269 lines
31 KiB
XML
269 lines
31 KiB
XML
<document id="AC2BD57635CC67837EE423E1D493D310" ID-CLB-Dataset="2767" ID-DOI="10.11646/zootaxa.5087.3.2" ID-GBIF-Dataset="43c1b1e7-1797-4ed7-817d-606114b807ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="5826858" ID-ZooBank="6253FEED-A4FB-4832-AE51-91E4F51655C4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641539129469" checkinUser="plazi" docAuthor="Paz, Fernando, Ratcliffe, Brett C. & Figueroa, Luis" docDate="2022" docId="E61687C1FFA2FFD17BB4BCE9FB121B3F" docLanguage="en" docName="zootaxa.5087.3.2.pdf" docOrigin="Zootaxa 5087 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyclocephala mateoi Paz & Ratcliffe 2022, new species" docType="treatment" docUuid="CCED91AA-7B00-4830-B6E2-BF5E648294A7" docUuidSource="ZooBank" docVersion="6" lastPageNumber="431" masterDocId="1A2FFFB9FFA1FFD57B23BF3BFFAE1A5A" masterDocTitle="Three new species of Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae) from Amazonian Peru and a checklist of Cyclocephala species in Peru" masterLastPageNumber="440" masterPageNumber="427" pageNumber="430" updateTime="1699296570230" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="D30EB1A23D2D07B97618AE9BDBC2376B" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="5468B95FC8BCDD7E19E40F8D1E4A2684">
|
||
<mods:title id="FAF6428609EA70B84A69477C14F42657">Three new species of Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae) from Amazonian Peru and a checklist of Cyclocephala species in Peru</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="C985D7946E26359F2C1506ED6F274C7C" type="personal">
|
||
<mods:role id="E3CA900769F300110E5BE584EF8239F9">
|
||
<mods:roleTerm id="960FAEDD43C9A96A1750BB59DEE8849D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B5C09A6080C9CA74A23F26310FA30C2A">Paz, Fernando</mods:namePart>
|
||
<mods:nameIdentifier id="F7A937BDD2DAE695B82D14D477510107" type="ORCID">0000-0002-6331-4416</mods:nameIdentifier>
|
||
<mods:affiliation id="EEE0AE57080020AD435E8004CA859191">Departamento de Entomología, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Apartado Postal 14 - 0434, Lima, Peru. fernandopaz 683 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6331 - 4416</mods:affiliation>
|
||
<mods:nameIdentifier id="05A077E8E2F97547E1482DB9315D0A48" type="email">fernandopaz683@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="E79C6BC2F581810A808C78B1206DB473" type="personal">
|
||
<mods:role id="837C24D88CBA0D9A4AE49B899DC1E5EC">
|
||
<mods:roleTerm id="FF9CBB61809DFA3EB51AEBCEBDDB843B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="BC384C2E3F779919F174C6CFF0EEB434">Ratcliffe, Brett C.</mods:namePart>
|
||
<mods:nameIdentifier id="C39F3110BED444B36B32AF22CCB28EAE" type="ORCID">0000-0002-0589-8791</mods:nameIdentifier>
|
||
<mods:affiliation id="D7181EEA135AC52A64043F135B9D7A12">Systematics Research Collections, University of Nebraska State Museum, University of Nebraska, Lincoln, Nebraska, 68588 - 0338, United States of America. bratcliffe 1 @ unl. edu; http: // orcid. org / 0000 - 0002 - 0589 - 8791</mods:affiliation>
|
||
<mods:nameIdentifier id="9D8C9745516D1750469321859E6037DB" type="email">bratcliffe1@unl.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="572D3C7D00B44B73C38A37E7D8DECC45" type="personal">
|
||
<mods:role id="DFA1A0EDDCC77CB858B27154781525B8">
|
||
<mods:roleTerm id="B6721AA5D599552A74179BFFDAA6D0BC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3BC4FE91427A2DF4FF3FA77709EE147A">Figueroa, Luis</mods:namePart>
|
||
<mods:affiliation id="4D95672202B79E62C0EF636B7DE58B1E">Departamento de Entomología, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Lima, Peru.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="F80153D8D730379CAE0DC3FF5A32A898">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F95E804298793E46F48619D62F24A990" type="host">
|
||
<mods:titleInfo id="711FCA590D4B6E6A456279562D8D138E">
|
||
<mods:title id="2EC3613C0D6D3950A55BC1E3E89E46AA">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="BC653EC24D4B7847D5A55300354DA047">
|
||
<mods:date id="6E929F60A6EA426538BF829A72E155A3">2022</mods:date>
|
||
<mods:detail id="797E4553F4F56ECBF17654F48CA87C47" type="pubDate">
|
||
<mods:number id="49CE551C9CFF4FF7DC79CBE950F9ADAA">2022-01-07</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="257F82ACB649602EEF22608A4E03527E" type="volume">
|
||
<mods:number id="67A517D852A1EA0C7853DF3E4113E980">5087</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="801215F1CF2C33B4BA88EBF8493E8CD7" type="issue">
|
||
<mods:number id="3E0F559C3B2256445888E9287AB09039">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="D3C9113ED2E4B6811851D96824015179" unit="page">
|
||
<mods:start id="6E12B79A45E706CC9219192113523C37">427</mods:start>
|
||
<mods:end id="32933D78FE54D5EF7B9D7A9F6D1490BB">440</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="F4A56679E968F1466202645332B6B559">journal article</mods:classification>
|
||
<mods:identifier id="F74FA6634FF24C4ECF070E366BB1B8CB" type="CLB-Dataset">2767</mods:identifier>
|
||
<mods:identifier id="A13B4D87266DB2AA0854E088FF8B3360" type="DOI">10.11646/zootaxa.5087.3.2</mods:identifier>
|
||
<mods:identifier id="1D8DAC1DDC96451CD213436FB3E5E409" type="GBIF-Dataset">43c1b1e7-1797-4ed7-817d-606114b807ce</mods:identifier>
|
||
<mods:identifier id="82F4B9AE1C93A513E037E41E6DAD3D7F" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="406B96E019F4BDB2EC039195C8ED971D" type="Zenodo-Dep">5826858</mods:identifier>
|
||
<mods:identifier id="3D6BF14A091A4C8AB5CC581856EDDC2C" type="ZooBank">6253FEED-A4FB-4832-AE51-91E4F51655C4</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="E61687C1FFA2FFD17BB4BCE9FB121B3F" ID-DOI="http://doi.org/10.5281/zenodo.5826868" ID-GBIF-Taxon="191928391" ID-Zenodo-Dep="5826868" LSID="urn:lsid:zoobank.org:act:CCED91AA-7B00-4830-B6E2-BF5E648294A7" httpUri="http://treatment.plazi.org/id/E61687C1FFA2FFD17BB4BCE9FB121B3F" lastPageId="4" lastPageNumber="431" pageId="3" pageNumber="430">
|
||
<subSubSection id="26A5655CFFA2FFD67BB4BCE9FD5719B6" box="[151,761,978,1005]" pageId="3" pageNumber="430" type="nomenclature">
|
||
<paragraph id="6E0036D7FFA2FFD67BB4BCE9FD5719B6" blockId="3.[151,761,978,1041]" box="[151,761,978,1005]" pageId="3" pageNumber="430">
|
||
<heading id="354881BBFFA2FFD67BB4BCE9FD5719B6" bold="true" box="[151,761,978,1005]" fontSize="11" level="1" pageId="3" pageNumber="430" reason="1">
|
||
<emphasis id="5CCBEAC5FFA2FFD67BB4BCE9FD5719B6" bold="true" box="[151,761,978,1005]" pageId="3" pageNumber="430">
|
||
<taxonomicName id="A9BF4D54FFA2FFD67BB4BCE9FDF219B6" authority="Paz & Ratcliffe" authorityName="Paz & Ratcliffe" authorityYear="2022" box="[151,604,978,1005]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="mateoi" status="sp. nov.">
|
||
<emphasis id="5CCBEAC5FFA2FFD67BB4BCE9FE3C19B7" bold="true" box="[151,402,978,1005]" italics="true" pageId="3" pageNumber="430">Cyclocephala mateoi</emphasis>
|
||
Paz & Ratcliffe
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="47F857BEFFA2FFD6794ABCE8FD5719B6" box="[617,761,978,1005]" pageId="3" pageNumber="430" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="26A5655CFFA2FFD67BB4BCCCFEE41E4B" box="[151,330,1015,1041]" pageId="3" pageNumber="430" type="description">
|
||
<paragraph id="6E0036D7FFA2FFD67BB4BCCCFEE41E4B" blockId="3.[151,761,978,1041]" box="[151,330,1015,1041]" pageId="3" pageNumber="430">
|
||
(
|
||
<figureCitation id="F6842A52FFA2FFD67BBCBCCCFEB71E4B" box="[159,281,1015,1041]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1893,1917]" captionTargetBox="[183,1413,388,1865]" captionTargetId="figure-118@4.[159,1427,381,1870]" captionTargetPageId="4" captionText="FIGURES 7–12. Cyclocephala mateoi. 7, Male holotype; 8, female allotype; 9–10, parameres of holotype; 11, protarsal claw of holotype; 12, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826864" httpUri="https://zenodo.org/record/5826864/files/figure.png" pageId="3" pageNumber="430">Figs. 7–12</figureCitation>
|
||
,
|
||
<figureCitation id="F6842A52FFA2FFD67A06BCCCFEEC1E4B" box="[293,322,1015,1041]" captionStart="FIGURE 20" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[198,1390,181,1919]" captionTargetId="figure-18@7.[198,1390,181,1920]" captionTargetPageId="7" captionText="FIGURE 20. Distribution map of the new species of Peruvian Cyclocephala." figureDoi="http://doi.org/10.5281/zenodo.5826870" httpUri="https://zenodo.org/record/5826870/files/figure.png" pageId="3" pageNumber="430">20</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="26A5655CFFA2FFD67BB4BB04FEEC1EFB" pageId="3" pageNumber="430" type="materials_examined">
|
||
<paragraph id="6E0036D7FFA2FFD67BB4BB04FEEC1EFB" blockId="3.[151,1437,1086,2013]" pageId="3" pageNumber="430">
|
||
<emphasis id="5CCBEAC5FFA2FFD67BB4BB04FEE01E02" bold="true" box="[151,334,1086,1113]" pageId="3" pageNumber="430">Type material:</emphasis>
|
||
<materialsCitation id="DED73C8AFFA2FFD67A78BB04FBEB1E27" ID-GBIF-Occurrence="3427663325" collectionCode="UNAS" country="Peru" elevation="693" latitude="-9.312417" location="Tingo Maria" longLatPrecision="1" longitude="-75.99525" municipality="Campus" pageId="3" pageNumber="430" specimenCount="693" typeStatus="holotype">
|
||
<typeStatus id="B1048875FFA2FFD67A78BB04FE6D1E03" box="[347,451,1087,1113]" pageId="3" pageNumber="430" type="holotype">Holotype</typeStatus>
|
||
male and
|
||
<typeStatus id="B1048875FFA2FFD67969BB04FD0A1E03" box="[586,676,1087,1113]" pageId="3" pageNumber="430" type="allotype">allotype</typeStatus>
|
||
female labeled: “
|
||
<collectingCountry id="16A87647FFA2FFD67858BB04FC661E03" box="[891,968,1087,1113]" name="Peru" pageId="3" pageNumber="430">PERU</collectingCountry>
|
||
, HU.
|
||
<location id="6B60600CFFA2FFD67F35BB04FB051E03" LSID="urn:lsid:plazi:treatment:E61687C1FFA2FFD17BB4BCE9FB121B3F:6B60600CFFA2FFD67F35BB04FB051E03" box="[1046,1195,1087,1113]" country="Peru" latitude="-9.312417" longLatPrecision="1" longitude="-75.99525" municipality="Campus" name="Tingo Maria" pageId="3" pageNumber="430">Tingo Maria</location>
|
||
, /
|
||
<collectingMunicipality id="8E64ACADFFA2FFD67FF1BB04FA801E03" box="[1234,1326,1087,1113]" pageId="3" pageNumber="430">Campus</collectingMunicipality>
|
||
<collectionCode id="08AEAE12FFA2FFD67E18BB04FA291E03" box="[1339,1415,1087,1113]" pageId="3" pageNumber="430">UNAS</collectionCode>
|
||
/
|
||
<geoCoordinate id="0B8B5010FFA2FFD67BB4BB58FEB21E27" box="[151,284,1123,1149]" degrees="9" direction="south" minutes="18" orientation="latitude" pageId="3" pageNumber="430" precision="1" seconds="44.7" value="-9.312417">9°18'44.7"S</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="0B8B5010FFA2FFD67A0EBB59FE651E27" box="[301,459,1122,1149]" degrees="75" direction="west" minutes="59" orientation="longitude" pageId="3" pageNumber="430" precision="1" seconds="42.9" value="-75.99525">75°59'42.9"W</geoCoordinate>
|
||
/
|
||
<specimenCount id="78B9FD5EFFA2FFD67AC1BB58FD9C1E27" box="[482,562,1123,1149]" pageId="3" pageNumber="430" type="generic">
|
||
<quantity id="A9479B32FFA2FFD67AC1BB58FD801E27" box="[482,558,1123,1149]" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="3" pageNumber="430" unit="m" value="693.0">
|
||
<elevation id="E592D1E4FFA2FFD67AC1BB58FD801E27" box="[482,558,1123,1149]" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="3" pageNumber="430" unit="m" value="693.0">693 m</elevation>
|
||
</quantity>
|
||
.
|
||
</specimenCount>
|
||
” and with our red
|
||
<typeStatus id="B1048875FFA2FFD67826BB58FCC91E27" box="[773,871,1123,1149]" pageId="3" pageNumber="430" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="B1048875FFA2FFD678BCBB58FC571E27" box="[927,1017,1123,1149]" pageId="3" pageNumber="430" type="allotype">allotype</typeStatus>
|
||
labels
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="DED73C8AFFA2FFD67F73BB58FE901EFB" ID-GBIF-Occurrence="3427663309" collectionCode="MUSM" country="Peru" pageId="3" pageNumber="430" specimenCount="2" typeStatus="holotype">
|
||
<typeStatus id="B1048875FFA2FFD67F73BB58FB161E27" box="[1104,1208,1123,1149]" pageId="3" pageNumber="430" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<typeStatus id="B1048875FFA2FFD67FD3BB58FAE41E27" box="[1264,1354,1123,1149]" pageId="3" pageNumber="430" type="allotype">allotype</typeStatus>
|
||
deposited in
|
||
<collectionCode id="08AEAE12FFA2FFD67BC6BBBCFE901EFB" box="[229,318,1159,1185]" pageId="3" pageNumber="430">MUSM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="26A5655CFFA2FFD67BE4BB91FA3F1CBB" pageId="3" pageNumber="430" type="description">
|
||
<paragraph id="6E0036D7FFA2FFD67BE4BB91FD701C0B" blockId="3.[151,1437,1086,2013]" pageId="3" pageNumber="430">
|
||
<emphasis id="5CCBEAC5FFA2FFD67BE4BB91FE7C1E9F" bold="true" box="[199,466,1194,1221]" pageId="3" pageNumber="430">
|
||
Description.
|
||
<typeStatus id="B1048875FFA2FFD67A7CBB90FE601E9F" box="[351,462,1195,1221]" pageId="3" pageNumber="430" type="holotype">Holotype</typeStatus>
|
||
,
|
||
</emphasis>
|
||
male. (
|
||
<figureCitation id="F6842A52FFA2FFD67906BB90FDC61E9F" box="[549,616,1195,1221]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1893,1917]" captionTargetBox="[183,1413,388,1865]" captionTargetId="figure-118@4.[159,1427,381,1870]" captionTargetPageId="4" captionText="FIGURES 7–12. Cyclocephala mateoi. 7, Male holotype; 8, female allotype; 9–10, parameres of holotype; 11, protarsal claw of holotype; 12, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826864" httpUri="https://zenodo.org/record/5826864/files/figure.png" pageId="3" pageNumber="430">Fig. 7</figureCitation>
|
||
). Length
|
||
<quantity id="A9479B32FFA2FFD679F1BB90FC961E9F" box="[722,824,1195,1221]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="3" pageNumber="430" unit="mm" value="14.4">14.4 mm</quantity>
|
||
; width
|
||
<quantity id="A9479B32FFA2FFD678AABB90FC4F1E9F" box="[905,993,1195,1221]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="3" pageNumber="430" unit="mm" value="6.6">6.6 mm</quantity>
|
||
. Color testaceous except for black frons and apices of tibial teeth.
|
||
<emphasis id="5CCBEAC5FFA2FFD67A9BBBF4FDAF1EB3" bold="true" box="[440,513,1231,1257]" pageId="3" pageNumber="430">Head:</emphasis>
|
||
Frons and clypeus with punctures moderate in size and density, glabrous. Clypeal apex subtruncate, slightly reflexed. Interocular width equals 3.0 transverse eye diameters. Antenna with 10 antennomeres, club subequal in length to antennomeres 2–7.
|
||
<emphasis id="5CCBEAC5FFA2FFD679C0BA2CFCCB1F6B" bold="true" box="[739,869,1303,1329]" pageId="3" pageNumber="430">Pronotum:</emphasis>
|
||
Surface with small, sparse punctures. Base lacking marginal bead.
|
||
<emphasis id="5CCBEAC5FFA2FFD67A60BA00FE371F0F" bold="true" box="[323,409,1339,1365]" pageId="3" pageNumber="430">Elytra:</emphasis>
|
||
Surface weakly punctate-striate, punctures slightly larger than those of pronotum, glabrous.
|
||
<emphasis id="5CCBEAC5FFA2FFD67BB4BA65FEBA1F22" bold="true" box="[151,276,1374,1400]" pageId="3" pageNumber="430">Pygidium:</emphasis>
|
||
Disc with minute, sparse punctures; lateral angles with moderately large, dense punctures and partially rugose. In lateral view, surface convex.
|
||
<emphasis id="5CCBEAC5FFA2FFD67976BAB8FD381FC7" bold="true" box="[597,662,1411,1437]" pageId="3" pageNumber="430">Legs:</emphasis>
|
||
Protibia tridentate, basal tooth small, distant from middle tooth. Protarsomere 5 enlarged, slightly bent, venter concave; medial claw large, bent, apex split (
|
||
<figureCitation id="F6842A52FFA2FFD67F4BBA9CFB1C1F9B" box="[1128,1202,1447,1473]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1893,1917]" captionTargetBox="[183,1413,388,1865]" captionTargetId="figure-118@4.[159,1427,381,1870]" captionTargetPageId="4" captionText="FIGURES 7–12. Cyclocephala mateoi. 7, Male holotype; 8, female allotype; 9–10, parameres of holotype; 11, protarsal claw of holotype; 12, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826864" httpUri="https://zenodo.org/record/5826864/files/figure.png" pageId="3" pageNumber="430">Fig 11</figureCitation>
|
||
). Metatarsus slightly longer than metatibia.
|
||
<emphasis id="5CCBEAC5FFA2FFD67ABABAF0FE5C1FBF" bold="true" box="[409,498,1483,1509]" pageId="3" pageNumber="430">Venter:</emphasis>
|
||
Prosternal process long, columnar, apex obliquely flattened into transversely oval disc with transverse, elevated “button” on anterior half.
|
||
<emphasis id="5CCBEAC5FFA2FFD67823BAD4FC241C53" bold="true" box="[768,906,1519,1545]" pageId="3" pageNumber="430">Parameres:</emphasis>
|
||
In caudal view, form elongate, base subtriangular, shaft subparallel, lateral margins of each paramere slightly flared before rounded apex (
|
||
<figureCitation id="F6842A52FFA2FFD67FB9B928FB711C77" box="[1178,1247,1555,1581]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1893,1917]" captionTargetBox="[183,1413,388,1865]" captionTargetId="figure-118@4.[159,1427,381,1870]" captionTargetPageId="4" captionText="FIGURES 7–12. Cyclocephala mateoi. 7, Male holotype; 8, female allotype; 9–10, parameres of holotype; 11, protarsal claw of holotype; 12, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826864" httpUri="https://zenodo.org/record/5826864/files/figure.png" pageId="3" pageNumber="430">Fig. 9</figureCitation>
|
||
). In lateral view, basal piece slightly longer than parameres (
|
||
<figureCitation id="F6842A52FFA2FFD6795EB90CFD7E1C0B" box="[637,720,1591,1617]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1893,1917]" captionTargetBox="[183,1413,388,1865]" captionTargetId="figure-118@4.[159,1427,381,1870]" captionTargetPageId="4" captionText="FIGURES 7–12. Cyclocephala mateoi. 7, Male holotype; 8, female allotype; 9–10, parameres of holotype; 11, protarsal claw of holotype; 12, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826864" httpUri="https://zenodo.org/record/5826864/files/figure.png" pageId="3" pageNumber="430">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6E0036D7FFA2FFD67BE4B961FA3F1CBB" blockId="3.[151,1437,1086,2013]" pageId="3" pageNumber="430">
|
||
<emphasis id="5CCBEAC5FFA2FFD67BE4B961FE9D1C2E" bold="true" box="[199,307,1626,1652]" pageId="3" pageNumber="430">
|
||
<typeStatus id="B1048875FFA2FFD67BE4B961FE801C2E" box="[199,302,1626,1652]" pageId="3" pageNumber="430" type="allotype">Allotype</typeStatus>
|
||
,
|
||
</emphasis>
|
||
female (
|
||
<figureCitation id="F6842A52FFA2FFD67AB5B960FE751C2F" box="[406,475,1627,1653]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1893,1917]" captionTargetBox="[183,1413,388,1865]" captionTargetId="figure-118@4.[159,1427,381,1870]" captionTargetPageId="4" captionText="FIGURES 7–12. Cyclocephala mateoi. 7, Male holotype; 8, female allotype; 9–10, parameres of holotype; 11, protarsal claw of holotype; 12, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826864" httpUri="https://zenodo.org/record/5826864/files/figure.png" pageId="3" pageNumber="430">Fig. 8</figureCitation>
|
||
). Length
|
||
<quantity id="A9479B32FFA2FFD6796BB960FD011C2F" box="[584,687,1627,1653]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="3" pageNumber="430" unit="mm" value="16.3">16.3 mm</quantity>
|
||
; width across humeri
|
||
<quantity id="A9479B32FFA2FFD67885B960FC511C2F" box="[934,1023,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="3" pageNumber="430" unit="mm" value="7.4">7.4 mm</quantity>
|
||
. As
|
||
<typeStatus id="B1048875FFA2FFD67F0CB960FB3F1C2F" box="[1071,1169,1627,1653]" pageId="3" pageNumber="430" type="holotype">holotype</typeStatus>
|
||
except in the following respects:
|
||
<emphasis id="5CCBEAC5FFA2FFD67A21B944FEF61CC3" bold="true" box="[258,344,1663,1689]" pageId="3" pageNumber="430">Elytra:</emphasis>
|
||
Epipleuron (ventral view) expanded at level of abdominal sternites 1–2 and then abruptly constricted with a subacute tooth at level of abdominal sternite 3 (
|
||
<figureCitation id="F6842A52FFA2FFD6787DB998FC1C1CE7" box="[862,946,1699,1725]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1893,1917]" captionTargetBox="[183,1413,388,1865]" captionTargetId="figure-118@4.[159,1427,381,1870]" captionTargetPageId="4" captionText="FIGURES 7–12. Cyclocephala mateoi. 7, Male holotype; 8, female allotype; 9–10, parameres of holotype; 11, protarsal claw of holotype; 12, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826864" httpUri="https://zenodo.org/record/5826864/files/figure.png" pageId="3" pageNumber="430">Fig. 12</figureCitation>
|
||
). In dorsal view, lateral margin slightly expanded above sternite 3.
|
||
<emphasis id="5CCBEAC5FFA2FFD67A8DB9FDFD851CBA" bold="true" box="[430,555,1734,1760]" pageId="3" pageNumber="430">Pygidium:</emphasis>
|
||
In lateral view, surface weakly convex.
|
||
<emphasis id="5CCBEAC5FFA2FFD678CDB9FCFB811CBB" bold="true" box="[1006,1071,1735,1761]" pageId="3" pageNumber="430">Legs:</emphasis>
|
||
Protarsus simple, not enlarged.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="26A5655CFFA2FFD67BE4B9D0FB0B1D5F" box="[199,1189,1771,1797]" pageId="3" pageNumber="430" type="etymology">
|
||
<paragraph id="6E0036D7FFA2FFD67BE4B9D0FB0B1D5F" blockId="3.[151,1437,1086,2013]" box="[199,1189,1771,1797]" pageId="3" pageNumber="430">
|
||
<emphasis id="5CCBEAC5FFA2FFD67BE4B9D0FEE01D5F" bold="true" box="[199,334,1771,1797]" pageId="3" pageNumber="430">Etymology.</emphasis>
|
||
This new species is dedicated to Fernando Paz’s nephew, Mateo Causo Paz.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="26A5655CFFA2FFD67BE4B835FD931D2B" pageId="3" pageNumber="430" type="distribution">
|
||
<paragraph id="6E0036D7FFA2FFD67BE4B835FDA11D17" blockId="3.[151,1437,1086,2013]" pageId="3" pageNumber="430">
|
||
<emphasis id="5CCBEAC5FFA2FFD67BE4B835FECF1D72" bold="true" box="[199,353,1806,1832]" pageId="3" pageNumber="430">Distribution.</emphasis>
|
||
<specimenCount id="78B9FD5EFFA2FFD67A4AB834FE581D73" box="[361,502,1807,1833]" pageId="3" pageNumber="430" type="generic">2 specimens</specimenCount>
|
||
examined (
|
||
<figureCitation id="F6842A52FFA2FFD6795FB834FD7D1D73" box="[636,723,1807,1833]" captionStart="FIGURE 20" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[198,1390,181,1919]" captionTargetId="figure-18@7.[198,1390,181,1920]" captionTargetPageId="7" captionText="FIGURE 20. Distribution map of the new species of Peruvian Cyclocephala." figureDoi="http://doi.org/10.5281/zenodo.5826870" httpUri="https://zenodo.org/record/5826870/files/figure.png" pageId="3" pageNumber="430">Fig. 20</figureCitation>
|
||
).
|
||
<collectingCountry id="16A87647FFA2FFD679CBB834FCB11D7C" box="[744,799,1807,1833]" name="Peru" pageId="3" pageNumber="430">PERU</collectingCountry>
|
||
(2):
|
||
<collectingRegion id="AC7BF835FFA2FFD67879B82BFC481D71" box="[858,998,1808,1835]" country="Peru" name="Huanuco" pageId="3" pageNumber="430">HUÁNUCO</collectingRegion>
|
||
(2): Campus of Universidad Nacional Agraria de la Selva, Tingo María.
|
||
</paragraph>
|
||
<paragraph id="6E0036D7FFA2FFD67BE4B86CFD931D2B" blockId="3.[151,1437,1086,2013]" box="[199,573,1878,1905]" pageId="3" pageNumber="430">
|
||
<emphasis id="5CCBEAC5FFA2FFD67BE4B86CFE791D2A" bold="true" box="[199,471,1878,1905]" pageId="3" pageNumber="430">Temporal distribution.</emphasis>
|
||
No data.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="26A5655CFFA2FFD17BE4B841FB121B3F" lastPageId="4" lastPageNumber="431" pageId="3" pageNumber="430" type="diagnosis">
|
||
<paragraph id="6E0036D7FFA2FFD17BE4B841FF551B1B" blockId="3.[151,1437,1086,2013]" lastBlockId="4.[151,1437,151,357]" lastPageId="4" lastPageNumber="431" pageId="3" pageNumber="430">
|
||
<emphasis id="5CCBEAC5FFA2FFD67BE4B841FEEA1DCE" bold="true" box="[199,324,1914,1940]" pageId="3" pageNumber="430">Diagnosis:</emphasis>
|
||
<taxonomicName id="A9BF4D54FFA2FFD67A69B840FD991DCE" box="[330,567,1915,1941]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="mateoi">
|
||
<emphasis id="5CCBEAC5FFA2FFD67A69B840FD991DCE" box="[330,567,1915,1941]" italics="true" pageId="3" pageNumber="430">Cyclocephala mateoi</emphasis>
|
||
</taxonomicName>
|
||
is similar in
|
||
<emphasis id="5CCBEAC5FFA2FFD679E4B840FCBC1DCF" box="[711,786,1915,1941]" italics="true" pageId="3" pageNumber="430">gestalt</emphasis>
|
||
to
|
||
<taxonomicName id="A9BF4D54FFA2FFD67817B840FB291DCF" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[820,1159,1915,1941]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="verticalis">
|
||
<emphasis id="5CCBEAC5FFA2FFD67817B840FC121DCF" box="[820,956,1915,1941]" italics="true" pageId="3" pageNumber="430">C. verticalis</emphasis>
|
||
Burmeister, 1847
|
||
</taxonomicName>
|
||
. However,
|
||
<taxonomicName id="A9BF4D54FFA2FFD67E27B840FADE1DCE" box="[1284,1392,1915,1941]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="mateoi">
|
||
<emphasis id="5CCBEAC5FFA2FFD67E27B840FADE1DCE" box="[1284,1392,1915,1941]" italics="true" pageId="3" pageNumber="430">C. mateoi</emphasis>
|
||
</taxonomicName>
|
||
has a slightly more subtruncate clypeal apex while the clypeal apex of
|
||
<taxonomicName id="A9BF4D54FFA2FFD678A9B8A4FBBD1DE3" authorityName="Burmeister" authorityYear="1847" box="[906,1043,1951,1977]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="verticalis">
|
||
<emphasis id="5CCBEAC5FFA2FFD678A9B8A4FBBD1DE3" box="[906,1043,1951,1977]" italics="true" pageId="3" pageNumber="430">C. verticalis</emphasis>
|
||
</taxonomicName>
|
||
is more parabolic; the protibia has a small, basal tooth that is distant from the others in
|
||
<taxonomicName id="A9BF4D54FFA2FFD679C7B8F8FCFF1D86" box="[740,849,1987,2013]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="mateoi">
|
||
<emphasis id="5CCBEAC5FFA2FFD679C7B8F8FCFF1D86" box="[740,849,1987,2013]" italics="true" pageId="3" pageNumber="430">C. mateoi</emphasis>
|
||
</taxonomicName>
|
||
, but the protibal teeth in
|
||
<taxonomicName id="A9BF4D54FFA2FFD67F4EB8F8FB581D87" authorityName="Burmeister" authorityYear="1847" box="[1133,1270,1987,2013]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="verticalis">
|
||
<emphasis id="5CCBEAC5FFA2FFD67F4EB8F8FB581D87" box="[1133,1270,1987,2013]" italics="true" pageId="3" pageNumber="430">C. verticalis</emphasis>
|
||
</taxonomicName>
|
||
are all equally spaced; and the parameres are subtriangular at the base, subparallel, and flared before the apices (
|
||
<figureCitation id="F6842A52FFA5FFD17FF2BFACFABD1AEB" box="[1233,1299,151,177]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1893,1917]" captionTargetBox="[183,1413,388,1865]" captionTargetId="figure-118@4.[159,1427,381,1870]" captionTargetPageId="4" captionText="FIGURES 7–12. Cyclocephala mateoi. 7, Male holotype; 8, female allotype; 9–10, parameres of holotype; 11, protarsal claw of holotype; 12, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826864" httpUri="https://zenodo.org/record/5826864/files/figure.png" pageId="4" pageNumber="431">Fig. 9</figureCitation>
|
||
) while in
|
||
<taxonomicName id="A9BF4D54FFA5FFD17EA0BFACFF511A8F" authorityName="Burmeister" authorityYear="1847" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="verticalis">
|
||
<emphasis id="5CCBEAC5FFA5FFD17EA0BFACFF511A8F" italics="true" pageId="4" pageNumber="431">C. verticalis</emphasis>
|
||
</taxonomicName>
|
||
the parameres are elongate-oval with a tuft of setae on the inner margin of each apex. The female epipleuron of
|
||
<taxonomicName id="A9BF4D54FFA5FFD17BC0BFE4FEFF1AA2" box="[227,337,223,249]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="mateoi">
|
||
<emphasis id="5CCBEAC5FFA5FFD17BC0BFE4FEFF1AA2" box="[227,337,223,249]" italics="true" pageId="4" pageNumber="431">C. mateoi</emphasis>
|
||
</taxonomicName>
|
||
is expanded at the level of abdominal sternites 1–2 and then abruptly constricted with a subacute tooth at the level of abdominal sternite 3 (
|
||
<figureCitation id="F6842A52FFA5FFD1795EBE38FD7F1B47" box="[637,721,259,285]" captionStart="FIGURES 7–12" captionStartId="4.[151,264,1893,1917]" captionTargetBox="[183,1413,388,1865]" captionTargetId="figure-118@4.[159,1427,381,1870]" captionTargetPageId="4" captionText="FIGURES 7–12. Cyclocephala mateoi. 7, Male holotype; 8, female allotype; 9–10, parameres of holotype; 11, protarsal claw of holotype; 12, epipleuron (ventral view) of allotype. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5826864" httpUri="https://zenodo.org/record/5826864/files/figure.png" pageId="4" pageNumber="431">Fig. 12</figureCitation>
|
||
), while the epipleuron of
|
||
<taxonomicName id="A9BF4D54FFA5FFD178DEBE38FB261B47" authorityName="Burmeister" authorityYear="1847" box="[1021,1160,259,285]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="verticalis">
|
||
<emphasis id="5CCBEAC5FFA5FFD178DEBE38FB261B47" box="[1021,1160,259,285]" italics="true" pageId="4" pageNumber="431">C. verticalis</emphasis>
|
||
</taxonomicName>
|
||
is simple and gradually tapering.
|
||
</paragraph>
|
||
<paragraph id="6E0036D7FFA5FFD17BE4BE70FB121B3F" blockId="4.[151,1437,151,357]" box="[199,1212,330,357]" pageId="4" pageNumber="431">
|
||
<emphasis id="5CCBEAC5FFA5FFD17BE4BE70FE2B1B3E" bold="true" box="[199,389,330,357]" pageId="4" pageNumber="431">Natural history.</emphasis>
|
||
<taxonomicName id="A9BF4D54FFA5FFD17AAFBE70FDD71B3E" box="[396,633,331,357]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="mateoi">
|
||
<emphasis id="5CCBEAC5FFA5FFD17AAFBE70FDD71B3E" box="[396,633,331,357]" italics="true" pageId="4" pageNumber="431">Cyclocephala mateoi</emphasis>
|
||
</taxonomicName>
|
||
specimens were collected at an elevation of
|
||
<specimenCount id="78B9FD5EFFA5FFD17F4DBE70FB121B3F" box="[1134,1212,331,357]" pageId="4" pageNumber="431" type="generic">
|
||
<quantity id="A9479B32FFA5FFD17F4DBE70FB161B3F" box="[1134,1208,331,357]" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="4" pageNumber="431" unit="m" value="693.0">693 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |