327 lines
39 KiB
XML
327 lines
39 KiB
XML
<document ID-DOI="10.1590/S1679-62252005000200001" ID-ISSN="1982-0224" ID-Zenodo-Dep="5418569" approvalRequired="104" approvalRequired_for_taxonomicNames="6" approvalRequired_for_textStreams="91" approvalRequired_for_treatments="7" checkinTime="1630369289621" checkinUser="carolina" docAuthor="Vari, Richard P., Ferraris Jr, Carl J. & de Pinna, Mário C. C." docDate="2005" docId="E56BC71F0E73FFE63E76FAB4FBF4E6C3" docLanguage="en" docName="NeotropIchthyol.2005.8.pdf" docOrigin="Neotropical Ichthyology 3 (2)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1679-62252005000200001&lng=en&tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Cetopsis sandrae Vari & Ferraris Jr & de Pinna 2005, new species" docType="treatment" docVersion="2" lastPageId="70" lastPageNumber="197" masterDocId="1952BF670E37FFA03F4CFFA8FFCEE255" masterDocTitle="The Neotropical whale catfishes (Siluriformes: Cetopsidae: Cetopsinae), a revisionary study" masterLastPageNumber="238" masterPageNumber="127" pageId="68" pageNumber="195" updateTime="1630681654510" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo>
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<mods:title>The Neotropical whale catfishes (Siluriformes: Cetopsidae: Cetopsinae), a revisionary study</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Vari, Richard P.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ferraris Jr, Carl J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>de Pinna, Mário C. C.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Neotropical Ichthyology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2005</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2005-06-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>127</mods:start>
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<mods:end>238</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1679-62252005000200001&lng=en&tlng=en</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1590/S1679-62252005000200001</mods:identifier>
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<mods:identifier type="ISSN">1982-0224</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5418569</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:E56BC71F0E73FFE63E76FAB4FBF4E6C3" httpUri="http://treatment.plazi.org/id/E56BC71F0E73FFE63E76FAB4FBF4E6C3" lastPageId="70" lastPageNumber="197" pageId="68" pageNumber="195">
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<subSubSection box="[314,645,1308,1332]" pageId="68" pageNumber="195" type="nomenclature">
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<paragraph blockId="68.[314,645,1308,1363]" box="[314,645,1308,1332]" pageId="68" pageNumber="195">
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<heading box="[314,645,1308,1332]" centered="true" fontSize="10" level="2" pageId="68" pageNumber="195" reason="2">
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<emphasis bold="true" box="[314,645,1308,1332]" pageId="68" pageNumber="195">
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<taxonomicName authority="Vari & Ferraris Jr & de Pinna, 2005" authorityName="Vari & Ferraris Jr & de Pinna" authorityYear="2005" box="[314,501,1308,1332]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="68" pageNumber="195" phylum="Chordata" rank="species" species="sandrae" status="new species">
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<emphasis bold="true" box="[314,501,1308,1332]" italics="true" pageId="68" pageNumber="195">Cetopsis sandrae</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[514,645,1308,1331]" pageId="68" pageNumber="195" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[356,603,1340,1363]" pageId="68" pageNumber="195" type="description">
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<paragraph blockId="68.[314,645,1308,1363]" box="[356,603,1340,1363]" pageId="68" pageNumber="195">
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<figureCitation box="[356,475,1340,1363]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="69.[151,190,614,637]" captionStartId-1="69.[151,190,1110,1133]" captionStartId-2="70.[158,197,831,854]" captionTargetBox-0="[244,1372,207,582]" captionTargetBox-1="[252,1389,711,1085]" captionTargetBox-2="[163,798,210,798]" captionTargetId-0="figure-415@69.[204,1413,184,602]" captionTargetId-1="figure-417@69.[215,1425,688,1103]" captionTargetId-2="figure-704@70.[158,798,170,798]" captionTargetPageId-0="69" captionTargetPageId-1="69" captionTargetPageId-2="70" captionText-0="Fig. 39. Cetopsis sandrae, new species, holotype, male, MZUSP 61091, 64 mm SL; Brazil, Mato Grosso, Nova Mutum, unnamed Córrego, Fazenda Buriti (13°51’52"S, 56°11’36"W)." captionText-1="Fig. 40. Cetopsis sandrae, new species, paratype, female, MZUSP 83227, 75 mm SL; Brazil, Mato Grosso, Nova Mutum, unnamed Córrego, Fazenda Buriti (13°51’52"S, 56°11’36"W)." captionText-2="Fig. 41. Map of central and northern SouthAmerica showing geographic distribution of Cetopsis sandrae (1 = holotype locality), Cetopsis sarcodes (2 = holotype locality), Cetopsis starnesi (3 = holotype locality), and Cetopsis umbrosa (4 = holotype locality) (some symbols represent more that one locality and/or lot of specimens)." figureDoi-0="http://doi.org/10.5281/zenodo.5418656" figureDoi-1="http://doi.org/10.5281/zenodo.5418658" figureDoi-2="http://doi.org/10.5281/zenodo.5418660" httpUri-0="https://zenodo.org/record/5418656/files/figure.png" httpUri-1="https://zenodo.org/record/5418658/files/figure.png" httpUri-2="https://zenodo.org/record/5418660/files/figure.png" pageId="68" pageNumber="195">Figs. 39-41</figureCitation>
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,
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<tableCitation box="[483,569,1340,1363]" captionStart="Table 9" captionStartId="76.[158,220,197,220]" captionText="Table 9. Number of specimens of the species of Cetopsis with cited number of dorsal- and pectoral-fin rays. Bold indicates value for holotype or lectotype when examined. Dorsal fin of C. jurubidae damaged and count of rays uncertain; 6 branched dorsal-fin rays cited for holotype of species by Fowler (1944:235)." pageId="68" pageNumber="195">Tables 9</tableCitation>
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-15
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</paragraph>
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</subSubSection>
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<subSubSection pageId="68" pageNumber="195" type="diagnosis">
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<paragraph blockId="68.[158,801,1409,1977]" pageId="68" pageNumber="195">
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<emphasis bold="true" box="[158,276,1410,1433]" pageId="68" pageNumber="195">Diagnosis.</emphasis>
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<taxonomicName authorityName="Vari & Ferraris Jr & de Pinna" authorityYear="2005" box="[280,464,1409,1433]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="68" pageNumber="195" phylum="Chordata" rank="species" species="sandrae">
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<emphasis box="[280,464,1409,1433]" italics="true" pageId="68" pageNumber="195">Cetopsis sandrae</emphasis>
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</taxonomicName>
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can be distinguished from all of its congeners by the combination of the presence of an eye, the conical teeth on the vomer and the dentary, the rounded posterior nares that is distinctly separated from the contralateral nares by a distance greater than the width of the posterior nares, the absence of a dark humeral spot, the presence of a posteriorly-rounded, variably-developed, bilobed patch of dark pigmentation at the base of the caudal fin, the presence of a spot of dark pigmentation on the base of the dorsal fin with prominent dark pigmentation present along the membrane behind the first ray of the dorsal fin, the possession of 40 or 41 total vertebrae (with 40 most common), the lack of a distinct demarcation between the dark pigmentation on the dorsal and dorsolateral surfaces of the body
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<emphasis box="[620,688,1826,1849]" italics="true" pageId="68" pageNumber="195">versus</emphasis>
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more ventral regions, and the presence of eye size or larger, dark spots on the ventral and ventrolateral surfaces of the body in the region below the horizontal extending through the ventral margin of the caudal peduncle.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="70" lastPageNumber="197" pageId="68" pageNumber="195" type="description">
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<paragraph blockId="68.[839,1482,194,1977]" pageId="68" pageNumber="195">
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<emphasis bold="true" box="[839,980,194,217]" pageId="68" pageNumber="195">Description.</emphasis>
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Body moderately stout, slightly-compressed laterally anteriorly and becoming progressively distinctlycompressed posteriorly. Body depth at dorsal-fin origin approximately 0.29-0.30 of SL, and slightly greater than HL. Lateral line on body complete, unbranched, and midlateral; extending from vertical through pectoral-fin base to hypural plate. Dorsal profile of body slightly convex from nape to dorsal-fin origin and nearly straight from dorsal-fin origin to caudal-fin base. Ventral profile of body convex along abdomen, approximately straight, but posterodorsally-slanted, along base of anal fin. Caudal-peduncle depth slightly greater than caudal-peduncle length.
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</paragraph>
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<paragraph blockId="68.[839,1482,194,1977]" pageId="68" pageNumber="195">Head in profile acutely triangular overall with bluntlyrounded snout. Dorsal profile of head slightly convex from tip of snout to nape. Ventral profile of head slightly convex. Margin of snout in dorsal view bluntly triangular. Postorbital margins of head aligned nearly in parallel from dorsal view. Enlarged jaw musculature apparent externally on dorsal surface of postorbital portion of head.</paragraph>
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<paragraph blockId="68.[839,1482,194,1977]" pageId="68" pageNumber="195">Opercular membrane attaching to isthmus only anterior of vertical through pectoral-fin insertion. Opercular opening moderate; extending ventral of pectoral-fin insertion by distance equal to distance from tip of snout to middle of eye and extending dorsal of pectoral-fin insertion by distance approximately equal to snout length.</paragraph>
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<paragraph blockId="68.[839,1482,194,1977]" pageId="68" pageNumber="195">Eye situated on lateral surface of head; located entirely dorsal to horizontal extending through pectoral-fin insertion; eye visible in dorsal view, but not in ventral view, of head. Middle of orbit at approximately anterior 0.30 of HL. Eye diameter approximately one-half of snout length. Interorbital width approximately equal to distance from tip of snout to posterior margin of orbit. Anterior narial opening circular, surrounded by short, anteriorly-directed, tubular rim of skin. Opening of anterior nares located along horizontal extending through maxillary-barbel origin and ventral of horizontal extending through tip of snout. Distance between anterior nares approximately equal to snout length. Posterior narial opening located on dorsal surface of head, situated along vertical through anterior margin of orbit; narial opening nearly round, with anterior two-thirds of aperture surrounded by flap of skin with anterior portion of flap highest.</paragraph>
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<paragraph blockId="68.[839,1482,194,1977]" pageId="68" pageNumber="195">Mouth inferior; its width approximately one-half of HL. Margin of lower jaw gently rounded, its posterior limit reaching to vertical through posterior margin of orbit. Premaxillary tooth patch in form of gently-arched band, continuous across midline and with anterior margin convex and posterior margin concave and running in parallel to anterior margin. Teeth on premaxilla small, conical, sharply-pointed, uniform-sized, and arranged in three regular rows. Vomerine teeth arranged in single irregular row apparently continuous across midline. Vomerine teeth stout, conical, and much larger than teeth on premaxilla and approximately as large as largest teeth on dentary. Dentary teeth comparable in shape to, but larger in size than, premaxillary teeth. Dentary dentition consisting of two or three irregular rows medially that taper to one row laterally.</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.5418656" ID-Zenodo-Dep="5418656" httpUri="https://zenodo.org/record/5418656/files/figure.png" pageId="69" pageNumber="196" startId="69.[151,190,614,637]" targetBox="[244,1372,207,582]" targetPageId="69">
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<paragraph blockId="69.[151,1474,613,669]" pageId="69" pageNumber="196">
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<emphasis bold="true" box="[151,237,614,637]" pageId="69" pageNumber="196">Fig. 39.</emphasis>
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<taxonomicName authority="Vari & Ferraris Jr & de Pinna, 2005" authorityName="Vari & Ferraris Jr & de Pinna" authorityYear="2005" box="[245,434,613,637]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="69" pageNumber="196" phylum="Chordata" rank="species" species="sandrae" status="new species">
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<emphasis box="[245,434,613,637]" italics="true" pageId="69" pageNumber="196">Cetopsis sandrae</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[448,583,614,637]" pageId="69" pageNumber="196" rank="species">new species</taxonomicNameLabel>
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, holotype, male, MZUSP 61091, 64 mm SL; Brazil, Mato Grosso, Nova Mutum, unnamed Córrego, Fazenda Buriti (13°51’52"S, 56°11’36"W).
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</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.5418658" ID-Zenodo-Dep="5418658" httpUri="https://zenodo.org/record/5418658/files/figure.png" pageId="69" pageNumber="196" startId="69.[151,190,1110,1133]" targetBox="[252,1389,711,1085]" targetPageId="69">
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<paragraph blockId="69.[151,1474,1109,1165]" pageId="69" pageNumber="196">
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<emphasis bold="true" box="[151,236,1110,1133]" pageId="69" pageNumber="196">Fig. 40.</emphasis>
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<taxonomicName authority="Vari & Ferraris Jr & de Pinna, 2005" authorityName="Vari & Ferraris Jr & de Pinna" authorityYear="2005" box="[243,430,1109,1133]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="69" pageNumber="196" phylum="Chordata" rank="species" species="sandrae" status="new species">
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<emphasis box="[243,430,1109,1133]" italics="true" pageId="69" pageNumber="196">Cetopsis sandrae</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[444,577,1110,1133]" pageId="69" pageNumber="196" rank="species">new species</taxonomicNameLabel>
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, paratype, female, MZUSP 83227, 75 mm SL; Brazil, Mato Grosso, Nova Mutum, unnamed Córrego, Fazenda Buriti (13°51’52"S, 56°11’36"W).
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</paragraph>
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</caption>
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<paragraph blockId="69.[151,794,1218,1977]" pageId="69" pageNumber="196">Maxillary barbel slender, its length approximately equal to distance from tip of snout to posterior margin of eye, and nearly one-half of HL; barbel origin located ventral to middle of orbit. Medial mental barbels approximately same length as lateral mental barbels and slightly shorter than maxillary barbels. Medial mental-barbel origin located along vertical through rictus. Lateral mental-barbel origin situated slightly posterior of vertical through medial mental-barbel origin. Tips of adpressed mental barbels falling just short of posterior margin of opercle.</paragraph>
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<paragraph blockId="69.[151,794,1218,1977]" pageId="69" pageNumber="196">
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Dorsal fin moderately large overall with length of dorsalfin base approximately 0.40-0.41 of HL. Longest branched dorsal-fin ray, excluding distal filament on first ray, equal in length to two-thirds of HL. Dorsal-fin spinelet absent. First dorsal-fin ray not spinous but with at least short, distal filament in all specimens, and with filament proportionally longer in apparently mature males (
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<figureCitation box="[452,531,1730,1753]" captionStart="Fig" captionStartId="69.[151,190,614,637]" captionTargetBox="[244,1372,207,582]" captionTargetId="figure-415@69.[204,1413,184,602]" captionTargetPageId="69" captionText="Fig. 39. Cetopsis sandrae, new species, holotype, male, MZUSP 61091, 64 mm SL; Brazil, Mato Grosso, Nova Mutum, unnamed Córrego, Fazenda Buriti (13°51’52"S, 56°11’36"W)." figureDoi="http://doi.org/10.5281/zenodo.5418656" httpUri="https://zenodo.org/record/5418656/files/figure.png" pageId="69" pageNumber="196">Fig. 39</figureCitation>
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). Distal margin of dorsal fin slightly sinusoidal with anterior portion concave and posterior portion convex. Dorsal-fin origin located at approximately anterior 0.36-0.37 of SL and along vertical extending through middle of adpressed pectoral fin. Tip of adpressed dorsal fin, excluding filament, nearly reaching to vertical through vent. Posterior most dorsal-fin ray without posterior, membranous attachment to body.
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</paragraph>
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<paragraph blockId="69.[831,1475,1218,1977]" pageId="69" pageNumber="196">Caudal fin moderately-forked, symmetrical; tips of lobes rounded. Length of longest caudal-fin ray approximately 1.75 times length of middle fin rays.</paragraph>
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<paragraph blockId="69.[831,1475,1218,1977]" pageId="69" pageNumber="196">
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Base of anal fin moderately long. Anal-fin origin located distinctly posterior of middle of SL and nearly at middle of TL.Anal-fin margin straight in females (
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<figureCitation box="[1245,1320,1378,1401]" captionStart="Fig" captionStartId="69.[151,190,1110,1133]" captionTargetBox="[252,1389,711,1085]" captionTargetId="figure-417@69.[215,1425,688,1103]" captionTargetPageId="69" captionText="Fig. 40. Cetopsis sandrae, new species, paratype, female, MZUSP 83227, 75 mm SL; Brazil, Mato Grosso, Nova Mutum, unnamed Córrego, Fazenda Buriti (13°51’52"S, 56°11’36"W)." figureDoi="http://doi.org/10.5281/zenodo.5418658" httpUri="https://zenodo.org/record/5418658/files/figure.png" pageId="69" pageNumber="196">Fig. 40</figureCitation>
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) and immature males, with posterior most unbranched anal-fin ray longest and with subsequent rays becoming gradually shorter. Analfin margin convex in mature males (
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<figureCitation box="[1205,1281,1474,1497]" captionStart="Fig" captionStartId="69.[151,190,614,637]" captionTargetBox="[244,1372,207,582]" captionTargetId="figure-415@69.[204,1413,184,602]" captionTargetPageId="69" captionText="Fig. 39. Cetopsis sandrae, new species, holotype, male, MZUSP 61091, 64 mm SL; Brazil, Mato Grosso, Nova Mutum, unnamed Córrego, Fazenda Buriti (13°51’52"S, 56°11’36"W)." figureDoi="http://doi.org/10.5281/zenodo.5418656" httpUri="https://zenodo.org/record/5418656/files/figure.png" pageId="69" pageNumber="196">Fig. 39</figureCitation>
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) and in apparently maturing male of
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<quantity box="[1021,1096,1506,1529]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="69" pageNumber="196" unit="mm" value="45.0">45 mm</quantity>
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SL that does not demonstrate other sexually-dimorphic male characteristics of dorsal and pectoral fins. Posterior most anal-fin ray without posterior, membranous attachment to body.
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</paragraph>
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<paragraph blockId="69.[831,1475,1218,1977]" pageId="69" pageNumber="196">Pelvic fin moderately long; distal margin distinctly convex, with first ray longest. Pelvic-fin insertion located anterior to middle of SL and along vertical through posterior terminus of base of dorsal fin. Tip of adpressed pelvic fin extending past middle of SL, but falling short of anterior margin of vent. Medial most pelvic-fin ray with membranous attachment to body along basal two-thirds of its length.</paragraph>
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<paragraph blockId="69.[831,1475,1218,1977]" pageId="69" pageNumber="196">Pectoral-fin length approximately 0.60 of HL. Pectoral-fin margin slightly convex, with first ray longest and prolonged into filament, filament proportionally longer in mature males. First pectoral-fin ray not spinous.</paragraph>
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<paragraph blockId="70.[839,1482,192,280]" pageId="70" pageNumber="197">
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<taxonomicName authorityName="Vari & Ferraris Jr & de Pinna" authorityYear="2005" box="[839,924,192,216]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="70" pageNumber="197" phylum="Chordata" rank="species" species="sandrae">
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<emphasis box="[839,924,192,216]" italics="true" pageId="70" pageNumber="197">sandrae</emphasis>
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</taxonomicName>
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also have the anal-fin margin distinctly convex rather than straight as occurs in females and immature males of that species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="70" pageNumber="197" type="biology_ecology">
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<paragraph blockId="70.[839,1482,320,504]" pageId="70" pageNumber="197">
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<emphasis bold="true" box="[839,934,321,344]" pageId="70" pageNumber="197">Ecology.</emphasis>
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<materialsCitation collectionCode="T, MZUSP" collectorName="All & F Chang" elevation="4" location="Lima" pageId="70" pageNumber="197" specimenCount="2" stateProvince="Lima" typeStatus="holotype">
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The
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<typeStatus box="[985,1078,321,344]" pageId="70" pageNumber="197">holotype</typeStatus>
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of
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<taxonomicName authorityName="Vari & Ferraris Jr & de Pinna" authorityYear="2005" box="[1111,1293,320,344]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="70" pageNumber="197" phylum="Chordata" rank="species" species="sandrae">
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<emphasis box="[1111,1293,320,344]" italics="true" pageId="70" pageNumber="197">Cetopsis sandrae</emphasis>
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</taxonomicName>
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and the
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<typeStatus box="[1379,1481,321,344]" pageId="70" pageNumber="197">paratypes</typeStatus>
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of the species collected with it were collected in a relatively fast-flowing forest stream, approximately
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<quantity box="[1275,1336,385,408]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="70" pageNumber="197" unit="m" value="3.5" valueMax="4.0" valueMin="3.0">
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<elevation box="[1275,1336,385,408]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="70" pageNumber="197" unit="m" value="3.5" valueMax="4.0" valueMin="3.0">3-4 m</elevation>
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</quantity>
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wide and 0.4- 0.8 m deep with a small waterfall.
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<collectorName box="[1208,1242,417,440]" pageId="70" pageNumber="197">All</collectorName>
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the samples were collected in riffles along the stream (
|
||
<collectorName box="[1188,1232,449,472]" name="F Chang" pageId="70" pageNumber="197">F.C.</collectorName>
|
||
<collectionCode box="[1232,1250,449,472]" name="Tavera, Department of Geology and Geophysics" pageId="70" pageNumber="197">T</collectionCode>
|
||
.
|
||
<collectingRegion box="[1259,1318,449,472]" country="Peru" name="Lima" pageId="70" pageNumber="197">Lima</collectingRegion>
|
||
,
|
||
<collectionCode box="[1327,1420,449,472]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="70" pageNumber="197">MZUSP</collectionCode>
|
||
;
|
||
<emphasis italics="true" pageId="70" pageNumber="197">pers. commun.,</emphasis>
|
||
2003)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="70" pageNumber="197" type="distribution">
|
||
<paragraph blockId="70.[839,1481,544,632]" pageId="70" pageNumber="197">
|
||
<emphasis bold="true" box="[839,987,545,568]" pageId="70" pageNumber="197">Distribution.</emphasis>
|
||
<taxonomicName authorityName="Vari & Ferraris Jr & de Pinna" authorityYear="2005" box="[994,1182,544,568]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="70" pageNumber="197" phylum="Chordata" rank="species" species="sandrae">
|
||
<emphasis box="[994,1182,544,568]" italics="true" pageId="70" pageNumber="197">Cetopsis sandrae</emphasis>
|
||
</taxonomicName>
|
||
is known only from the rio Arinos in the upper portions of the rio Tapajós system, Amazon basin (
|
||
<figureCitation box="[951,1026,609,632]" captionStart="Fig" captionStartId="70.[158,197,831,854]" captionTargetBox="[163,798,210,798]" captionTargetId="figure-704@70.[158,798,170,798]" captionTargetPageId="70" captionText="Fig. 41. Map of central and northern SouthAmerica showing geographic distribution of Cetopsis sandrae (1 = holotype locality), Cetopsis sarcodes (2 = holotype locality), Cetopsis starnesi (3 = holotype locality), and Cetopsis umbrosa (4 = holotype locality) (some symbols represent more that one locality and/or lot of specimens)." figureDoi="http://doi.org/10.5281/zenodo.5418660" httpUri="https://zenodo.org/record/5418660/files/figure.png" pageId="70" pageNumber="197">Fig. 41</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.5418660" ID-Zenodo-Dep="5418660" httpUri="https://zenodo.org/record/5418660/files/figure.png" pageId="70" pageNumber="197" startId="70.[158,197,831,854]" targetBox="[163,798,210,798]" targetPageId="70">
|
||
<paragraph blockId="70.[158,801,831,1014]" pageId="70" pageNumber="197">
|
||
<emphasis bold="true" box="[158,241,831,854]" pageId="70" pageNumber="197">Fig. 41.</emphasis>
|
||
Map of central and northern South America showing geographic distribution of
|
||
<taxonomicName authorityName="Vari & Ferraris Jr & de Pinna" authorityYear="2005" box="[453,642,862,886]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="70" pageNumber="197" phylum="Chordata" rank="species" species="sandrae">
|
||
<emphasis box="[453,642,862,886]" italics="true" pageId="70" pageNumber="197">Cetopsis sandrae</emphasis>
|
||
</taxonomicName>
|
||
(1 = holotype locality),
|
||
<taxonomicName authorityName="Vari & Ferraris Jr & de Pinna" authorityYear="2005" box="[260,454,894,918]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="70" pageNumber="197" phylum="Chordata" rank="species" species="sarcodes">
|
||
<emphasis box="[260,454,894,918]" italics="true" pageId="70" pageNumber="197">Cetopsis sarcodes</emphasis>
|
||
</taxonomicName>
|
||
(2 = holotype locality),
|
||
<taxonomicName authorityName="Vari & Ferraris Jr & de Pinna" authorityYear="2005" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="70" pageNumber="197" phylum="Chordata" rank="species" species="starnesi">
|
||
<emphasis italics="true" pageId="70" pageNumber="197">Cetopsis starnesi</emphasis>
|
||
</taxonomicName>
|
||
(3 = holotype locality), and
|
||
<taxonomicName authorityName="Vari & Ferraris Jr & de Pinna" authorityYear="2005" box="[555,749,926,950]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="70" pageNumber="197" phylum="Chordata" rank="species" species="umbrosa">
|
||
<emphasis box="[555,749,926,950]" italics="true" pageId="70" pageNumber="197">Cetopsis umbrosa</emphasis>
|
||
</taxonomicName>
|
||
(4 = holotype locality) (some symbols represent more that one locality and/or lot of specimens).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="70.[158,802,1058,1817]" pageId="70" pageNumber="197">
|
||
<emphasis bold="true" box="[158,411,1058,1081]" pageId="70" pageNumber="197">Coloration in alcohol.</emphasis>
|
||
Dark pigmentation covering dorsal portion of head from tip of snout to posterior margin of head. Dark, dorsal pigmentation continuous with dark region posterodorsal and posterior to orbit. Remaining portions of head pale. Dorsal portion of body dark. Lateral surface of body irregularly covered with fine, dark spots; spots often coalesce into eye-size blotches, with such coalescence more prevalent ventral of lateral line. Heavily pigmented individuals with lateral surface of body nearly uniformly dark. Ventral surface of body pale, with few, scattered, dark spots present only in most heavily pigmented individuals.
|
||
</paragraph>
|
||
<paragraph blockId="70.[158,802,1058,1817]" pageId="70" pageNumber="197">Dorsal fin irregularly covered with dark pigmentation; pigmentation more concentrated basally and forming semicircular dark spot above base of fin. Anterior margin and distal portions of dorsal fin pale, but with dark pigmentation along first interradial membrane. Caudal-fin rays outlined by scattered, dark pigmentation except for narrow, clear band along distal margin of fin. Anal fin with scattered, dark pigmentation along base and with dark pigmentation rarely extending varying distances onto main body of fin. Pectoral fin pale other than for occasional dark pigment spots on basal or distal portions of first fin ray. Pelvic fin pale.</paragraph>
|
||
<paragraph blockId="70.[158,802,1058,1817]" pageId="70" pageNumber="197">Maxillary and mental barbel dusky basally and with distal portion pale.</paragraph>
|
||
<paragraph blockId="70.[158,801,1857,1977]" lastBlockId="70.[839,1481,672,824]" pageId="70" pageNumber="197">
|
||
<emphasis bold="true" box="[158,384,1858,1881]" pageId="70" pageNumber="197">Sexual dimorphism.</emphasis>
|
||
Mature males of
|
||
<taxonomicName authorityName="Vari & Ferraris Jr & de Pinna" authorityYear="2005" box="[564,746,1857,1881]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="70" pageNumber="197" phylum="Chordata" rank="species" species="sandrae">
|
||
<emphasis box="[564,746,1857,1881]" italics="true" pageId="70" pageNumber="197">Cetopsis sandrae</emphasis>
|
||
</taxonomicName>
|
||
have the filaments on the first rays of the dorsal and pectoral fins proportionally more elongate than are the extensions on those ray in conspecific females and immature males. Males of
|
||
<emphasis box="[776,801,1954,1977]" italics="true" pageId="70" pageNumber="197">C.</emphasis>
|
||
<emphasis bold="true" box="[839,968,673,696]" pageId="70" pageNumber="197">Etymology.</emphasis>
|
||
The specific name,
|
||
<emphasis box="[1193,1286,672,696]" italics="true" pageId="70" pageNumber="197">
|
||
<taxonomicName authorityName="Vari & Ferraris Jr & de Pinna" authorityYear="2005" box="[1193,1283,672,696]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsis" kingdom="Animalia" order="Siluriformes" pageId="70" pageNumber="197" phylum="Chordata" rank="species" species="sandrae">sandrae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
is in reference to Sandra J. Raredon, of the Division of Fishes, National Museum of Natural History, in thanks for all of her assistance to the authors, particularly the first author, over the course of the years in this and many other projects.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="70" pageNumber="197" type="materials_examined">
|
||
<paragraph blockId="70.[839,1482,865,1174]" pageId="70" pageNumber="197">
|
||
<emphasis bold="true" box="[839,1044,865,886]" pageId="70" pageNumber="197">Material examined.</emphasis>
|
||
<materialsCitation collectingDate="2000-02-16" collectionCode="MZUSP" country="Brazil" county="Nova Mutum" latitude="-13.864445" location="Fazenda" longLatPrecision="21" longitude="-56.193333" municipality="Fazenda Buriti" pageId="70" pageNumber="197" specimenCode="MZUSP 61091, 1" specimenCount="6" stateProvince="Mato Grosso" typeStatus="holotype">
|
||
<specimenCount box="[1053,1174,865,886]" pageId="70" pageNumber="197" type="generic">6 specimens</specimenCount>
|
||
(
|
||
<quantity box="[1191,1321,865,886]" metricMagnitude="-2" metricUnit="m" metricValue="4.0725" metricValueMax="7.5" metricValueMin="0.645" pageId="70" pageNumber="197" unit="mm" value="40.725" valueMax="75.0" valueMin="6.45">6, 45-75 mm</quantity>
|
||
SL).
|
||
<emphasis bold="true" box="[1380,1475,865,886]" pageId="70" pageNumber="197">
|
||
<typeStatus box="[1380,1475,865,886]" pageId="70" pageNumber="197">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<collectingCountry box="[839,902,893,914]" name="Brazil" pageId="70" pageNumber="197">Brazil</collectingCountry>
|
||
.
|
||
<collectingRegion box="[914,1048,893,914]" country="Brazil" name="Mato Grosso" pageId="70" pageNumber="197">Mato Grosso</collectingRegion>
|
||
:
|
||
<collectingCounty box="[1061,1197,893,914]" pageId="70" pageNumber="197">Nova Mutum</collectingCounty>
|
||
, unnamed córrego,
|
||
<collectingMunicipality pageId="70" pageNumber="197">
|
||
<location LSID="urn:lsid:plazi:treatment:E56BC71F0E73FFE63E76FAB4FBF4E6C3:681D20D20E71FFE63A34FCD5FA07E1C7" box="[1400,1481,893,914]" country="Brazil" county="Nova Mutum" latitude="-13.864445" longLatPrecision="21" longitude="-56.193333" municipality="Fazenda Buriti" name="Fazenda" pageId="70" pageNumber="197" stateProvince="Mato Grosso">Fazenda</location>
|
||
Buriti
|
||
</collectingMunicipality>
|
||
, approximately 1.5 km
|
||
<locationDeviation box="[1179,1327,922,943]" location="Fazenda" pageId="70" pageNumber="197">S of Fazenda</locationDeviation>
|
||
(
|
||
<geoCoordinate box="[1347,1476,922,943]" degrees="13" direction="south" minutes="51" orientation="latitude" pageId="70" pageNumber="197" precision="15" seconds="52" value="-13.864445">13°51’52"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[839,965,951,972]" degrees="56" direction="west" minutes="11" orientation="longitude" pageId="70" pageNumber="197" precision="15" seconds="36" value="-56.193333">56°11’36"W</geoCoordinate>
|
||
),
|
||
<collectionCode box="[986,1067,951,972]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="70" pageNumber="197">MZUSP</collectionCode>
|
||
<location LSID="urn:lsid:plazi:treatment:E56BC71F0E73FFE63E76FAB4FBF4E6C3:681D20D20E71FFE63B7FFC1FFB67E199" box="[1075,1193,951,972]" country="Brazil" county="Nova Mutum" latitude="-13.864445" longLatPrecision="21" longitude="-56.193333" municipality="Fazenda Buriti" name="Ichthyology" pageId="70" pageNumber="197" stateProvince="Mato Grosso">Ichthyology</location>
|
||
students,
|
||
<date box="[1297,1477,951,972]" pageId="70" pageNumber="197" value="2000-02-16">
|
||
<collectingDate box="[1297,1477,951,972]" pageId="70" pageNumber="197" value="2000-02-16">16 February 2000</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCode box="[839,990,980,1001]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="70" pageNumber="197">MZUSP 61091</specimenCode>
|
||
,
|
||
<specimenCode box="[1001,1013,980,1001]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="70" pageNumber="197">1</specimenCode>
|
||
, mature male (64)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation collectingDate="2000-02-20" collectionCode="MZUSP" country="Brazil" county="Nova Mutum" latitude="-13.864445" location="Fazenda" longLatPrecision="21" longitude="-56.193333" municipality="Fazenda Buriti" pageId="70" pageNumber="197" specimenCode="MZUSP 83227, 3, USNM 372766, 1, MZUSP 61051" specimenCount="5" stateProvince="Mato Grosso" typeStatus="paratype">
|
||
<emphasis bold="true" box="[1209,1313,980,1001]" pageId="70" pageNumber="197">
|
||
<typeStatus box="[1209,1313,980,1001]" pageId="70" pageNumber="197">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<specimenCount box="[1327,1448,980,1001]" pageId="70" pageNumber="197" type="generic">5 specimens</specimenCount>
|
||
(
|
||
<quantity metricMagnitude="-2" metricUnit="m" metricValue="4.0225" metricValueMax="7.5" metricValueMin="0.545" pageId="70" pageNumber="197" unit="mm" value="40.225" valueMax="75.0" valueMin="5.45">5, 45-75 mm</quantity>
|
||
SL).
|
||
<collectingCountry box="[1008,1073,1009,1030]" name="Brazil" pageId="70" pageNumber="197">Brazil</collectingCountry>
|
||
.
|
||
<collectingRegion box="[1086,1224,1009,1030]" country="Brazil" name="Mato Grosso" pageId="70" pageNumber="197">Mato Grosso</collectingRegion>
|
||
:
|
||
<collectingCounty box="[1237,1377,1009,1030]" pageId="70" pageNumber="197">Nova Mutum</collectingCounty>
|
||
, unnamed córrego,
|
||
<collectingMunicipality box="[926,1073,1037,1058]" pageId="70" pageNumber="197">
|
||
<location LSID="urn:lsid:plazi:treatment:E56BC71F0E73FFE63E76FAB4FBF4E6C3:681D20D20E71FFE63CD2FBA5FC21E677" box="[926,1007,1037,1058]" country="Brazil" county="Nova Mutum" municipality="Fazenda Buriti" name="Fazenda" pageId="70" pageNumber="197" stateProvince="Mato Grosso">Fazenda</location>
|
||
Buriti
|
||
</collectingMunicipality>
|
||
, approximately 1.5 km
|
||
<locationDeviation box="[1304,1434,1037,1058]" location="Fazenda" pageId="70" pageNumber="197">S of Fazenda</locationDeviation>
|
||
, collected with holotype,
|
||
<specimenCode box="[1054,1206,1066,1087]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="70" pageNumber="197">MZUSP 83227</specimenCode>
|
||
,
|
||
<specimenCode box="[1217,1229,1066,1087]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="70" pageNumber="197">3</specimenCode>
|
||
(45-75);
|
||
<specimenCode box="[1324,1478,1066,1087]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="70" pageNumber="197" type="Museum">USNM 372766</specimenCode>
|
||
,
|
||
<specimenCode box="[839,851,1095,1116]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="70" pageNumber="197" type="Museum">1</specimenCode>
|
||
(63).
|
||
<location LSID="urn:lsid:plazi:treatment:E56BC71F0E73FFE63E76FAB4FBF4E6C3:681D20D20E71FFE63CC7FBEFFBC0E609" box="[907,1038,1095,1116]" country="Brazil" county="Nova Mutum" municipality="Fazenda Buriti" name="Nova Mutum" pageId="70" pageNumber="197" stateProvince="Mato Grosso">Nova Mutum</location>
|
||
, riacho tributary to
|
||
<location LSID="urn:lsid:plazi:treatment:E56BC71F0E73FFE63E76FAB4FBF4E6C3:681D20D20E71FFE63B85FBEFFAE2E609" box="[1225,1324,1095,1116]" country="Brazil" county="Nova Mutum" municipality="Fazenda Buriti" name="rio Arinos" pageId="70" pageNumber="197" stateProvince="Mato Grosso">rio Arinos</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:E56BC71F0E73FFE63E76FAB4FBF4E6C3:681D20D20E71FFE63A78FBEFFA0BE609" box="[1332,1477,1095,1116]" country="Brazil" county="Nova Mutum" municipality="Fazenda Buriti" name="Fazenda Buriti" pageId="70" pageNumber="197" stateProvince="Mato Grosso">
|
||
<location LSID="urn:lsid:plazi:treatment:E56BC71F0E73FFE63E76FAB4FBF4E6C3:681D20D20E71FFE63A78FBEFFA4BE609" box="[1332,1413,1095,1116]" country="Brazil" county="Nova Mutum" municipality="Fazenda Buriti" name="Fazenda" pageId="70" pageNumber="197" stateProvince="Mato Grosso">Fazenda</location>
|
||
Buriti
|
||
</location>
|
||
,
|
||
<collectionCode box="[839,920,1124,1145]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="70" pageNumber="197">MZUSP</collectionCode>
|
||
<location LSID="urn:lsid:plazi:treatment:E56BC71F0E73FFE63E76FAB4FBF4E6C3:681D20D20E71FFE63CEEFBCCFBD7E62C" box="[930,1049,1124,1145]" country="Brazil" county="Nova Mutum" municipality="Fazenda Buriti" name="Ichthyology" pageId="70" pageNumber="197" stateProvince="Mato Grosso">Ichthyology</location>
|
||
students,
|
||
<date box="[1156,1288,1124,1145]" pageId="70" pageNumber="197" value="2000-02-20">
|
||
<collectingDate box="[1156,1288,1124,1145]" pageId="70" pageNumber="197" value="2000-02-20">20 Feb 2000</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCode box="[1301,1456,1124,1145]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="70" pageNumber="197">MZUSP 61051</specimenCode>
|
||
,
|
||
<specimenCode box="[1469,1481,1124,1145]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="70" pageNumber="197">1</specimenCode>
|
||
(46, cleared and stained)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |