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<document id="AAC0E019FF78BF531C4E8EE858145A93" ID-DOI="10.25226/bboc.v142i1.2022.a3" ID-ISSN="2513-9894" ID-ZooBank="4C9216EC-E822-4CC7-A163-6E96CFB3078F" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1718072142794" checkinUser="felipe" docAuthor="Suárez, William" docDate="2022" docId="E575C653FF940830FE26A7AB5693FE23" docLanguage="en" docName="BullBritOrnitholClub.142.1.10-74.pdf" docOrigin="Bulletin of the British Ornithologists Club 142 (1)" docStyle="DocumentStyle:EFA7F293C51FD4806C5290561C16F223.2:BullBritOrnitholClub.2017-.journal_article" docStyleId="EFA7F293C51FD4806C5290561C16F223" docStyleName="BullBritOrnitholClub.2017-.journal_article" docStyleVersion="2" docTitle="Tyto pollens Wetmore 1937" docType="treatment" docVersion="4" lastPageNumber="42" masterDocId="194CBE2BFFB40811FFE4A53D575AFFBA" masterDocTitle="Catalogue of Cuban fossil and subfossil birds" masterLastPageNumber="248" masterPageNumber="247" pageNumber="41" updateTime="1726258990606" updateUser="karina" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="8CD18AAD571ADC48CB72C57CE7A15F16">Catalogue of Cuban fossil and subfossil birds</mods:title>
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<subSubSection id="25C624CEFF940831FE26A7AB540FFD0A" box="[450,853,662,689]" pageId="32" pageNumber="41" type="nomenclature">
<paragraph id="6D637745FF940831FE26A7AB540FFD0A" blockId="32.[144,1036,662,826]" box="[450,853,662,689]" pageId="32" pageNumber="41">
<heading id="362BC029FF940831FE26A7AB540FFD0A" box="[450,853,662,689]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="41" reason="8">
<emphasis id="5FA8AB57FF940831FE26A7AB540FFD0A" bold="true" box="[450,853,662,689]" pageId="32" pageNumber="41">
22. †
<taxonomicName id="AADC0CC6FF940831FE1FA7AA540FFD0A" authority="Wetmore, 1937" authorityName="Wetmore" authorityYear="1937" box="[507,853,662,689]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="32" pageNumber="41" phylum="Chordata" rank="species" species="pollens">
<emphasis id="5FA8AB57FF940831FE1FA7AA55C2FD0B" bold="true" box="[507,664,662,689]" italics="true" pageId="32" pageNumber="41">Tyto pollens</emphasis>
<bibRefCitation id="094D0AB4FF940831FD44A7AA540FFD0A" author="Wetmore, A." box="[672,853,662,689]" pageId="32" pageNumber="41" pagination="427 - 441" refId="ref57784" refString="Wetmore, A. 1937. Bird remains from cave deposits on Great Exuma Island in the Bahamas. Bull. Mus. Comp. Zool. 80: 427 - 441." type="journal article" year="1937">Wetmore, 1937</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="25C624CEFF940831FEF4A78B5356FD6B" box="[272,1036,694,721]" pageId="32" pageNumber="41" type="description">
<paragraph id="6D637745FF940831FEF4A78B5356FD6B" blockId="32.[144,1036,662,826]" box="[272,1036,694,721]" pageId="32" pageNumber="41">
Bahamian Giant Barn Owl (Lechuza Gigante de las
<collectingCountry id="15CB37D5FF940831FC76A78B5359FD6B" box="[914,1027,694,721]" name="Bahamas" pageId="32" pageNumber="41">Bahamas</collectingCountry>
)
</paragraph>
</subSubSection>
<subSubSection id="25C624CEFF940831FF74A7E05506FB01" pageId="32" pageNumber="41" type="reference_group">
<paragraph id="6D637745FF940831FF74A7E055E9FD55" blockId="32.[144,1036,662,826]" box="[144,691,732,752]" pageId="32" pageNumber="41">
<taxonomicName id="AADC0CC6FF940831FF74A7E056D5FD55" authority="Wetmore, 1937" authorityName="Wetmore" authorityYear="1937" box="[144,399,732,752]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="32" pageNumber="41" phylum="Chordata" rank="species" species="pollens">
<emphasis id="5FA8AB57FF940831FF74A7E057A0FD55" box="[144,250,732,751]" italics="true" pageId="32" pageNumber="41">Tyto pollens</emphasis>
<bibRefCitation id="094D0AB4FF940831FEE4A7E056D5FD55" author="Wetmore, A." box="[256,399,732,752]" pageId="32" pageNumber="41" pagination="427 - 441" refId="ref57784" refString="Wetmore, A. 1937. Bird remains from cave deposits on Great Exuma Island in the Bahamas. Bull. Mus. Comp. Zool. 80: 427 - 441." type="journal article" year="1937">Wetmore, 1937</bibRefCitation>
</taxonomicName>
,
<emphasis id="5FA8AB57FF940831FE73A7E1553AFD55" box="[407,608,732,752]" italics="true" pageId="32" pageNumber="41">
Bull.
<collectingRegion id="AF18B9A7FF940831FE23A7E056A8FD55" box="[455,498,733,751]" country="Turkey" name="Mus" pageId="32" pageNumber="41">Mus</collectingRegion>
. Comp. Zoöl
</emphasis>
. 80: 436.
</paragraph>
<paragraph id="6D637745FF940831FF24A7CB5553FCB2" blockId="32.[144,1036,662,826]" box="[192,521,757,776]" pageId="32" pageNumber="41">
<treatmentCitationGroup id="4DCC506BFF940831FF24A7CB5553FCB2" box="[192,521,757,776]" pageId="32" pageNumber="41">
<taxonomicName id="AADC0CC6FF940831FF24A7CB567DFCB2" authorityName="', Arredondo" authorityYear="1972" box="[192,295,758,776]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="32" pageNumber="41" phylum="Chordata" rank="species" species="riveroi">
<emphasis id="5FA8AB57FF940831FF24A7CB567DFCB2" box="[192,295,758,776]" italics="true" pageId="32" pageNumber="41">Tyto riveroi</emphasis>
</taxonomicName>
:
<treatmentCitation id="EC7D5154FF940831FED5A7C8555EFCB2" author="Arredondo, O." box="[305,516,757,776]" page="131" pageId="32" pageNumber="41" year="1972">
<bibRefCitation id="094D0AB4FF940831FED5A7C8555EFCB2" author="Arredondo, O." box="[305,516,757,776]" pageId="32" pageNumber="41" pagination="129 - 140" refId="ref47651" refString="Arredondo, O. 1972 b. Especie nueva de lechuza gigante (Strigiformes: Tytonidae) del Pleistoceno cubano. Bol. Soc. Venez. Cienc. Nat. 30 (124 - 125): 129 - 140." type="journal article" year="1972">Arredondo 1972b: 131</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6D637745FF940831FF24A63255BBFC9B" blockId="32.[144,1036,662,826]" box="[192,737,782,801]" pageId="32" pageNumber="41">
<emphasis id="5FA8AB57FF940831FF24A6325797FC9B" box="[192,205,783,801]" italics="true" pageId="32" pageNumber="41">T</emphasis>
[
<emphasis id="5FA8AB57FF940831FF30A63257B5FC9B" box="[212,239,783,801]" italics="true" pageId="32" pageNumber="41">yto</emphasis>
].
<emphasis id="5FA8AB57FF940831FEE5A633561DFC9A" box="[257,327,782,800]" italics="true" pageId="32" pageNumber="41">Triveroi</emphasis>
: Arredondo 1982: 36 (typographical error).
</paragraph>
<paragraph id="6D637745FF940831FF24A61A5576FC80" blockId="32.[144,1036,662,826]" box="[192,556,807,826]" pageId="32" pageNumber="41">
<treatmentCitationGroup id="4DCC506BFF940831FF24A61A5576FC80" box="[192,556,807,826]" pageId="32" pageNumber="41">
<taxonomicName id="AADC0CC6FF940831FF24A61A5670FC80" box="[192,298,807,826]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="32" pageNumber="41" phylum="Chordata" rank="species" species="pollens">
<emphasis id="5FA8AB57FF940831FF24A61A5670FC80" box="[192,298,807,826]" italics="true" pageId="32" pageNumber="41">Tyto pollens</emphasis>
</taxonomicName>
:
<treatmentCitation id="EC7D5154FF940831FED1A61A5573FC80" ID-DOI="http://doi.org/10.5281/zenodo.5625592" ID-Zenodo-Dep="5625592" author="Suarez, W. &amp; Olson, S. L." box="[309,553,807,826]" httpUri="http://treatment.plazi.org/id/03A4E42AFFE1143E26F96562F05F83D8" page="539" pageId="32" pageNumber="41" year="2015">
<bibRefCitation id="094D0AB4FF940831FED1A61A5573FC80" author="Suarez, W. &amp; Olson, S. L." box="[309,553,807,826]" pageId="32" pageNumber="41" pagination="533 - 553" refId="ref57070" refString="Suarez, W. &amp; Olson, S. L. 2015. Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa 4020: 533 - 553." type="journal article" year="2015">Suárez &amp; Olson 2015: 539</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="6D637745FF940831FF24A65E5506FB01" blockId="32.[144,1164,867,1691]" pageId="32" pageNumber="41">
<emphasis id="5FA8AB57FF940831FF24A65E564DFCC1" bold="true" box="[192,279,867,891]" italics="true" pageId="32" pageNumber="41">History</emphasis>
.—
<emphasis id="5FA8AB57FF940831FED3A65E5630FCC1" box="[311,362,867,891]" italics="true" pageId="32" pageNumber="41">1937</emphasis>
: Vivienne Knowles collects the
<typeStatus id="B267C9E7FF940831FD03A6595443FCC6" box="[743,793,868,892]" pageId="32" pageNumber="41">type</typeStatus>
series in
<collectingCountry id="15CB37D5FF940831FC76A65E537CFCC1" box="[914,1062,867,891]" name="Bahamas" pageId="32" pageNumber="41">the Bahamas</collectingCountry>
and the original description (October) of
<taxonomicName id="AADC0CC6FF940831FDF9A6B955D2FC21" authority="Wetmore, 1937" authorityName="Wetmore" authorityYear="1937" box="[541,648,899,923]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="32" pageNumber="41" phylum="Chordata" rank="species" species="pollens">
<emphasis id="5FA8AB57FF940831FDF9A6B955D2FC21" box="[541,648,899,923]" italics="true" pageId="32" pageNumber="41">T. pollens</emphasis>
</taxonomicName>
is published (
<bibRefCitation id="094D0AB4FF940831FCDDA6BE54BFFC21" author="Wetmore, A." box="[825,997,899,923]" pageId="32" pageNumber="41" pagination="427 - 441" refId="ref57784" refString="Wetmore, A. 1937. Bird remains from cave deposits on Great Exuma Island in the Bahamas. Bull. Mus. Comp. Zool. 80: 427 - 441." type="journal article" year="1937">Wetmore 1937</bibRefCitation>
).
<emphasis id="5FA8AB57FF940831FC19A6BE53DCFC21" box="[1021,1158,899,923]" italics="true" pageId="32" pageNumber="41">
<date id="19625185FF940831FC19A6BE53DCFC21" box="[1021,1158,899,923]" pageId="32" pageNumber="41" value="1959-06-03">3 June 1959</date>
</emphasis>
: additional specimens from Banana Hole, New Providence,
<collectingCountry id="15CB37D5FF940831FC8BA69E548DFC01" box="[879,983,931,955]" name="Bahamas" pageId="32" pageNumber="41">Bahamas</collectingCountry>
, are described and illustrated by
<bibRefCitation id="094D0AB4FF940831FE96A6FE55CFFC61" author="Brodkorb, P." box="[370,661,963,987]" pageId="32" pageNumber="41" pagination="349 - 371" refId="ref48589" refString="Brodkorb, P. 1959. Pleistocene birds from New Providence Island, Bahamas. Bull. Fla. State Mus. 4: 349 - 371." type="journal article" year="1959">Brodkorb (1959: 357358</bibRefCitation>
, pl. 1, figs. 5 = tibiotarsus [anterior], 6 = tarsometatarsus [anterior]).
<emphasis id="5FA8AB57FF940831FE2DA6DE5563FC41" box="[457,569,995,1019]" italics="true" pageId="32" pageNumber="41">
<date id="19625185FF940831FE2DA6DE5563FC41" box="[457,569,995,1019]" pageId="32" pageNumber="41" value="1970-04">April 1970</date>
</emphasis>
: Manuel Rivero de la Calle presents to O. Arredondo the first known material from
<collectingCountry id="15CB37D5FF940831FDE7A13E551BFBA1" box="[515,577,1027,1051]" name="Cuba" pageId="32" pageNumber="41">Cuba</collectingCountry>
, collected in Galería de los Megalocnus, Cueva de Bellamar, western
<collectingCountry id="15CB37D5FF940831FE76A11E5694FB81" box="[402,462,1059,1083]" name="Cuba" pageId="32" pageNumber="41">Cuba</collectingCountry>
(
<bibRefCitation id="094D0AB4FF940831FE06A11E55B2FB81" author="Arredondo, O." box="[482,744,1059,1083]" pageId="32" pageNumber="41" pagination="129 - 140" refId="ref47651" refString="Arredondo, O. 1972 b. Especie nueva de lechuza gigante (Strigiformes: Tytonidae) del Pleistoceno cubano. Bol. Soc. Venez. Cienc. Nat. 30 (124 - 125): 129 - 140." type="journal article" year="1972">Arredondo 1972b: 131</bibRefCitation>
).
<emphasis id="5FA8AB57FF940831FCE4A11E5469FB81" box="[768,819,1059,1083]" italics="true" pageId="32" pageNumber="41">1972</emphasis>
: original description of
<taxonomicName id="AADC0CC6FF940831FBB8A1195789FBE1" authorityName="', Arredondo" authorityYear="1972" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="32" pageNumber="41" phylum="Chordata" rank="species" species="riveroi">
<emphasis id="5FA8AB57FF940831FBB8A1195789FBE1" italics="true" pageId="32" pageNumber="41">Tyto riveroi</emphasis>
</taxonomicName>
published (
<bibRefCitation id="094D0AB4FF940831FE87A17E5575FBE1" author="Arredondo, O." box="[355,559,1091,1115]" pageId="32" pageNumber="41" pagination="129 - 140" refId="ref47651" refString="Arredondo, O. 1972 b. Especie nueva de lechuza gigante (Strigiformes: Tytonidae) del Pleistoceno cubano. Bol. Soc. Venez. Cienc. Nat. 30 (124 - 125): 129 - 140." type="journal article" year="1972">Arredondo 1972b</bibRefCitation>
).
<emphasis id="5FA8AB57FF940831FDA6A17E5450FBE1" box="[578,778,1091,1115]" italics="true" pageId="32" pageNumber="41">
<date id="19625185FF940831FDA6A17E5450FBE1" box="[578,778,1091,1115]" pageId="32" pageNumber="41" value="2015-09-23">23 September 2015</date>
</emphasis>
: first direct comparisons between Bahamian and Cuban giant barn owls (including previously unrecorded material) reveal
<taxonomicName id="AADC0CC6FF940831FF73A1B9564AFB21" authorityName="', Arredondo" authorityYear="1972" box="[151,272,1156,1179]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="32" pageNumber="41" phylum="Chordata" rank="species" species="riveroi">
<emphasis id="5FA8AB57FF940831FF73A1B9564AFB21" box="[151,272,1156,1179]" italics="true" pageId="32" pageNumber="41">Tyto riveroi</emphasis>
</taxonomicName>
to be a junior subjective synonym of
<taxonomicName id="AADC0CC6FF940831FD51A1B95443FB21" authority="Wetmore, 1937" authorityName="Wetmore" authorityYear="1937" box="[693,793,1155,1179]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="32" pageNumber="41" phylum="Chordata" rank="species" species="pollens">
<emphasis id="5FA8AB57FF940831FD51A1B9559EFB21" box="[693,708,1156,1179]" italics="true" pageId="32" pageNumber="41">T</emphasis>
.
<emphasis id="5FA8AB57FF940831FD35A1BE5443FB21" box="[721,793,1155,1179]" italics="true" pageId="32" pageNumber="41">pollens</emphasis>
</taxonomicName>
, and expand the ancient range of the latter to
<collectingCountry id="15CB37D5FF940831FEFCA19E560EFB01" box="[280,340,1187,1211]" name="Cuba" pageId="32" pageNumber="41">Cuba</collectingCountry>
(
<bibRefCitation id="094D0AB4FF940831FE86A19E550BFB01" author="Suarez, W. &amp; Olson, S. L." box="[354,593,1187,1211]" pageId="32" pageNumber="41" pagination="533 - 553" refId="ref57070" refString="Suarez, W. &amp; Olson, S. L. 2015. Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa 4020: 533 - 553." type="journal article" year="2015">Suárez &amp; Olson 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="25C624CEFF940831FF24A1FE54DBF981" pageId="32" pageNumber="41" type="materials_examined">
<paragraph id="6D637745FF940831FF24A1FE554DFAA1" blockId="32.[144,1164,867,1691]" pageId="32" pageNumber="41">
<emphasis id="5FA8AB57FF940831FF24A1FE5670FB61" bold="true" box="[192,298,1219,1243]" italics="true" pageId="32" pageNumber="41">
<typeStatus id="B267C9E7FF940831FF24A1FE5670FB61" box="[192,298,1219,1243]" pageId="32" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
.—Left femur, MCZ 2262 (
<bibRefCitation id="094D0AB4FF940831FD86A1FE542FFB61" author="Wetmore, A." box="[610,885,1219,1243]" pageId="32" pageNumber="41" pagination="427 - 441" refId="ref57784" refString="Wetmore, A. 1937. Bird remains from cave deposits on Great Exuma Island in the Bahamas. Bull. Mus. Comp. Zool. 80: 427 - 441." type="journal article" year="1937">Wetmore 1937: 436437</bibRefCitation>
, figs. 10* [anterior], 11* [posterior], 12* [lateral], 13* [distal], 14* [medial]). Collected during early 1937 by Vivienne Knowles (
<bibRefCitation id="094D0AB4FF940831FEE4A03E568CFAA1" author="Wetmore, A." box="[256,470,1283,1307]" pageId="32" pageNumber="41" pagination="427 - 441" refId="ref57784" refString="Wetmore, A. 1937. Bird remains from cave deposits on Great Exuma Island in the Bahamas. Bull. Mus. Comp. Zool. 80: 427 - 441." type="journal article" year="1937">Wetmore 1937: 427</bibRefCitation>
, 437).
</paragraph>
<paragraph id="6D637745FF940831FF24A01E54D2FAE1" blockId="32.[144,1164,867,1691]" pageId="32" pageNumber="41">
<emphasis id="5FA8AB57FF940831FF24A01E5607FA81" bold="true" box="[192,349,1315,1339]" italics="true" pageId="32" pageNumber="41">Type locality</emphasis>
.—Cave deposit on Great
<collectingRegion id="AF18B9A7FF940831FD7BA01E55B4FA80" box="[671,750,1315,1338]" country="Bahamas" name="Exuma" pageId="32" pageNumber="41">Exuma</collectingRegion>
[= Little
<collectingRegion id="AF18B9A7FF940831FC81A01E54EEFA80" box="[869,948,1315,1338]" country="Bahamas" name="Exuma" pageId="32" pageNumber="41">Exuma</collectingRegion>
<emphasis id="5FA8AB57FF940831FC26A01E54BCFA81" box="[962,998,1315,1339]" italics="true" pageId="32" pageNumber="41">fide</emphasis>
<bibRefCitation id="094D0AB4FF940831FC17A01E5327FA81" author="Hecht, M. K." box="[1011,1149,1315,1339]" pageId="32" pageNumber="41" pagination="133 - 135" refId="ref50289" refString="Hecht, M. K. 1955. The comparison of recent and fossil amphibian, reptilian, and mammalian faunas in the Bahamas. Yb. Amer. Philos. Soc. 1954: 133 - 135." type="journal article" year="1955">Hecht 1955</bibRefCitation>
], Bahama Islands (
<bibRefCitation id="094D0AB4FF940831FEB4A07E557CFAE1" author="Wetmore, A." box="[336,550,1347,1371]" pageId="32" pageNumber="41" pagination="427 - 441" refId="ref57784" refString="Wetmore, A. 1937. Bird remains from cave deposits on Great Exuma Island in the Bahamas. Bull. Mus. Comp. Zool. 80: 427 - 441." type="journal article" year="1937">Wetmore 1937: 437</bibRefCitation>
; see
<bibRefCitation id="094D0AB4FF940831FDBFA07E5427FAE1" author="Suarez, W. &amp; Olson, S. L." box="[603,893,1347,1371]" pageId="32" pageNumber="41" pagination="533 - 553" refId="ref57070" refString="Suarez, W. &amp; Olson, S. L. 2015. Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa 4020: 533 - 553." type="journal article" year="2015">Suárez &amp; Olson 2015: 540</bibRefCitation>
).
</paragraph>
<paragraph id="6D637745FF940831FF24A05E54DBF981" blockId="32.[144,1164,867,1691]" pageId="32" pageNumber="41">
<emphasis id="5FA8AB57FF940831FF24A05E56D1FAC1" bold="true" box="[192,395,1379,1403]" italics="true" pageId="32" pageNumber="41">Referred material</emphasis>
.—
<emphasis id="5FA8AB57FF940831FE4FA05E553DFAC1" bold="true" box="[427,615,1379,1403]" pageId="32" pageNumber="41">Tarsometatarsus</emphasis>
: proximal half of right,
<collectionCode id="0BCDEF80FF940831FC69A05E54BDFAC1" box="[909,999,1379,1403]" country="Cuba" name="Coleccion Zoologia, Academia de Ciencias de Cuba" pageId="32" pageNumber="41">CZACC</collectionCode>
unnumbered; proximal end of right,
<collectionCode id="0BCDEF80FF940831FE45A0BE5693FA21" box="[417,457,1411,1435]" pageId="32" pageNumber="41">OA</collectionCode>
3215 (
<bibRefCitation id="094D0AB4FF940831FDFEA0BE5416FA21" author="Suarez, W. &amp; Olson, S. L." box="[538,844,1411,1435]" pageId="32" pageNumber="41" pagination="533 - 553" refId="ref57070" refString="Suarez, W. &amp; Olson, S. L. 2015. Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa 4020: 533 - 553." type="journal article" year="2015">Suárez &amp; Olson 2015: 539</bibRefCitation>
, fig. 6: D [anterior]); distal half of left, DPUH 1252 (
<typeStatus id="B267C9E7FF940831FE4EA09E5554FA01" box="[426,526,1443,1467]" pageId="32" pageNumber="41" type="holotype">holotype</typeStatus>
of
<taxonomicName id="AADC0CC6FF940831FDD8A09954F4FA01" authority="', Arredondo 1972 b: 132" authorityName="', Arredondo" authorityPageNumber="132" authorityYear="1972" box="[572,942,1443,1467]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="32" pageNumber="41" phylum="Chordata" rank="species" species="riveroi">
<emphasis id="5FA8AB57FF940831FDD8A0995511FA01" box="[572,587,1444,1467]" italics="true" pageId="32" pageNumber="41">T</emphasis>
.
<emphasis id="5FA8AB57FF940831FDBEA09955C7FA01" box="[602,669,1444,1467]" italics="true" pageId="32" pageNumber="41">riveroi</emphasis>
,
<bibRefCitation id="094D0AB4FF940831FD55A09E54F4FA01" author="Arredondo, O." box="[689,942,1443,1467]" pageId="32" pageNumber="41" pagination="129 - 140" refId="ref47651" refString="Arredondo, O. 1972 b. Especie nueva de lechuza gigante (Strigiformes: Tytonidae) del Pleistoceno cubano. Bol. Soc. Venez. Cienc. Nat. 30 (124 - 125): 129 - 140." type="journal article" year="1972">Arredondo 1972b: 132</bibRefCitation>
</taxonomicName>
, figs. 1 [anterior], 2 [posterior], 3 [lateral], 4* = reversed: D, top row [distal], bottom row [anterior],
<bibRefCitation id="094D0AB4FF940831FBEBA0FE57ADFA41" author="Arredondo, O." pageId="32" pageNumber="41" refId="ref47789" refString="Arredondo, O. 1976. The great predatory birds of the Pleistocene of Cuba. Pp. 169 - 187 in Olson, S. L. (ed.) Collected papers in avian paleontology honoring the 90 th birthday of Alexander Wetmore. Smiths. Contrib. Paleobiol. 27." type="journal volume" year="1976">Arredondo 1976: 185</bibRefCitation>
, fig. 11: C [anterior], D [posterior], E [lateral], 1982: 39, fig. 2*: left [anterior], right [posterior]). Collected by Manuel Rivero de la Calle in Galería de los Megalocnus, Cueva de Bellamar (see
<bibRefCitation id="094D0AB4FF940831FEABA31E5513F981" author="Arredondo, O." box="[335,585,1571,1595]" pageId="32" pageNumber="41" pagination="129 - 140" refId="ref47651" refString="Arredondo, O. 1972 b. Especie nueva de lechuza gigante (Strigiformes: Tytonidae) del Pleistoceno cubano. Bol. Soc. Venez. Cienc. Nat. 30 (124 - 125): 129 - 140." type="journal article" year="1972">Arredondo 1972b: 131</bibRefCitation>
,
<bibRefCitation id="094D0AB4FF940831FDB0A31E542FF981" author="Suarez, W. &amp; Olson, S. L." box="[596,885,1571,1595]" pageId="32" pageNumber="41" pagination="533 - 553" refId="ref57070" refString="Suarez, W. &amp; Olson, S. L. 2015. Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa 4020: 533 - 553." type="journal article" year="2015">Suárez &amp; Olson 2015: 540</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="25C624CEFF940830FF24A37F5693FE23" lastPageId="33" lastPageNumber="42" pageId="32" pageNumber="41" type="distribution">
<paragraph id="6D637745FF940830FF24A37F57ACFF03" blockId="32.[144,1164,867,1691]" lastBlockId="33.[144,1164,129,409]" lastPageId="33" lastPageNumber="42" pageId="32" pageNumber="41">
<emphasis id="5FA8AB57FF940831FF24A37F5614F9E0" bold="true" box="[192,334,1602,1626]" italics="true" pageId="32" pageNumber="41">Distribution</emphasis>
.—Cave deposit in west
<collectingCountry id="15CB37D5FF940831FD80A37E55FAF9E1" box="[612,672,1603,1627]" name="Cuba" pageId="32" pageNumber="41">Cuba</collectingCountry>
(see Appendix).
<emphasis id="5FA8AB57FF940831FC80A3795493F9E1" box="[868,969,1604,1627]" italics="true" pageId="32" pageNumber="41">
<collectingRegion id="AF18B9A7FF940831FC80A3795493F9E1" box="[868,969,1604,1627]" country="Cuba" name="Matanzas" pageId="32" pageNumber="41">Matanzas</collectingRegion>
</emphasis>
.
<collectingRegion id="AF18B9A7FF940831FC3CA37E531DF9E0" box="[984,1095,1603,1626]" country="Cuba" name="Matanzas" pageId="32" pageNumber="41">Matanzas</collectingRegion>
:
<emphasis id="5FA8AB57FF940831FBB7A37E53D2F9E1" bold="true" box="[1107,1160,1603,1627]" pageId="32" pageNumber="41">MBE</emphasis>
(
<bibRefCitation id="094D0AB4FF940831FF73A35E56D7F9C1" author="Arredondo, O." box="[151,397,1635,1659]" pageId="32" pageNumber="41" pagination="129 - 140" refId="ref47651" refString="Arredondo, O. 1972 b. Especie nueva de lechuza gigante (Strigiformes: Tytonidae) del Pleistoceno cubano. Bol. Soc. Venez. Cienc. Nat. 30 (124 - 125): 129 - 140." type="journal article" year="1972">Arredondo 1972b: 132</bibRefCitation>
[
<taxonomicName id="AADC0CC6FF940831FE46A3595541F9C1" authority="Suárez, 2022" authorityName="Suárez" authorityYear="2022" box="[418,539,1636,1659]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="32" pageNumber="41" phylum="Chordata" rank="species" species="riveroi" status="sp. nov.">
<emphasis id="5FA8AB57FF940831FE46A3595541F9C1" box="[418,539,1636,1659]" italics="true" pageId="32" pageNumber="41">Tyto riveroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="449B162CFF940831FDC4A35E5526F9C1" box="[544,636,1635,1659]" pageId="32" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
],
<bibRefCitation id="094D0AB4FF940831FD77A35E54F4F9C1" author="Suarez, W. &amp; Olson, S. L." box="[659,942,1635,1659]" pageId="32" pageNumber="41" pagination="533 - 553" refId="ref57070" refString="Suarez, W. &amp; Olson, S. L. 2015. Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa 4020: 533 - 553." type="journal article" year="2015">Suárez &amp; Olson 2015: 539</bibRefCitation>
). For fossil localities in
<collectingCountry id="15CB37D5FF940831FF54A3BE5619F921" box="[176,323,1667,1691]" name="Bahamas" pageId="32" pageNumber="41">the Bahamas</collectingCountry>
see
<bibRefCitation id="094D0AB4FF940831FE9FA3BE5533F921" author="Wetmore, A." box="[379,617,1667,1691]" pageId="32" pageNumber="41" pagination="427 - 441" refId="ref57784" refString="Wetmore, A. 1937. Bird remains from cave deposits on Great Exuma Island in the Bahamas. Bull. Mus. Comp. Zool. 80: 427 - 441." type="journal article" year="1937">Wetmore (1937: 437)</bibRefCitation>
,
<bibRefCitation id="094D0AB4FF940831FD9CA3BE54C6F921" author="Brodkorb, P." box="[632,924,1667,1691]" pageId="32" pageNumber="41" pagination="349 - 371" refId="ref48589" refString="Brodkorb, P. 1959. Pleistocene birds from New Providence Island, Bahamas. Bull. Fla. State Mus. 4: 349 - 371." type="journal article" year="1959">Brodkorb (1959: 357358)</bibRefCitation>
,
<bibRefCitation id="094D0AB4FF940830FC4FA3BE567FFF23" author="Olson, S. L. &amp; Hilgartner, W. B." lastPageId="33" lastPageNumber="42" pageId="32" pageNumber="41" refId="ref53860" refString="Olson, S. L. &amp; Hilgartner, W. B. 1982. Fossil and subfossil birds from the Bahamas. Pp. 22 - 56 in Olson, S. L. (ed.) Fossil vertebrates from the Bahamas. Smiths. Contrib. Paleobiol. 48." type="journal volume" year="1982">Olson &amp; Hilgartner (1982: 3637)</bibRefCitation>
,
<bibRefCitation id="094D0AB4FF950830FED0A5BC5535FF23" author="Suarez, W. &amp; Olson, S. L." box="[308,623,129,153]" pageId="33" pageNumber="42" pagination="533 - 553" refId="ref57070" refString="Suarez, W. &amp; Olson, S. L. 2015. Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa 4020: 533 - 553." type="journal article" year="2015">Suárez &amp; Olson (2015: 540)</bibRefCitation>
and
<bibRefCitation id="094D0AB4FF950830FD4BA5BC54B4FF23" author="Steadman, D. W. &amp; Franklin, J." box="[687,1006,129,153]" pageId="33" pageNumber="42" pagination="26833 - 26841" refId="ref55570" refString="Steadman, D. W. &amp; Franklin, J. 2020. Bird populations and species lost to Late Quaternary environmental change and human impact in the Bahamas. Proc. Natl. Acad. Sci. USA 117: 26833 - 26841." type="journal article" year="2020">Steadman &amp; Franklin (2020</bibRefCitation>
: SI appendix, table S1).
</paragraph>
<paragraph id="6D637745FF950830FF24A5FC56B8FF62" blockId="33.[144,1164,129,409]" box="[192,482,191,217]" pageId="33" pageNumber="42">
<emphasis id="5FA8AB57FF950830FF24A5FC5627FF62" bold="true" box="[192,381,191,217]" italics="true" pageId="33" pageNumber="42">
Direct
<superScript id="9AA9DA0DFF950830FEE9A5825646FF77" attach="right" box="[269,284,191,205]" fontSize="6" pageId="33" pageNumber="42">14</superScript>
<taxonomicName id="AADC0CC6FF950830FEF8A5FC5627FF62" box="[284,381,192,217]" class="Aves" family="Caprimulgidae" genus="Caprimulgus" kingdom="Animalia" order="Caprimulgiformes" pageId="33" pageNumber="42" phylum="Chordata" rank="species" species="dating">C dating</taxonomicName>
</emphasis>
.—None.
</paragraph>
<paragraph id="6D637745FF950830FF24A5DC5693FE23" blockId="33.[144,1164,129,409]" pageId="33" pageNumber="42">
<emphasis id="5FA8AB57FF950830FF24A5DC5658FF42" bold="true" box="[192,258,225,248]" italics="true" pageId="33" pageNumber="42">Notes</emphasis>
.—The largest
<taxonomicName id="AADC0CC6FF950830FE4EA5DF5680FF43" authorityName="Billberg" authorityYear="1828" box="[426,474,226,249]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="33" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="5FA8AB57FF950830FE4EA5DF5680FF43" box="[426,474,226,249]" italics="true" pageId="33" pageNumber="42">Tyto</emphasis>
</taxonomicName>
in the Americas (
<bibRefCitation id="094D0AB4FF950830FD4EA5DC54A8FF43" author="Olson, S. L. &amp; Hilgartner, W. B." box="[682,1010,225,249]" pageId="33" pageNumber="42" refId="ref53860" refString="Olson, S. L. &amp; Hilgartner, W. B. 1982. Fossil and subfossil birds from the Bahamas. Pp. 22 - 56 in Olson, S. L. (ed.) Fossil vertebrates from the Bahamas. Smiths. Contrib. Paleobiol. 48." type="journal volume" year="1982">Olson &amp; Hilgartner 1982: 36</bibRefCitation>
,
<bibRefCitation id="094D0AB4FF950830FC1BA5DC5635FEA3" author="Steadman, D. W. &amp; Hilgartner, W. B." pageId="33" pageNumber="42" refId="ref55615" refString="Steadman, D. W. &amp; Hilgartner, W. B. 1999. A new species of extinct barn owl (Aves: Tyto) from Barbuda, Lesser Antilles. Pp. 75 - 83 in Olson, S. L., Wellnhofer, P., Mourer-Chauvire, C., Steadman, D. W. &amp; Martin, L. D. (eds.) Avian paleontology at the close of the 20 th century: Proc. 4 th Intern. Meet. Soc. Avian Paleontology &amp; Evolution, Washington, DC, 1996. Smiths. Contrib. Paleobiol. 89." type="journal volume" year="1999">Steadman &amp; Hilgartner 1999: 79</bibRefCitation>
,
<bibRefCitation id="094D0AB4FF950830FE99A43C55F3FEA3" author="Suarez, W. &amp; Olson, S. L." box="[381,681,257,281]" pageId="33" pageNumber="42" pagination="533 - 553" refId="ref57070" refString="Suarez, W. &amp; Olson, S. L. 2015. Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa 4020: 533 - 553." type="journal article" year="2015">Suárez &amp; Olson 2015: 540</bibRefCitation>
) and the rarest of the giant barn owls in Cuban fossil deposits (see Notes under
<taxonomicName id="AADC0CC6FF950830FD8CA41F5580FE83" box="[616,730,290,313]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="33" pageNumber="42" phylum="Chordata" rank="species" species="cravesae">
<emphasis id="5FA8AB57FF950830FD8CA41F5580FE83" box="[616,730,290,313]" italics="true" pageId="33" pageNumber="42">T. cravesae</emphasis>
</taxonomicName>
).
<taxonomicName id="AADC0CC6FF950830FD16A41F5402FE83" box="[754,856,289,313]" class="Aves" family="Tytonidae" genus="Tyto" kingdom="Animalia" order="Strigiformes" pageId="33" pageNumber="42" phylum="Chordata" rank="species" species="pollens">
<emphasis id="5FA8AB57FF950830FD16A41F5402FE83" box="[754,856,289,313]" italics="true" pageId="33" pageNumber="42">T. pollens</emphasis>
</taxonomicName>
probably evolved in
<collectingCountry id="15CB37D5FF950830FBB4A41C53D6FE83" box="[1104,1164,289,313]" name="Cuba" pageId="33" pageNumber="42">Cuba</collectingCountry>
and subsequently colonised
<collectingCountry id="15CB37D5FF950830FE35A47C553BFEE3" box="[465,609,321,345]" name="Bahamas" pageId="33" pageNumber="42">the Bahamas</collectingCountry>
(
<bibRefCitation id="094D0AB4FF950830FD95A47C54CFFEE3" author="Suarez, W. &amp; Olson, S. L." box="[625,917,321,345]" pageId="33" pageNumber="42" pagination="533 - 553" refId="ref57070" refString="Suarez, W. &amp; Olson, S. L. 2015. Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa 4020: 533 - 553." type="journal article" year="2015">Suárez &amp; Olson 2015: 549</bibRefCitation>
), where it is recorded from Quaternary deposits on some islands of the Great Bahama Bank (
<bibRefCitation id="094D0AB4FF950830FC7BA45C53D2FEC3" author="Suarez, W. &amp; Olson, S. L." box="[927,1160,353,377]" pageId="33" pageNumber="42" pagination="533 - 553" refId="ref57070" refString="Suarez, W. &amp; Olson, S. L. 2015. Systematics and distribution of the giant fossil barn owls of the West Indies (Aves: Strigiformes: Tytonidae). Zootaxa 4020: 533 - 553." type="journal article" year="2015">Suárez &amp; Olson 2015</bibRefCitation>
,
<bibRefCitation id="094D0AB4FF950830FF74A4BC56E4FE23" author="Steadman, D. W. &amp; Franklin, J." box="[144,446,385,409]" pageId="33" pageNumber="42" pagination="26833 - 26841" refId="ref55570" refString="Steadman, D. W. &amp; Franklin, J. 2020. Bird populations and species lost to Late Quaternary environmental change and human impact in the Bahamas. Proc. Natl. Acad. Sci. USA 117: 26833 - 26841." type="journal article" year="2020">Steadman &amp; Franklin 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>