AB93D026F5794D0864197F9">
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<mods:date id="4199C45CC205EC1C27A567601F757F77">2024</mods:date>
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<mods:number id="0330EE60C37059F4C457EA00FB8A8845">2024-01-30</mods:number>
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<mods:number id="9D803E32F1AA91483E0C0992792387F6">5405</mods:number>
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<mods:number id="2C1366A35D4CD9BFFB307935B160298F">2</mods:number>
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<mods:url id="403F21959F3E5DFB2611CF24CA4BBC2B">http://dx.doi.org/10.11646/zootaxa.5405.2.7</mods:url>
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<mods:classification id="7A8665DE2D467C0F963A222283C69EB1">journal article</mods:classification>
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<mods:identifier id="5754BAACC40E2EB3B42DED54DD4D8D38" type="DOI">10.11646/zootaxa.5405.2.7</mods:identifier>
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<mods:identifier id="4891EEA064D230CF117772B558ED933D" type="ISSN">1175-5326</mods:identifier>
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<treatment id="E73187B3813C6F6D799C2BA7E49625B0" ID-DOI="http://doi.org/10.5281/zenodo.10603407" ID-Zenodo-Dep="10603407" LSID="urn:lsid:plazi:treatment:E73187B3813C6F6D799C2BA7E49625B0" httpUri="http://treatment.plazi.org/id/E73187B3813C6F6D799C2BA7E49625B0" lastPageId="6" lastPageNumber="278" pageId="5" pageNumber="278">
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<subSubSection id="2782652E813C6F6E799C2BA7E2A725D4" box="[151,733,151,178]" pageId="5" pageNumber="278" type="nomenclature">
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<paragraph id="6F2736A5813C6F6E799C2BA7E2A725D4" blockId="5.[151,733,151,213]" box="[151,733,151,178]" pageId="5" pageNumber="278">
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<heading id="346F81C9813C6F6E799C2BA7E2A725D4" bold="true" box="[151,733,151,178]" fontSize="11" level="1" pageId="5" pageNumber="278" reason="1">
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<emphasis id="5DECEAB7813C6F6E799C2BA7E2A725D4" bold="true" box="[151,733,151,178]" pageId="5" pageNumber="278">
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<taxonomicName id="A8984D26813C6F6E799C2BA7E20225D4" authority="Hong and James" authorityName="Hong and James" authorityYear="2024" box="[151,632,151,178]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="278" phylum="Annelida" rank="species" species="piyakarnae" status="sp. nov.">
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<emphasis id="5DECEAB7813C6F6E799C2BA7E1DA25D7" bold="true" box="[151,416,151,178]" italics="true" pageId="5" pageNumber="278">Amynthas piyakarnae</emphasis>
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||
Hong and James
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||
</taxonomicName>
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||
<taxonomicNameLabel id="46DF57CC813C6F6E7B742BA7E2A725D4" box="[639,733,151,177]" pageId="5" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="2782652E813C6F6E799C2B8BE09125B0" box="[151,235,187,213]" pageId="5" pageNumber="278" type="description">
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<paragraph id="6F2736A5813C6F6E799C2B8BE09125B0" blockId="5.[151,733,151,213]" box="[151,235,187,213]" pageId="5" pageNumber="278">
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(
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<figureCitation id="F7A32A20813C6F6E79942B8BE09925B0" box="[159,227,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1218,1242]" captionTargetBox="[471,1111,259,1161]" captionTargetId="figure-52@6.[466,1123,234,1197]" captionTargetPageId="6" captionText="FIGURE 4. Amynthas piyakarnae sp. nov. (A, B). A: ventral view; B: intestinal caeca; C: spermatheca. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10603405" httpUri="https://zenodo.org/record/10603405/files/figure.png" pageId="5" pageNumber="278">Fig. 4</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="2782652E813C6F6E799C2A33E3B224EC" pageId="5" pageNumber="278" type="materials_examined">
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||
<paragraph id="6F2736A5813C6F6E799C2A33E3B224EC" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="278">
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||
<emphasis id="5DECEAB7813C6F6E799C2A33E17C2478" bold="true" box="[151,262,259,285]" pageId="5" pageNumber="278">Material.</emphasis>
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||
<materialsCitation id="DFF03CF8813C6F6E78072A33E3952400" collectingDate="2005-07-23" collectionCode="BDNUL" collectorName="Boutong & Phimphet & Son & K. Inkhavilay" country="Laos" county="Viengthong District" elevation="2273" latitude="20.27205" location="Summit of Phouleoi Mountain" longLatPrecision="1" longitude="103.02953" municipality="Nam Et-Phouleoi" pageId="5" pageNumber="278" specimenCode="BDNUL 0060" specimenCount="1" stateProvince="Houaphan" typeStatus="holotype">
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<typeStatus id="B0238807813C6F6E78072A33E1032478" box="[268,377,259,285]" pageId="5" pageNumber="278" type="holotype">Holotype</typeStatus>
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||
: 1 clitellate (
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||
<specimenCode id="3F3E9EDE813C6F6E7B082A33E2D22478" box="[515,680,259,285]" collectionCode="BDNUL" pageId="5" pageNumber="278">BDNUL 0060</specimenCode>
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):
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<collectingCountry id="178F7635813C6F6E7BB32A34E28B247B" box="[696,753,260,286]" name="Laos" pageId="5" pageNumber="278">Laos</collectingCountry>
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,
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||
<collectingRegion id="AD5CF847813C6F6E7BF02A33E3922478" box="[763,1000,259,285]" country="Laos" name="Houaphan" pageId="5" pageNumber="278">Houaphanh Province</collectingRegion>
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||
,
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<collectingCounty id="86464E29813C6F6E7AF92A33E4AB2478" box="[1010,1233,259,285]" pageId="5" pageNumber="278">Viengthong District</collectingCounty>
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||
,
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<collectingMunicipality id="8F43ACDF813C6F6E7DD12A34E5E62478" box="[1242,1436,259,286]" pageId="5" pageNumber="278">Nam Et-Phouleoi</collectingMunicipality>
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NPA,
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||
<location id="6A47607E813C6F6E79D62A17E2412424" LSID="urn:lsid:plazi:treatment:E73187B3813C6F6D799C2BA7E49625B0:6A47607E813C6F6E79D62A17E2412424" box="[221,571,295,321]" country="Laos" county="Viengthong District" latitude="20.27205" longLatPrecision="1" longitude="103.02953" municipality="Nam Et-Phouleoi" name="Summit of Phouleoi Mountain" pageId="5" pageNumber="278" stateProvince="Houaphan">Summit of Phouleoi Mountain</location>
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||
,
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||
<quantity id="A8609B40813C6F6E7B432A17E2DC2427" box="[584,678,295,322]" metricMagnitude="3" metricUnit="m" metricValue="2.273" pageId="5" pageNumber="278" unit="m" value="2273.0">
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||
<elevation id="E4B5D196813C6F6E7B432A17E2DC2427" box="[584,678,295,322]" metricMagnitude="3" metricUnit="m" metricValue="2.273" pageId="5" pageNumber="278" unit="m" value="2273.0">2,273 m</elevation>
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||
</quantity>
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||
(
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||
<geoCoordinate id="0AAC5062813C6F6E7BBC2A19E3302421" box="[695,842,297,324]" degrees="20" direction="north" minutes="16.323" orientation="latitude" pageId="5" pageNumber="278" precision="1" value="20.27205">20°16.323′N</geoCoordinate>
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||
,
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||
<geoCoordinate id="0AAC5062813C6F6E7A5C2A19E39C2421" box="[855,998,297,324]" degrees="103" direction="east" minutes="1.772" orientation="longitude" pageId="5" pageNumber="278" precision="1" value="103.02953">103°1.772′E</geoCoordinate>
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), wet mossy forest, very thick organic soil surface,
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||
<date id="1B261065813C6F6E782F2A7BE1C42403" box="[292,446,331,358]" pageId="5" pageNumber="278" value="2005-07-23">
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||
<collectingDate id="0B62E98D813C6F6E782F2A7BE1C42403" box="[292,446,331,358]" pageId="5" pageNumber="278" value="2005-07-23">July 23, 2005</collectingDate>
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||
</date>
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||
,
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||
<collectorName id="C26D5373813C6F6E78C32A7CE2512403" box="[456,555,332,358]" pageId="5" pageNumber="278">Boutong</collectorName>
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,
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<collectorName id="C26D5373813C6F6E7B3D2A7BE2DE2400" box="[566,676,331,357]" pageId="5" pageNumber="278">Phimphet</collectorName>
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,
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<collectorName id="C26D5373813C6F6E7BA42A7BE2A72400" box="[687,733,331,357]" pageId="5" pageNumber="278">Son</collectorName>
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, and
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<collectorName id="C26D5373813C6F6E7A1C2A7CE3CB2400" box="[791,945,331,358]" pageId="5" pageNumber="278">K. Inkhavilay</collectorName>
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colls
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</materialsCitation>
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.
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<materialsCitation id="DFF03CF8813C6F6E7AF22A7CE1E824EC" collectingDate="2005-07-23" collectionCode="BDNUL" collectorName="Boutong & Phimphet & Son & K. Inkhavilay" country="Laos" county="Viengthong District" elevation="2273" latitude="20.27205" location="Summit of Phouleoi Mountain" longLatPrecision="1" longitude="103.02953" municipality="Nam Et-Phouleoi" pageId="5" pageNumber="278" specimenCode="BDNUL 0061" specimenCount="1" stateProvince="Houaphan" typeStatus="paratype">
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<typeStatus id="B0238807813C6F6E7AF22A7CE4252403" box="[1017,1119,332,358]" pageId="5" pageNumber="278" type="paratype">Paratype</typeStatus>
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: 1 clitellate (
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<specimenCode id="3F3E9EDE813C6F6E7DE72A7BE5EA2400" box="[1260,1424,331,357]" collectionCode="BDNUL" pageId="5" pageNumber="278">BDNUL 0061</specimenCode>
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), same data as holotype
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</materialsCitation>
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.
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<materialsCitation id="DFF03CF8813C6F6E78972A40E3BF24EC" box="[412,965,367,394]" collectingDate="2005-07-23" collectionCode="BDNUL" collectorName="Boutong & Phimphet & Son & K. Inkhavilay" country="Laos" county="Viengthong District" elevation="2273" latitude="20.27205" location="Summit of Phouleoi Mountain" longLatPrecision="1" longitude="103.02953" municipality="Nam Et-Phouleoi" pageId="5" pageNumber="278" specimenCount="1" stateProvince="Houaphan" typeStatus="paratype">Non-types: 1 clitellate, same data as for holotype</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="2782652E813C6F6E79CC2AA3E4C624C8" box="[199,1212,403,429]" pageId="5" pageNumber="278" type="etymology">
|
||
<paragraph id="6F2736A5813C6F6E79CC2AA3E4C624C8" blockId="5.[151,1437,259,2014]" box="[199,1212,403,429]" pageId="5" pageNumber="278">
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||
<emphasis id="5DECEAB7813C6F6E79CC2AA3E13424C8" bold="true" box="[199,334,403,429]" pageId="5" pageNumber="278">Etymology.</emphasis>
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The species is named after the second author’s daughter, Piyakarn Inkhavilay.
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</paragraph>
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</subSubSection>
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<subSubSection id="2782652E813C6F6E79CC2A87E3512704" pageId="5" pageNumber="278" type="diagnosis">
|
||
<paragraph id="6F2736A5813C6F6E79CC2A87E3512704" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="278">
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||
<emphasis id="5DECEAB7813C6F6E79CC2A87E13824B4" bold="true" box="[199,322,439,465]" pageId="5" pageNumber="278">Diagnosis.</emphasis>
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Spermathecal pores paired in 5/6–8/9 at 9
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<superScript id="98ED9BED813C6F6E7A332A87E33F24A0" attach="left" box="[824,837,439,453]" fontSize="6" pageId="5" pageNumber="278">th</superScript>
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setal line; male pores superficial at lateral ends of transverse elongate oval pads elevated equatorially on XVIII, pads surrounded by 3–4 small outer rings, 0.25–0.31 circumference apart; genital markings circular, pre-setal VIII, IX; row of 3–5 small round genital papillae pre-setal in XVIII, evenly spaced; 2 or 4 genital papillae in row, pre-setal in XIX; pinnate intestinal caeca, additional small caecal lobes on XXVIII, first dorsal pores in 5/6 or nearby.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="2782652E813C6F6E79CC295BE53620A4" pageId="5" pageNumber="278" type="description">
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||
<paragraph id="6F2736A5813C6F6E79CC295BE1772638" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="278">
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<emphasis id="5DECEAB7813C6F6E79CC295BE12327E0" bold="true" box="[199,345,619,645]" pageId="5" pageNumber="278">Description.</emphasis>
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Dimensions
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||
<quantity id="A8609B40813C6F6E78FA295BE20127E3" box="[497,635,619,646]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" metricValueMax="1.42" metricValueMin="0.98" pageId="5" pageNumber="278" unit="mm" value="120.0" valueMax="142.0" valueMin="98.0">98–142 mm</quantity>
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||
by 5.0–
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||
<quantity id="A8609B40813C6F6E7BD0295BE34827E3" box="[731,818,619,646]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="278" unit="mm" value="5.5">5.5 mm</quantity>
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||
at segment X,
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||
<quantity id="A8609B40813C6F6E7AEF295BE41427E3" box="[996,1134,619,646]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.2" metricValueMin="4.5" pageId="5" pageNumber="278" unit="mm" value="4.85" valueMax="5.2" valueMin="4.5">4.5–5.2 mm</quantity>
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||
at segment XXX,
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||
<quantity id="A8609B40813C6F6E7C43295BE0B927CF" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.7" metricValueMin="4.5" pageId="5" pageNumber="278" unit="mm" value="4.6" valueMax="4.7" valueMin="4.5">4.5–4.7 mm</quantity>
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||
at clitellum; body cylindrical, segments 66–129 (possibly some natural amputees). Setae regularly distributed around segmental equators, numbering 34–40 at VII, 57–58 at XX, 13–16 between the male pores, setal formula AA:AB:ZZ:YZ = 4:3:5:3 at XIII. Female pore single XIV on
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||
<quantity id="A8609B40813C6F6E7A6A29E7E3902797" box="[865,1002,727,754]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="278" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
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oval porophore. Prostomium epilobic with tongue open, pigment dark red-brown on dorsal anterior, ventral I–IV only; clitellum coffee color, formalin preservation. First dorsal pore at 5/6 (2 individuals), 6/7 (1 individual). Clitellum annular XIV–XVI; setae invisible externally.
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||
</paragraph>
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||
<paragraph id="6F2736A5813C6F6E79CC2857E142213C" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="278">
|
||
Male pores superficial at lateral margins of ventrum in XVIII at 9
|
||
<superScript id="98ED9BED813C6F6E7AB32857E3BF2610" attach="left" box="[952,965,871,885]" fontSize="6" pageId="5" pageNumber="278">th</superScript>
|
||
setal line, 0.25–0.31 circumference apart ventrally at lateral ends of transverse elongate elevated oval pads equatorial on XVIII, pads surrounded by 3– 4 small outer rings; row of 3–5 small round genital papillae pre-setal in XVIII, evenly spaced; 2 or 4 genital papillae in a row, pre-setal in XIX, 2 pre-setal in XX (one individual). Spermathecal pores paired in 5/6–8/9 at 9
|
||
<superScript id="98ED9BED813C6F6E7C9B28E3E5E72684" attach="none" box="[1424,1437,979,993]" fontSize="6" pageId="5" pageNumber="278">th</superScript>
|
||
setal line, inconspicuous at lateral margins of ventrum, 0.27–0.29 circumference apart ventrally. Genital markings circular, paired pre-setal VIII and IX (2 individuals); pre-setal mid-ventral VIII and IX plus single left pre-setal IX (1 individual).
|
||
</paragraph>
|
||
<paragraph id="6F2736A5813C6F6E79CC2F53E4A92030" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="278">Septa 5/6/7 thick, 7/8 thinly muscular, 8/9/10 absent, 10/11–12/13 thinly muscular. Gizzard large in VIII– X. Intestine beginning in XV, lymph glands not found. Typhlosole thick low ridge from XXVII. Intestinal caeca originating in XXVII and extending anteriorly about to XXV, pinnate, each divided into 13–17 small finger-shaped lobes; 2–3 dorsal caecal lobes in XXVIII. Hearts esophageal X–XIII. Male sexual system holandric, testes and funnels in ventrally joined sacs in X and XI. Seminal vesicles paired in XI and XII; dorsal lobes lacking. Prostates in XVIII, and extending from XVII–XXII; glandular portions consist of 2 main lobes each divided again into 2–4 small lobes. Genital papillae glands in segments XVIII and XIX with very short stalks or sessile.</paragraph>
|
||
<paragraph id="6F2736A5813C6F6E79CC2E6FE53620A4" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="278">Ovaries in XIII. Spermathecae paired VI–IX; 2 pairs in VII; ampulla oval, small black spots on surface, slender ducts shorter than ampulla; diverticulum chamber chili-shaped, with stalk as long as ampulla attached near body wall, no nephridia on ducts. Mushroom-shaped genital marking glands in VIII and IX on ventral body wall.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2782652E813C6F6D79CC2EFBE49625B0" lastPageId="6" lastPageNumber="279" pageId="5" pageNumber="278" type="discussion">
|
||
<paragraph id="6F2736A5813C6F6E79CC2EFBE303236F" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="278">
|
||
<emphasis id="5DECEAB7813C6F6E79CC2EFBE1412080" bold="true" box="[199,315,1483,1509]" pageId="5" pageNumber="278">Remarks.</emphasis>
|
||
Like the other species described here, the distinctions from the main body of the
|
||
<taxonomicName id="A8984D26813C6F6E7DD62EFBE5282080" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[1245,1362,1483,1509]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="278" phylum="Annelida" rank="species" species="corticis">
|
||
<emphasis id="5DECEAB7813C6F6E7DD62EFBE5282080" box="[1245,1362,1483,1509]" italics="true" pageId="5" pageNumber="278">A. corticis</emphasis>
|
||
</taxonomicName>
|
||
-group are the pinnate caeca and the forward position of the dorsal pores.
|
||
</paragraph>
|
||
<paragraph id="6F2736A5813C6F6E79CC2D23E2622260" blockId="5.[151,1437,259,2014]" pageId="5" pageNumber="278">
|
||
The new species similarly bears some superficial similarities to
|
||
<taxonomicName id="A8984D26813C6F6E7A872D23E4042348" baseAuthorityName="Thai et Samphon" baseAuthorityYear="1988" box="[908,1150,1555,1581]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="278" phylum="Annelida" rank="species" species="duplidiverticulatus">
|
||
<emphasis id="5DECEAB7813C6F6E7A872D23E4042348" box="[908,1150,1555,1581]" italics="true" pageId="5" pageNumber="278">A. duplidiverticulatus</emphasis>
|
||
</taxonomicName>
|
||
, with respect to the shape of the male pore region and presence of genital papilla between the male pores. However,
|
||
<taxonomicName id="A8984D26813C6F6E7DA32D07E5E72334" baseAuthorityName="Thai et Samphon" baseAuthorityYear="1988" box="[1192,1437,1591,1617]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="278" phylum="Annelida" rank="species" species="duplidiverticulatus">
|
||
<emphasis id="5DECEAB7813C6F6E7DA32D07E5E72334" box="[1192,1437,1591,1617]" italics="true" pageId="5" pageNumber="278">A. duplidiverticulatus</emphasis>
|
||
</taxonomicName>
|
||
has a greater number of setae compared to the new species (81 vs. 34–40 at VIII, and 66 at XIX vs. 58 at XX).
|
||
<taxonomicName id="A8984D26813C6F6E799C2D4FE1F623FC" baseAuthorityName="Thai et Samphon" baseAuthorityYear="1988" box="[151,396,1663,1689]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="278" phylum="Annelida" rank="species" species="duplidiverticulatus">
|
||
<emphasis id="5DECEAB7813C6F6E799C2D4FE1F623FC" box="[151,396,1663,1689]" italics="true" pageId="5" pageNumber="278">A. duplidiverticulatus</emphasis>
|
||
</taxonomicName>
|
||
has simple intestinal caeca, but
|
||
<taxonomicName id="A8984D26813C6F6E7BF52D4FE38923FC" authority="Hong & Inkhavilay & James, 2024" authorityName="Hong & Inkhavilay & James" authorityYear="2024" box="[766,1011,1663,1689]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="278" phylum="Annelida" rank="species" species="piyakarnae" status="sp. nov.">
|
||
<emphasis id="5DECEAB7813C6F6E7BF52D4FE38923FC" box="[766,1011,1663,1689]" italics="true" pageId="5" pageNumber="278">Amynthas piyakarnae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="5DECEAB7813C6F6E7AF72D4FE42D23FC" bold="true" box="[1020,1111,1663,1689]" pageId="5" pageNumber="278">
|
||
<taxonomicNameLabel id="46DF57CC813C6F6E7AF72D4FE42D23FC" box="[1020,1111,1663,1689]" pageId="5" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has pinnate intestinal caeca. Spermathecal forms differ and
|
||
<taxonomicName id="A8984D26813C6F6E7B0B2D93E28C23D8" baseAuthorityName="Thai et Samphon" baseAuthorityYear="1988" box="[512,758,1699,1725]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="278" phylum="Annelida" rank="species" species="duplidiverticulatus">
|
||
<emphasis id="5DECEAB7813C6F6E7B0B2D93E28C23D8" box="[512,758,1699,1725]" italics="true" pageId="5" pageNumber="278">A. duplidiverticulatus</emphasis>
|
||
</taxonomicName>
|
||
lacks genital marking glands, but the new species has the mushroom-shaped genital marking glands in VIII and IX on ventral body wall. The male pore region and genital papillae are unique to this species.
|
||
</paragraph>
|
||
<paragraph id="6F2736A5813C6F6D79CC2C3FE49625B0" blockId="5.[151,1437,259,2014]" lastBlockId="6.[151,1436,151,213]" lastPageId="6" lastPageNumber="279" pageId="5" pageNumber="278">
|
||
The
|
||
<collectingCountry id="178F7635813C6F6E79F12C20E14A224F" box="[250,304,1808,1834]" name="Laos" pageId="5" pageNumber="278">Laos</collectingCountry>
|
||
<taxonomicName id="A8984D26813C6F6E78322C3FE1D6224C" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[313,428,1807,1833]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="278" phylum="Annelida" rank="species" species="corticis">
|
||
<emphasis id="5DECEAB7813C6F6E78322C3FE1D6224C" box="[313,428,1807,1833]" italics="true" pageId="5" pageNumber="278">A. corticis</emphasis>
|
||
</taxonomicName>
|
||
-group appears in various forms. In particular, individuals of many species show different external characteristics, such as genital markings, and genital papillae. Even in the same location, they show different appearances depending on their habitat, or even on the same mountain in one region, depending on altitude. Even in the genus
|
||
<taxonomicName id="A8984D26813C6F6E780D2C4BE10922F0" authorityName="Kinberg" authorityYear="1867" box="[262,371,1915,1941]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="278" phylum="Annelida" rank="genus">
|
||
<emphasis id="5DECEAB7813C6F6E780D2C4BE10922F0" box="[262,371,1915,1941]" italics="true" pageId="5" pageNumber="278">Amynthas</emphasis>
|
||
</taxonomicName>
|
||
, the
|
||
<taxonomicName id="A8984D26813C6F6E78A22C4CE18622F0" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[425,508,1916,1941]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="278" phylum="Annelida" rank="species" species="corticis">
|
||
<emphasis id="5DECEAB7813C6F6E78A22C4CE18622F0" box="[425,508,1916,1941]" italics="true" pageId="5" pageNumber="278">corticis</emphasis>
|
||
</taxonomicName>
|
||
-group, which has four pairs of spermathecal pores, is highly variable. In the
|
||
<collectingCountry id="178F7635813C6F6E7C432C4CE50422F3" box="[1352,1406,1916,1942]" name="Laos" pageId="5" pageNumber="278">Laos</collectingCountry>
|
||
<taxonomicName id="A8984D26813C6F6E7C8F2C4BE09022DC" baseAuthorityName="Kinberg" baseAuthorityYear="1866" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="5" pageNumber="278" phylum="Annelida" rank="species" species="corticis">
|
||
<emphasis id="5DECEAB7813C6F6E7C8F2C4BE09022DC" italics="true" pageId="5" pageNumber="278">A. corticis</emphasis>
|
||
</taxonomicName>
|
||
-group, the morphology of the intestinal caeca was important characteristic. Along with the simple, manicate, and pinnate
|
||
<typeStatus id="B0238807813C6F6E78162CF4E12022BB" box="[285,346,1988,2014]" pageId="5" pageNumber="278">types</typeStatus>
|
||
, it is one of the characteristics necessary for species recognition. In future research, research on the mutual evolution between these different
|
||
<typeStatus id="B0238807813F6F6D7B6A2BA7E2E625D4" box="[609,668,151,177]" pageId="6" pageNumber="279">types</typeStatus>
|
||
through molecular biological analysis of intestinal caeca
|
||
<typeStatus id="B0238807813F6F6D7C1E2BA7E52A25D4" box="[1301,1360,151,177]" pageId="6" pageNumber="279">types</typeStatus>
|
||
within the genus
|
||
<taxonomicName id="A8984D26813F6F6D78032B8BE1B525B1" baseAuthorityName="Kinberg" baseAuthorityYear="1866" box="[264,463,187,213]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Crassiclitellata" pageId="6" pageNumber="279" phylum="Annelida" rank="species" species="corticis">
|
||
<emphasis id="5DECEAB7813F6F6D78032B8BE1B525B1" box="[264,463,187,213]" italics="true" pageId="6" pageNumber="279">Amynthas corticis</emphasis>
|
||
</taxonomicName>
|
||
-group is expected to be an important clue for understanding this group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |