247 lines
22 KiB
XML
247 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.81.57792" ID-GBIF-Dataset="7c1bea40-8c93-4ba1-9faa-97fcf81ffdad" ID-Pensoft-Pub="1314-2607-81-43" ID-Pensoft-UUID="D48C8138C7E35800953C86D433828718" ID-Zenodo-Dep="4584245" ID-ZooBank="6E3EC3E6F3EF4AA59B2A10C4D719CE19" ModsDocID="1314-2607-81-43" checkinTime="1614373120647" checkinUser="pensoft" docAuthor="Olmi, Massimo, Chen, Hua-Yan, Shih, Chungkun, Mueller, Patrick, Capradossi, Leonardo, Ren, Dong, Perkovsky, Evgeny E. & Guglielmino, Adalgisa" docDate="2021" docId="E6A1EA2BC18958CB96CEA03C0F49BC3A" docLanguage="en" docName="JourHymenoptRes 81: 43-55" docOrigin="Journal of Hymenoptera Research 81" docPubDate="2021-02-25" docSource="http://dx.doi.org/10.3897/jhr.81.57792" docTitle="Hybristodryinus castaneus Olmi, Guglielmino & Chen 2021, sp. nov." docType="treatment" docUuid="5CA10E8A-2DE2-4FD6-B4C4-9D3F580BA92F" docUuidSource="ZooBank" docVersion="4" id="D48C8138C7E35800953C86D433828718" lastPageNumber="43" masterDocId="D48C8138C7E35800953C86D433828718" masterDocTitle="New species of Hybristodryinus Engel (Hymenoptera, Dryinidae) from mid-Cretaceous amber of northern Myanmar, with notes on their possible hosts" masterLastPageNumber="55" masterPageNumber="43" pageNumber="43" updateTime="1678755185687" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species of Hybristodryinus Engel (Hymenoptera, Dryinidae) from mid-Cretaceous amber of northern Myanmar, with notes on their possible hosts</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Olmi, Massimo</mods:namePart>
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<mods:affiliation>Tropical Entomology Research Center, Viterbo 01100, Italy</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chen, Hua-Yan</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0382-1635</mods:nameIdentifier>
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<mods:affiliation>State Key Laboratory of Biocontrol, School of Life Sciences / School of Ecology, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
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<mods:nameIdentifier type="email">huayanc@scbg.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shih, Chungkun</mods:namePart>
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<mods:affiliation>College of Life Sciences, Capital Normal University, Haidian District, Beijing 100048, China & Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013 - 7012, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mueller, Patrick</mods:namePart>
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<mods:affiliation>Friedhofstrasse 9, Kaeshofen 66894, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Capradossi, Leonardo</mods:namePart>
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<mods:affiliation>Via Pericle Scriboni 28, Tuscania 01017, Italy</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ren, Dong</mods:namePart>
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<mods:affiliation>College of Life Sciences, Capital Normal University, Haidian District, Beijing 100048, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Perkovsky, Evgeny E.</mods:namePart>
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<mods:affiliation>Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev 01601, Ukraine & Borissiak Paleontological Institute of the Russian Academy of Sciences, Moscow 117997, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Guglielmino, Adalgisa</mods:namePart>
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<mods:affiliation>Department of Agriculture and Forest Sciences (DAFNE), University of Tuscia, Viterbo 01100, Italy</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-02-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>81</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>43</mods:start>
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<mods:end>55</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/jhr.81.57792</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.81.57792</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2607-81-43</mods:identifier>
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<mods:identifier type="ZooBank">6E3EC3E6F3EF4AA59B2A10C4D719CE19</mods:identifier>
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<mods:identifier type="Pensoft-UUID">D48C8138C7E35800953C86D433828718</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4584245</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="179918099" LSID="urn:lsid:zoobank.org:act:5CA10E8A-2DE2-4FD6-B4C4-9D3F580BA92F" httpUri="http://treatment.plazi.org/id/E6A1EA2BC18958CB96CEA03C0F49BC3A" lastPageNumber="43" pageId="0" pageNumber="43">
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<subSubSection pageId="0" pageNumber="43" type="nomenclature">
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<paragraph pageId="0" pageNumber="43">
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<taxonomicName LSID="http://zoobank.org/5CA10E8A-2DE2-4FD6-B4C4-9D3F580BA92F" authority="Olmi, Guglielmino & Chen" authorityName="Olmi, Guglielmino & Chen" authorityYear="2021" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hybristodryinus castaneus" order="Hymenoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="castaneus" status="sp. nov.">Hybristodryinus castaneus Olmi, Guglielmino & Chen</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="43">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="description">
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<paragraph pageId="0" pageNumber="43">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hybristodryinus castaneus sp. nov., male, holotype A habitus, dorsal view B habitus, ventral view C head and mesosoma, dorsal view D head and mesosoma, ventral view E antenna F fore wing." figureDoi="10.3897/jhr.81.57792.figure1" httpUri="https://binary.pensoft.net/fig/512986" pageId="0" pageNumber="43">Figure 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="diagnosis">
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<paragraph pageId="0" pageNumber="43">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" pageId="0" pageNumber="43">Male</emphasis>
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: macropterous, with antennomere 9 about six times as long as broad (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hybristodryinus castaneus sp. nov., male, holotype A habitus, dorsal view B habitus, ventral view C head and mesosoma, dorsal view D head and mesosoma, ventral view E antenna F fore wing." figureDoi="10.3897/jhr.81.57792.figure1" httpUri="https://binary.pensoft.net/fig/512986" pageId="0" pageNumber="43">1E</figureCitation>
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); face without long deep longitudinal furrow (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hybristodryinus castaneus sp. nov., male, holotype A habitus, dorsal view B habitus, ventral view C head and mesosoma, dorsal view D head and mesosoma, ventral view E antenna F fore wing." figureDoi="10.3897/jhr.81.57792.figure1" httpUri="https://binary.pensoft.net/fig/512986" pageId="0" pageNumber="43">1D</figureCitation>
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); posterior margin of vertex not deeply excavated medially (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hybristodryinus castaneus sp. nov., male, holotype A habitus, dorsal view B habitus, ventral view C head and mesosoma, dorsal view D head and mesosoma, ventral view E antenna F fore wing." figureDoi="10.3897/jhr.81.57792.figure1" httpUri="https://binary.pensoft.net/fig/512986" pageId="0" pageNumber="43">1C</figureCitation>
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); lateral ocelli not touching occipital carina; notauli complete, posteriorly separated (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hybristodryinus castaneus sp. nov., male, holotype A habitus, dorsal view B habitus, ventral view C head and mesosoma, dorsal view D head and mesosoma, ventral view E antenna F fore wing." figureDoi="10.3897/jhr.81.57792.figure1" httpUri="https://binary.pensoft.net/fig/512986" pageId="0" pageNumber="43">1C</figureCitation>
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).
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</paragraph>
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<caption ID-Zenodo-Dep="4584247" doi="10.3897/jhr.81.57792.figure1" httpUri="https://binary.pensoft.net/fig/512986" pageId="0" pageNumber="43" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" pageId="0" pageNumber="43">Figure 1.</emphasis>
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<taxonomicName authorityName="Olmi, Guglielmino & Chen" authorityYear="2021" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hybristodryinus castaneus" order="Hymenoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="castaneus">
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<emphasis italics="true" pageId="0" pageNumber="43">Hybristodryinus castaneus</emphasis>
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</taxonomicName>
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sp. nov., male, holotype
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<emphasis bold="true" pageId="0" pageNumber="43">A</emphasis>
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habitus, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="43">B</emphasis>
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habitus, ventral view
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<emphasis bold="true" pageId="0" pageNumber="43">C</emphasis>
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head and mesosoma, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="43">D</emphasis>
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head and mesosoma, ventral view
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<emphasis bold="true" pageId="0" pageNumber="43">E</emphasis>
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antenna
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<emphasis bold="true" pageId="0" pageNumber="43">F</emphasis>
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fore wing.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="description">
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<paragraph pageId="0" pageNumber="43">Description.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" pageId="0" pageNumber="43">Male.</emphasis>
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Macropterous (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hybristodryinus castaneus sp. nov., male, holotype A habitus, dorsal view B habitus, ventral view C head and mesosoma, dorsal view D head and mesosoma, ventral view E antenna F fore wing." figureDoi="10.3897/jhr.81.57792.figure1" httpUri="https://binary.pensoft.net/fig/512986" pageId="0" pageNumber="43">1A, B</figureCitation>
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); length 1.4 mm. Completely dark brown. Antenna filiform, about as long as body, with setae very short. Scape about as broad as pedicel. Antennomeres in following proportions: 5:4:8:9:9:9:8:8:6:7. Antennomere 9 about six times as long as broad (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hybristodryinus castaneus sp. nov., male, holotype A habitus, dorsal view B habitus, ventral view C head and mesosoma, dorsal view D head and mesosoma, ventral view E antenna F fore wing." figureDoi="10.3897/jhr.81.57792.figure1" httpUri="https://binary.pensoft.net/fig/512986" pageId="0" pageNumber="43">1E</figureCitation>
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). Head with sculpture not visible; face without long deep median longitudinal furrow; mandible with teeth not visible; frontal line not visible; occipital carina complete, laterally not reaching eyes; ocellar ratio: POL:OL:OOL:OPL = 1:1:2:1; temple more than twice as long as OOL (5:2); greatest breadth of lateral ocellus as long as OOL; posterior margin of vertex not excavated medially (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hybristodryinus castaneus sp. nov., male, holotype A habitus, dorsal view B habitus, ventral view C head and mesosoma, dorsal view D head and mesosoma, ventral view E antenna F fore wing." figureDoi="10.3897/jhr.81.57792.figure1" httpUri="https://binary.pensoft.net/fig/512986" pageId="0" pageNumber="43">1C</figureCitation>
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); occiput deeply excavated. Palpal formula 6/3. Mesosoma much longer than head (30:10) and metasoma (30:20). Pronotum with lateral regions not protruding. Mesoscutum shiny, with sculpture not visible, slightly shorter than head (9:10), much longer than mesoscutellum (9:5). Notauli complete, posteriorly separated (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hybristodryinus castaneus sp. nov., male, holotype A habitus, dorsal view B habitus, ventral view C head and mesosoma, dorsal view D head and mesosoma, ventral view E antenna F fore wing." figureDoi="10.3897/jhr.81.57792.figure1" httpUri="https://binary.pensoft.net/fig/512986" pageId="0" pageNumber="43">1C</figureCitation>
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); minimum distance between notauli longer than OOL (3:2). Mesoscutellum shiny, with sculpture not visible. Metanotum and metapectal-propodeal complex not visible. Epicnemium not visible. Mesopleuron not visible. Fore wing hyaline, without dark transverse bands (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Hybristodryinus castaneus sp. nov., male, holotype A habitus, dorsal view B habitus, ventral view C head and mesosoma, dorsal view D head and mesosoma, ventral view E antenna F fore wing." figureDoi="10.3897/jhr.81.57792.figure1" httpUri="https://binary.pensoft.net/fig/512986" pageId="0" pageNumber="43">1F</figureCitation>
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). Stigmal vein (2r-rs&Rs) with angle between proximal and distal parts, with distal part much longer than pterostigma (12:8). Second radial 1 cell (2R1) open. Pterostigma much longer than broad (9:2). Metacarpus (PostabR1) slightly shorter than pterostigma (7:9). Fore leg ratio:? (coxa not visible, because it is hidden by an air bubble): 5 (trochanter): 16 (femur): 12 (tibia): 8 (protarsomere 1): 3 (protarsomere 2): 3 (protarsomere 3): 2 (protarsomere 4): 4 (protarsomere 5). Protrochanter more than four times as long as broad (5:1). Mid leg ratio: 7 (coxa): 4 (trochanter): 17 (femur): 15 (tibia): 9 (mesotarsomere 1): 3 (mesotarsomere 2): 2 (mesotarsomere 3): 1.5 (mesotarsomere 4): 3 (mesotarsomere 5). Mesotrochanter more than twice as long as broad (4:1.5). Hind leg ratio: 8 (coxa): 4 (trochanter): 17 (femur): 21 (tibia): 10 (metatarsomere 1): 4 (metatarsomere 2): 3 (metatarsomere 3): 2 (metatarsomere 4): 3 (metatarsomere 5). Tibial spurs 1/1/2.
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</paragraph>
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<paragraph pageId="0" pageNumber="43">
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<emphasis bold="true" pageId="0" pageNumber="43">Female.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="materials_examined">
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<paragraph pageId="0" pageNumber="43">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Myanmar" county="Burmese" location="Northern" municipality="Obtained" specimenCount="1" stateProvince="Kachin State" typeStatus="Holotype">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="43">
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<typeStatus>Holotype</typeStatus>
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:
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</emphasis>
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male (DAF3869),
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<collectingCountry name="Myanmar">Myanmar</collectingCountry>
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: specimen in mid-Cretaceous
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<collectingCounty>Burmese</collectingCounty>
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amber (about 99 Ma).
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<collectingMunicipality>Obtained</collectingMunicipality>
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from a mine situated in
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<location LSID="urn:lsid:plazi:treatment:E6A1EA2BC18958CB96CEA03C0F49BC3A:A907D8D3CD74E489585971350C1E215F" country="Myanmar" county="Burmese" municipality="Obtained" name="Northern" stateProvince="Kachin State">Northern</location>
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<collectingCountry name="Myanmar">Myanmar</collectingCountry>
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,
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<collectingRegion country="Myanmar" name="Kachin">Kachin State</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:E6A1EA2BC18958CB96CEA03C0F49BC3A:3A6A364C7228594F5202D0BD0D3E968D" country="Myanmar" county="Burmese" municipality="Obtained" name="Tanai Township" stateProvince="Kachin State">Tanai Township</location>
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,
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<location LSID="urn:lsid:plazi:treatment:E6A1EA2BC18958CB96CEA03C0F49BC3A:EFA5FA8F26A31EBDB834C2CD71EAF240" country="Myanmar" county="Burmese" municipality="Obtained" name="Hukawng Valley" stateProvince="Kachin State">Hukawng Valley</location>
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,
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<locationDeviation location="Tanai City">SW of Tanai City</locationDeviation>
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(DAF).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="etymology">
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<paragraph pageId="0" pageNumber="43">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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The species is named
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<taxonomicName authorityName="Olmi, Guglielmino & Chen" authorityYear="2021" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hybristodryinus castaneus" order="Hymenoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="castaneus">
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<emphasis italics="true" pageId="0" pageNumber="43">Hybristodryinus castaneus</emphasis>
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</taxonomicName>
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(Latin adjective meaning
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<normalizedToken originalValue="“brown”">"brown"</normalizedToken>
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) because of its dark brown colour.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="43" type="remarks">
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<paragraph pageId="0" pageNumber="43">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="43">
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<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. castaneus" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="castaneus">
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<emphasis italics="true" pageId="0" pageNumber="43">H. castaneus</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. konbaung" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="konbaung">
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<emphasis italics="true" pageId="0" pageNumber="43">H. konbaung</emphasis>
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</taxonomicName>
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Perkovsky, Olmi,
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<normalizedToken originalValue="Müller">Mueller</normalizedToken>
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& Martynova, 2019, known from Burmese amber, for the following characters: face without a long deep longitudinal furrow, lateral ocelli not touching the occipital carina, posterior margin of the vertex not excised medially, notauli complete and separated posteriorly. However, the antenna of
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<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. castaneus" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="castaneus">
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<emphasis italics="true" pageId="0" pageNumber="43">H. castaneus</emphasis>
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</taxonomicName>
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is slenderer, with antennomere 9 about six times as long as broad (less slender in
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<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. konbaung" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="konbaung">
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<emphasis italics="true" pageId="0" pageNumber="43">H. konbaung</emphasis>
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</taxonomicName>
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, with antennomere 9 about twice as long as broad).
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<bibRefCitation DOI="https://doi.org/10.1016/j.cretres.2019.01.023" author="Perkovsky, EE" journalOrPublisher="Cretaceous Research" pageId="0" pageNumber="43" pagination="169 - 189" refId="B21" refString="Perkovsky, EE, Olmi, M, Mueller, P, Martynova, KV, 2019. A review of the genus Hybristodryinus Engel, 2005 (Hymenoptera, Dryinidae) from mid-Cretaceous Burmese amber, with a discussion on its phylogenetic significance. Cretaceous Research 99: 169 - 189, DOI: https://doi.org/10.1016/j.cretres.2019.01.023" title="A review of the genus Hybristodryinus Engel, 2005 (Hymenoptera, Dryinidae) from mid-Cretaceous Burmese amber, with a discussion on its phylogenetic significance." url="https://doi.org/10.1016/j.cretres.2019.01.023" volume="99" year="2019">Perkovsky et al. (2019)</bibRefCitation>
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indicate "pronotum with lateral regions protruding" among the characters of the generic diagnosis. However, this character is not visible in all known males (as in
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<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. pyu" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="pyu">
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<emphasis italics="true" pageId="0" pageNumber="43">H. pyu</emphasis>
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</taxonomicName>
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Perkovsky, Olmi,
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<normalizedToken originalValue="Müller">Mueller</normalizedToken>
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& Martynova, 2019) and it is not present in
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<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. castaneus" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="castaneus">
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<emphasis italics="true" pageId="0" pageNumber="43">H. castaneus</emphasis>
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</taxonomicName>
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sp. nov. (see description above). Except this character,
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<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. castaneus" order="Hemiptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="castaneus">
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<emphasis italics="true" pageId="0" pageNumber="43">H. castaneus</emphasis>
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</taxonomicName>
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meets with all other generic diagnosis characters of
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<taxonomicName authorityName="Engel" authorityYear="2005" class="Insecta" family="Dryinidae" genus="Hybristodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hybristodryinus" order="Hymenoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="43">Hybristodryinus</emphasis>
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</taxonomicName>
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. Therefore, we update the diagnosis of the genus by excluding this character.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |