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<document ID-DOI="10.1590/S1679-62252005000200001" ID-ISSN="1982-0224" ID-Zenodo-Dep="5418569" approvalRequired="104" approvalRequired_for_taxonomicNames="6" approvalRequired_for_textStreams="91" approvalRequired_for_treatments="7" checkinTime="1630369289621" checkinUser="carolina" docAuthor="Vari, Richard P., Ferraris Jr, Carl J. &amp; de Pinna, Mário C. C." docDate="2005" docId="E56BC71F0E31FFA93C85FBD5FF00E42C" docLanguage="en" docName="NeotropIchthyol.2005.8.pdf" docOrigin="Neotropical Ichthyology 3 (2)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252005000200001&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Cetopsidium ferreirai Vari &amp; Ferraris Jr &amp; de Pinna 2005, new species" docType="treatment" docVersion="2" lastPageId="9" lastPageNumber="136" masterDocId="1952BF670E37FFA03F4CFFA8FFCEE255" masterDocTitle="The Neotropical whale catfishes (Siluriformes: Cetopsidae: Cetopsinae), a revisionary study" masterLastPageNumber="238" masterPageNumber="127" pageId="6" pageNumber="133" updateTime="1630681654510" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The Neotropical whale catfishes (Siluriformes: Cetopsidae: Cetopsinae), a revisionary study</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vari, Richard P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ferraris Jr, Carl J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>de Pinna, Mário C. C.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2005</mods:date>
<mods:detail type="pubDate">
<mods:number>2005-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>127</mods:start>
<mods:end>238</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/S1679-62252005000200001</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">5418569</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:E56BC71F0E31FFA93C85FBD5FF00E42C" httpUri="http://treatment.plazi.org/id/E56BC71F0E31FFA93C85FBD5FF00E42C" lastPageId="9" lastPageNumber="136" pageId="6" pageNumber="133">
<subSubSection box="[969,1351,1149,1173]" pageId="6" pageNumber="133" type="nomenclature">
<paragraph blockId="6.[969,1351,1149,1205]" box="[969,1351,1149,1173]" pageId="6" pageNumber="133">
<heading box="[969,1351,1149,1173]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="133" reason="2">
<emphasis bold="true" box="[969,1351,1149,1173]" pageId="6" pageNumber="133">
<emphasis bold="true" box="[969,1214,1149,1173]" italics="true" pageId="6" pageNumber="133">
<taxonomicName authority="Vari &amp; Ferraris Jr &amp; de Pinna, 2005" authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[969,1209,1149,1173]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="133" phylum="Chordata" rank="species" species="ferreirai" status="new species">Cetopsidium ferreirai</taxonomicName>
,
</emphasis>
<taxonomicNameLabel box="[1220,1351,1150,1173]" pageId="6" pageNumber="133" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1054,1266,1182,1205]" pageId="6" pageNumber="133" type="description">
<paragraph blockId="6.[969,1351,1149,1205]" box="[1054,1266,1182,1205]" pageId="6" pageNumber="133">
<figureCitation box="[1054,1148,1182,1205]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[151,190,538,561]" captionStartId-1="8.[839,878,886,909]" captionTargetBox-0="[247,1384,228,497]" captionTargetBox-1="[839,1474,190,852]" captionTargetId-0="figure-738@7.[215,1425,193,525]" captionTargetId-1="figure-687@8.[839,1480,187,852]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 1. Cetopsidium ferreirai, new species, holotype, male, INPA 10194, 25 mm SL; Brazil, Pará, rio Trombetas basin, rio Mapuera, last cachoeira before the rio Trombetas (approximately 1°05S, 57°01&quot;W)." captionText-1="Fig. 2. Map of central and northern South America showing geographic distribution of Cetopsidium ferreirai (1 = holotype locality), Cetopsidium minutum (2 = holotype locality of Hemicetopsis minutus), Cetopsidium morenoi, Cetopsidium orientale (3 = holotype locality of Pseudocetopsis orientale), Cetopsidium pemon (4 = holotype locality), and Cetopsidium roae (5 = holotype locality) (some symbols represent more that one locality and/or lot of specimens)." figureDoi-0="http://doi.org/10.5281/zenodo.5418573" figureDoi-1="http://doi.org/10.5281/zenodo.5418575" httpUri-0="https://zenodo.org/record/5418573/files/figure.png" httpUri-1="https://zenodo.org/record/5418575/files/figure.png" pageId="6" pageNumber="133">Figs. 1-2</figureCitation>
,
<tableCitation box="[1157,1266,1182,1205]" captionStart-0="Table 2" captionStart-1="Table 3" captionStart-2="Table 4" captionStart-3="Table 5" captionStart-4="Table 6" captionStart-5="Table 7" captionStart-6="Table 8" captionStartId-0="22.[158,221,212,235]" captionStartId-1="22.[158,221,567,590]" captionStartId-2="22.[158,221,925,948]" captionStartId-3="22.[158,221,1285,1308]" captionStartId-4="22.[158,221,1641,1664]" captionStartId-5="22.[839,902,212,235]" captionStartId-6="22.[839,901,569,592]" captionText-0="Table 2. Number of specimens of the species of Cetopsidium with cited number of dorsal andpectoral-fin rays. Bold indicates value for holotype when examined." captionText-1="Table 3. Number of specimens of the species of Cetopsidium with cited number of branched anal-fin rays. Bold indicates value for holotype when examined." captionText-2="Table 4. Number of specimens of the species of Cetopsidium with cited number of total anal-fin rays. Bold indicates value for holotype when examined." captionText-3="Table 5. Number of specimens of the species of Cetopsidium with cited number of preanal vertebrae. Bold indicates values for holotype when examined." captionText-4="Table 6. Number of species of the specimens of Cetopsidium with cited number of precaudal vertebrae.Bold indicates values for holotype when examined." captionText-5="Table 7. Number of specimens of the species of Cetopsidium with cited number of caudal vertebrae. Bold indicates value for holotype when examined." captionText-6="Table 8. Numbers of specimens of the species of Cetopsidium with cited number of total vertebrae. Bold indicates value for holotype when examined" pageId="6" pageNumber="133">Tables 2-8</tableCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="133" type="reference_group">
<paragraph blockId="6.[839,1481,1250,1306]" pageId="6" pageNumber="133">
<treatmentCitationGroup pageId="6" pageNumber="133">
<taxonomicName baseAuthorityName="Eigenmann" baseAuthorityYear="1912" box="[839,1086,1250,1275]" class="Actinopterygii" family="Cetopsidae" genus="Pseudocetopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="133" phylum="Chordata" rank="species" species="minuta">
<emphasis box="[839,1086,1250,1275]" italics="true" pageId="6" pageNumber="133">Pseudocetopsis minuta</emphasis>
</taxonomicName>
[not of Eigenmann, 1912].
<treatmentCitation author="Ferreira, E" page="52" pageId="6" pageNumber="133" year="1995">
<bibRefCitation author="Ferreira, E" pageId="6" pageNumber="133" pagination="1 - 4" refId="ref108846" refString="Ferreira, E. 1995. Composicao, distribuicao e aspectos ecologicos da ictiofauna de um trecho do rio Trombetas, na area de influencia da futura UHE Cachoeira Porteira, Estado do Para, Brasil. ActaAmazonica, 23 (1 - 4, Suplemento): 1 - 88. [issue for 1993, actual publication date 1995]" type="journal article" year="1995">Ferreira, 1995: 52</bibRefCitation>
</treatmentCitation>
, table 3 [
<collectingCountry box="[1058,1128,1283,1306]" name="Brazil" pageId="6" pageNumber="133">Brazil</collectingCountry>
: Pará, rio Trombetas basin].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="134" pageId="6" pageNumber="133" type="diagnosis">
<paragraph blockId="6.[839,1482,1346,1978]" lastBlockId="7.[151,794,642,1241]" lastPageId="7" lastPageNumber="134" pageId="6" pageNumber="133">
<emphasis bold="true" box="[839,957,1347,1370]" pageId="6" pageNumber="133">Diagnosis.</emphasis>
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[963,1198,1346,1370]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="133" phylum="Chordata" rank="species" species="ferreirai">
<emphasis box="[963,1198,1346,1370]" italics="true" pageId="6" pageNumber="133">Cetopsidium ferreirai</emphasis>
</taxonomicName>
differs from all of its congeners with the possible exception of
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[1232,1310,1379,1403]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="133" phylum="Chordata" rank="species" species="roae">
<emphasis box="[1232,1310,1379,1403]" italics="true" pageId="6" pageNumber="133">C. roae</emphasis>
</taxonomicName>
and
<taxonomicName box="[1359,1481,1379,1403]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="133" phylum="Chordata" rank="species" species="minutum">
<emphasis box="[1359,1481,1379,1403]" italics="true" pageId="6" pageNumber="133">C. minutum</emphasis>
</taxonomicName>
(see those species accounts) in the lack of distal filaments on the first rays of the dorsal and pectoral fins in adult males (
<emphasis box="[848,917,1476,1499]" italics="true" pageId="6" pageNumber="133">versus</emphasis>
the presence of such filaments in adult males as indicated by the possession of distinctly convex anal-fin margins, respectively).
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[1044,1273,1538,1562]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="133" phylum="Chordata" rank="species" species="ferreirai">
<emphasis box="[1044,1273,1538,1562]" italics="true" pageId="6" pageNumber="133">Cetopsidium ferreirai</emphasis>
</taxonomicName>
further differs from
<taxonomicName box="[839,966,1571,1595]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="133" phylum="Chordata" rank="species" species="minutum">
<emphasis box="[839,966,1571,1595]" italics="true" pageId="6" pageNumber="133">C. minutum</emphasis>
</taxonomicName>
in the length of the pelvic fin (falling short of anal-fin origin
<emphasis box="[997,1065,1604,1627]" italics="true" pageId="6" pageNumber="133">versus</emphasis>
extending at least to anal-fin origin, respectively).
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[976,1217,1634,1658]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="133" phylum="Chordata" rank="species" species="ferreirai">
<emphasis box="[976,1217,1634,1658]" italics="true" pageId="6" pageNumber="133">Cetopsidium ferreirai</emphasis>
</taxonomicName>
further differs from
<taxonomicName baseAuthorityName="Fernandez-Yepez" baseAuthorityYear="1972" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="133" phylum="Chordata" rank="species" species="morenoi">
<emphasis italics="true" pageId="6" pageNumber="133">C. morenoi</emphasis>
</taxonomicName>
in the overall form of the head (the head being not massive proportionally as a consequence of the less extensive development of the adductor mandibulae muscle onto the dorsal surface of head and with the snout distinctly rounded
<emphasis box="[934,1003,1796,1819]" italics="true" pageId="6" pageNumber="133">versus</emphasis>
the head being proportionally massive with well-developed, distinctly bulging adductor mandibulae muscles and the snout less rounded, respectively; compare
<figureCitation box="[839,900,1891,1914]" captionStart="Fig" captionStartId="7.[151,190,538,561]" captionTargetBox="[247,1384,228,497]" captionTargetId="figure-738@7.[215,1425,193,525]" captionTargetPageId="7" captionText="Fig. 1. Cetopsidium ferreirai, new species, holotype, male, INPA 10194, 25 mm SL; Brazil, Pará, rio Trombetas basin, rio Mapuera, last cachoeira before the rio Trombetas (approximately 1°05S, 57°01&quot;W)." figureDoi="http://doi.org/10.5281/zenodo.5418573" httpUri="https://zenodo.org/record/5418573/files/figure.png" pageId="6" pageNumber="133">Fig. 1</figureCitation>
with
<figureCitation box="[960,1034,1891,1914]" captionStart="Fig" captionStartId="12.[158,196,1002,1025]" captionTargetBox="[254,1395,677,963]" captionTargetId="figure-39@12.[217,1427,646,986]" captionTargetPageId="12" captionText="Fig. 5. Cetopsidium morenoi, INHS 34506, 20.5 mm SL; Venezuela, Guarico, río San Bartolo (8o03N, 66o42W)." figureDoi="http://doi.org/10.5281/zenodo.5418584" httpUri="https://zenodo.org/record/5418584/files/figure.png" pageId="6" pageNumber="133">Figs. 5</figureCitation>
,
<figureCitation box="[1045,1060,1891,1914]" captionStart="Fig" captionStartId="12.[158,197,1400,1423]" captionTargetBox="[257,1382,1055,1361]" captionTargetId="figure-61@12.[217,1427,1055,1379]" captionTargetPageId="12" captionText="Fig. 6. Cetopsidium morenoi, mature male, ANSP 165734, 43 mm SL; Venezuela,Apure, Caño Potrerito of río Cinaruco, 24 km S of río Cinaruco on San Fernando de Apure to Puerto Paez Highway (6°25N, 67°32W)." figureDoi="http://doi.org/10.5281/zenodo.5418586" httpUri="https://zenodo.org/record/5418586/files/figure.png" pageId="6" pageNumber="133">6</figureCitation>
), and the less extensive dark pigmentation on the body (the near absence of dark pigmentation on the lateral surface of body and the pigmentation limited to the middorsal region of body
<emphasis box="[423,491,642,665]" italics="true" pageId="7" pageNumber="134">versus</emphasis>
the dark pigmentation being distributed over much of the lateral and dorsal surfaces of the body, respectively).
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[361,588,705,729]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="134" phylum="Chordata" rank="species" species="ferreirai">
<emphasis box="[361,588,705,729]" italics="true" pageId="7" pageNumber="134">Cetopsidium ferreirai</emphasis>
</taxonomicName>
further differs from
<taxonomicName authorityName="Vari, Ferraris, &amp; Keith" authorityYear="2003" box="[151,280,737,761]" class="Actinopterygii" family="Cetopsidae" genus="Pseudocetopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="134" phylum="Chordata" rank="species" species="orientale">
<emphasis box="[151,280,737,761]" italics="true" pageId="7" pageNumber="134">C. orientale</emphasis>
</taxonomicName>
in the alignment of the dorsal and ventral profiles of the portion of the body posterior of the base of the dorsal fin (profiles converging posteriorly
<emphasis box="[607,676,802,825]" italics="true" pageId="7" pageNumber="134">versus</emphasis>
running in parallel, respectively) and in the overall coloration (limited dark pigmentation located largely along the middorsal region of the body
<emphasis box="[275,343,898,921]" italics="true" pageId="7" pageNumber="134">versus</emphasis>
dark pigmentation widely distributed over the dorsal and lateral portions of body, respectively).
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[151,388,961,985]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="134" phylum="Chordata" rank="species" species="ferreirai">
<emphasis box="[151,388,961,985]" italics="true" pageId="7" pageNumber="134">Cetopsidium ferreirai</emphasis>
</taxonomicName>
further differs from
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[617,723,962,985]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="134" phylum="Chordata" rank="species" species="pemon">
<emphasis box="[617,723,962,985]" italics="true" pageId="7" pageNumber="134">C. pemon</emphasis>
</taxonomicName>
in the overall form of the head (profile of head anteriorly rounded
<emphasis box="[151,220,1026,1049]" italics="true" pageId="7" pageNumber="134">versus</emphasis>
distinctly pointed, respectively) and in the proportional length of the snout (approximately 2 times diameter of the eye
<emphasis box="[239,308,1090,1113]" italics="true" pageId="7" pageNumber="134">versus</emphasis>
approximately 3 times diameter of the eye, respectively).
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[305,542,1121,1145]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="134" phylum="Chordata" rank="species" species="ferreirai">
<emphasis box="[305,542,1121,1145]" italics="true" pageId="7" pageNumber="134">Cetopsidium ferreirai</emphasis>
</taxonomicName>
further differs from
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="134" phylum="Chordata" rank="species" species="roae">
<emphasis italics="true" pageId="7" pageNumber="134">C. roae</emphasis>
</taxonomicName>
in the position of the vent (proximate to the base of the anterior most anal-fin ray
<emphasis box="[422,490,1186,1209]" italics="true" pageId="7" pageNumber="134">versus</emphasis>
distinctly separated from the base of the anterior most anal-fin ray, respectively).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5418573" ID-Zenodo-Dep="5418573" httpUri="https://zenodo.org/record/5418573/files/figure.png" pageId="7" pageNumber="134" startId="7.[151,190,538,561]" targetBox="[247,1384,228,497]" targetPageId="7">
<paragraph blockId="7.[151,1474,537,593]" pageId="7" pageNumber="134">
<emphasis bold="true" box="[151,222,538,561]" pageId="7" pageNumber="134">Fig. 1.</emphasis>
<taxonomicName authority="Vari &amp; Ferraris Jr &amp; de Pinna, 2005" authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[230,464,537,561]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="134" phylum="Chordata" rank="species" species="ferreirai" status="new species">
<emphasis box="[230,464,537,561]" italics="true" pageId="7" pageNumber="134">Cetopsidium ferreirai</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[478,611,538,561]" pageId="7" pageNumber="134" rank="species">new species</taxonomicNameLabel>
, holotype, male, INPA 10194, 25 mm SL; Brazil, Pará, rio Trombetas basin, rio Mapuera, last cachoeira before the rio Trombetas (approximately 1°05S, 57°01&quot;W).
</paragraph>
</caption>
<subSubSection lastPageId="8" lastPageNumber="135" pageId="7" pageNumber="134" type="description">
<paragraph blockId="7.[151,794,1282,1977]" pageId="7" pageNumber="134">
<emphasis bold="true" box="[151,291,1282,1305]" pageId="7" pageNumber="134">Description.</emphasis>
Body elongate, barely compressed laterally anteriorly and becoming progressively distinctly-compressed posteriorly. Body depth at dorsal-fin origin approximately 0.18- 0.20 of SL, and slightly less than HL. Lateral line on body incomplete, unbranched, and midlateral; extending posteriorly to over posterior portion of base of anal fin but falling short of caudal peduncle. Dorsal profile of body slightly convex from nape to dorsal-fin origin and straight from dorsal-fin origin to caudal-fin base. Ventral profile of body slightly convex along abdomen, approximately straight, and nearly running in parallel to dorsal profile along region of caudal peduncle. Caudal-peduncle depth approximately equal to caudal-peduncle length.
</paragraph>
<paragraph blockId="7.[151,794,1282,1977]" pageId="7" pageNumber="134">Head in profile acutely triangular overall with bluntlypointed snout. Dorsal profile of head convex from tip of snout to vertical through posterior margin of orbit and nearly straight from that point to nape. Ventral profile of head slightly convex. Margin of snout rounded from dorsal view. Lateral margins of postorbital portion of head slightly convex on each side from dorsal view. Enlarged jaw musculature very evident externally on dorsal and lateral surfaces of postorbital portion of head.</paragraph>
<paragraph blockId="7.[831,1474,642,1977]" pageId="7" pageNumber="134">Opercular membrane attaching to isthmus only anterior of vertical through pectoral-fin insertion. Opercular opening moderately-elongate; extending anteroventral of pectoral-fin insertion by distance approximately equal to one-third of HL and extending dorsal of pectoral-fin insertion by distance equal to diameter of eye.</paragraph>
<paragraph blockId="7.[831,1474,642,1977]" pageId="7" pageNumber="134">Eye situated on lateral surface of head; located entirely dorsal to horizontal through pectoral-fin insertion; eye visible in dorsal view, but not in ventral view, of head. Middle of orbit at approximately anterior one-third of HL. Eye diameter slightly more than one-half of snout length. Interorbital width approximately equal to distance from tip of snout to middle of orbit. Anterior narial opening circular, surrounded by short, anteriorly-directed, tubular rim of skin. Opening of anterior nares located slightly dorsal of horizontal extending through maxillary-barbel origin and at, or slightly ventral of, horizontal extending through tip of snout. Distance between anterior nares approximately equal to distance from tip of snout to posterior margin of orbit. Posterior narial opening located on dorsal surface of head, situated dorsal to anterior one-third of orbit; narial opening nearly round and with flap of skin extending along anterior one-half of rim of aperture, with flap highest anteriorly.</paragraph>
<paragraph blockId="7.[831,1474,642,1977]" pageId="7" pageNumber="134">Mouth subterminal, its width approximately two-fifths of HL. Margin of lower jaw gently rounded, its posterior limit reaching to vertical through middle of orbit. Premaxillary tooth patch in form of gently-arched band continuous across midline and with anterior margin convex and posterior margin concave and running in parallel to anterior margin. Teeth on premaxilla small, conical, and sharply-pointed, with teeth arranged in two, more or less regular, rows of uniform-sized teeth medially and reduced to one row at lateral limit of premaxilla. Vomerine teeth arranged in single, arched row with distinct gap in series at midline. Vomerine teeth stout and conical, and either of uniform size or becoming progressively slightly larger laterally, but with even smallest teeth in series larger than teeth on premaxilla. Dentary teeth comparable in shape to, but larger in size than, premaxillary teeth and consisting of two or three irregular rows medially that taper to one row laterally.</paragraph>
<paragraph blockId="7.[831,1474,642,1977]" lastBlockId="8.[158,802,194,1593]" lastPageId="8" lastPageNumber="135" pageId="7" pageNumber="134">Maxillary barbel slender, its length slightly greater than distance from anterior margin of orbit to rear of head and approximately three-fourths of HL; barbel origin located ventral to anterior margin of orbit. Medial mental barbel approximately equal in length to lateral mental barbel, with latter slightly shorter than maxillary barbel. Medial mental-barbel origin located along vertical through rictus. Lateral mentalbarbel origin situated slightly posterior of vertical through medial mental-barbel origin. Tips of adpressed mental barbels extending slightly beyond posterior margin of opercle.</paragraph>
<paragraph blockId="8.[158,802,194,1593]" pageId="8" pageNumber="135">Dorsal fin moderately large overall with length of dorsalfin base approximately 0.33-0.35 of HL. Length of longest branched dorsal-fin ray equal to approximately two-thirds of HL. Dorsal-fin spinelet present. First dorsal-fin ray spinous for basal one-half of length and flexible more distally. First dorsal-fin ray lacking distal filament in all examined specimens including mature males as evidenced by presence of distinctly convex anal-fin margin. Distal margin of dorsal fin convex, with second branched fin ray longest. Dorsal-fin origin located at approximately anterior 0.29-0.30 of SL and along vertical extending through posterior one-half of adpressed pectoral fin. Tip of adpressed dorsal fin extending to vertical through vent. Posterior most dorsal-fin ray with slight basal, posterior, membranous attachment to body.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5418575" ID-Zenodo-Dep="5418575" httpUri="https://zenodo.org/record/5418575/files/figure.png" pageId="8" pageNumber="135" startId="8.[839,878,886,909]" targetBox="[839,1474,190,852]" targetPageId="8">
<paragraph blockId="8.[839,1481,886,1133]" pageId="8" pageNumber="135">
<emphasis bold="true" box="[839,909,886,909]" pageId="8" pageNumber="135">Fig. 2.</emphasis>
Map of central and northern South America showing geographic distribution of
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[1128,1364,917,941]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="135" phylum="Chordata" rank="species" species="ferreirai">
<emphasis box="[1128,1364,917,941]" italics="true" pageId="8" pageNumber="135">Cetopsidium ferreirai</emphasis>
</taxonomicName>
(1 = holotype locality),
<taxonomicName box="[991,1223,949,973]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="135" phylum="Chordata" rank="species" species="minutum">
<emphasis box="[991,1223,949,973]" italics="true" pageId="8" pageNumber="135">Cetopsidium minutum</emphasis>
</taxonomicName>
(2 = holotype locality of
<taxonomicName authorityName="Eigenmann" authorityYear="1912" box="[839,1076,982,1005]" class="Actinopterygii" family="Cetopsidae" genus="Hemicetopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="135" phylum="Chordata" rank="species" species="minutus">
<emphasis box="[839,1076,982,1005]" italics="true" pageId="8" pageNumber="135">Hemicetopsis minutus</emphasis>
</taxonomicName>
),
<taxonomicName baseAuthorityName="Fernandez-Yepez" baseAuthorityYear="1972" box="[1099,1332,981,1005]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="135" phylum="Chordata" rank="species" species="morenoi">
<emphasis box="[1099,1332,981,1005]" italics="true" pageId="8" pageNumber="135">Cetopsidium morenoi</emphasis>
</taxonomicName>
,
<taxonomicName class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="135" phylum="Chordata" rank="species" species="orientale">
<emphasis italics="true" pageId="8" pageNumber="135">Cetopsidium orientale</emphasis>
</taxonomicName>
(3 = holotype locality of
<taxonomicName authorityName="Vari, Ferraris, &amp; Keith" authorityYear="2003" box="[1199,1466,1013,1037]" class="Actinopterygii" family="Cetopsidae" genus="Pseudocetopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="135" phylum="Chordata" rank="species" species="orientale">
<emphasis box="[1199,1466,1013,1037]" italics="true" pageId="8" pageNumber="135">Pseudocetopsis orientale</emphasis>
</taxonomicName>
),
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[839,1050,1045,1069]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="135" phylum="Chordata" rank="species" species="pemon">
<emphasis box="[839,1050,1045,1069]" italics="true" pageId="8" pageNumber="135">Cetopsidium pemon</emphasis>
</taxonomicName>
(4 = holotype locality), and
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="135" phylum="Chordata" rank="species" species="roae">
<emphasis italics="true" pageId="8" pageNumber="135">Cetopsidium roae</emphasis>
</taxonomicName>
(5 = holotype locality) (some symbols represent more that one locality and/or lot of specimens).
</paragraph>
</caption>
<paragraph blockId="8.[158,802,194,1593]" pageId="8" pageNumber="135">Caudal fin deeply-forked, symmetrical; tips of lobes bluntly pointed. Length of longest caudal-fin rays approximately two times length of middle fin rays.</paragraph>
<paragraph blockId="8.[158,802,194,1593]" pageId="8" pageNumber="135">
Base of anal fin moderately long. Anal-fin origin located posterior of middle of SL but falling short of middle of TL. Anal-fin margin nearly straight in apparent females, and broadly convex in apparent mature males (see
<figureCitation box="[564,624,1090,1113]" captionStart="Fig" captionStartId="7.[151,190,538,561]" captionTargetBox="[247,1384,228,497]" captionTargetId="figure-738@7.[215,1425,193,525]" captionTargetPageId="7" captionText="Fig. 1. Cetopsidium ferreirai, new species, holotype, male, INPA 10194, 25 mm SL; Brazil, Pará, rio Trombetas basin, rio Mapuera, last cachoeira before the rio Trombetas (approximately 1°05S, 57°01&quot;W)." figureDoi="http://doi.org/10.5281/zenodo.5418573" httpUri="https://zenodo.org/record/5418573/files/figure.png" pageId="8" pageNumber="135">Fig. 1</figureCitation>
). Posterior most anal-fin ray with slight membranous attachment to body.
</paragraph>
<paragraph blockId="8.[158,802,194,1593]" pageId="8" pageNumber="135">Pelvic fin small; distal margin slightly convex with middle fin rays longest. Pelvic-fin insertion located anterior to middle of SL and along vertical through posterior terminus of base of dorsal fin. Tip of adpressed pelvic fin extending past middle of SL and reaching anterior margin of vent. Medial most pelvic-fin ray with membranous attachment to body along basal one-half of its length.</paragraph>
<paragraph blockId="8.[158,802,194,1593]" pageId="8" pageNumber="135">Pectoral-fin length approximately two-thirds of HL. Pectoral-fin margin distinctly convex with middle ray longest. First pectoral-fin ray spinous with smooth margins; spinous portion of ray short, with length slightly more than one-half that of first branched ray and not prolonged as distal filament in any of examined specimens including apparently mature males (as indicated by broadly convex anal-fin margin).</paragraph>
<paragraph blockId="8.[158,802,1634,1977]" pageId="8" pageNumber="135">
<emphasis bold="true" box="[158,404,1634,1657]" pageId="8" pageNumber="135">Coloration in alcohol.</emphasis>
All available specimens with most of dark pigmentation apparently lost as consequence of postmortem bleaching. Overall coloration of head and body pale. Abdomen pale. Body with small, scattered, rounded, pale brown spots on dorsal and lateral surfaces in some specimens. Dorsal portion of head pale or with few scattered dark spots dorsally and around orbit. Lateral and ventral surfaces of head otherwise pale. Fine stripe of brown spots distributed along middorsal portion of body in
<specimenCount box="[589,730,1890,1913]" pageId="8" pageNumber="135" type="generic" typeStatus="paratype">one paratype</specimenCount>
. Other specimens with overall pigmentation very faint and middorsal stripe not apparent.
</paragraph>
<paragraph blockId="8.[839,1482,1186,1337]" pageId="8" pageNumber="135">Dorsal fin pale or with few, scattered, dark spots at base. Anal fin pale or with fine, very widely scattered, dark pigment spots basally. Caudal fin with few, scattered, dark spots basally with remainder of fin pale. Pelvic and pectoral fins pale. All barbels pale.</paragraph>
<paragraph blockId="8.[839,1482,1378,1753]" pageId="8" pageNumber="135">
<emphasis bold="true" box="[839,1070,1378,1401]" pageId="8" pageNumber="135">Sexual dimorphism.</emphasis>
A portion of the examined population samples of
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[967,1206,1409,1433]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="135" phylum="Chordata" rank="species" species="ferreirai">
<emphasis box="[967,1206,1409,1433]" italics="true" pageId="8" pageNumber="135">Cetopsidium ferreirai</emphasis>
</taxonomicName>
are characterized by the presence of an anal-fin margin that is broadly convex in contrast to the straight fin margin present in the remainder of the examined material of the species. Such a broadly convex anal-fin margin typifies mature males of other sexually dimorphic species in the
<taxonomicName authorityName="Bleeker" authorityYear="1858" box="[1098,1216,1570,1593]" class="Actinopterygii" family="Cetopsidae" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="135" phylum="Chordata" rank="subFamily" subFamily="Cetopsinae">Cetopsinae</taxonomicName>
and individuals with that attribute are consequently considered to be mature males. The presumed mature males lack, however, the distal filaments on the first rays of the dorsal and pectoral fins that usually characterize mature males of sexually-dimorphic species within the
<taxonomicName authorityName="Bleeker" authorityYear="1858" box="[880,1004,1730,1753]" class="Actinopterygii" family="Cetopsidae" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="135" phylum="Chordata" rank="subFamily" subFamily="Cetopsinae">Cetopsinae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="135" type="distribution">
<paragraph blockId="8.[839,1481,1793,1881]" pageId="8" pageNumber="135">
<emphasis bold="true" box="[839,987,1794,1817]" pageId="8" pageNumber="135">Distribution.</emphasis>
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[993,1227,1793,1817]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="135" phylum="Chordata" rank="species" species="ferreirai">
<emphasis box="[993,1227,1793,1817]" italics="true" pageId="8" pageNumber="135">Cetopsidium ferreirai</emphasis>
</taxonomicName>
is known only from the rio Trombetas, a left bank tributary of the lower rio
<collectingRegion box="[1369,1481,1826,1849]" country="Venezuela" name="Amazonas" pageId="8" pageNumber="135">Amazonas</collectingRegion>
(
<figureCitation box="[847,911,1858,1881]" captionStart="Fig" captionStartId="8.[839,878,886,909]" captionTargetBox="[839,1474,190,852]" captionTargetId="figure-687@8.[839,1480,187,852]" captionTargetPageId="8" captionText="Fig. 2. Map of central and northern South America showing geographic distribution of Cetopsidium ferreirai (1 = holotype locality), Cetopsidium minutum (2 = holotype locality of Hemicetopsis minutus), Cetopsidium morenoi, Cetopsidium orientale (3 = holotype locality of Pseudocetopsis orientale), Cetopsidium pemon (4 = holotype locality), and Cetopsidium roae (5 = holotype locality) (some symbols represent more that one locality and/or lot of specimens)." figureDoi="http://doi.org/10.5281/zenodo.5418575" httpUri="https://zenodo.org/record/5418575/files/figure.png" pageId="8" pageNumber="135">Fig. 2</figureCitation>
; see also under “Remarks”).
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="136" pageId="8" pageNumber="135" type="etymology">
<paragraph blockId="8.[839,1482,1921,1977]" pageId="8" pageNumber="135">
<emphasis bold="true" box="[839,968,1922,1945]" pageId="8" pageNumber="135">Etymology.</emphasis>
The species name,
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[1170,1260,1921,1945]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="135" phylum="Chordata" rank="species" species="ferreirai">
<emphasis box="[1170,1260,1921,1945]" italics="true" pageId="8" pageNumber="135">ferreirai</emphasis>
</taxonomicName>
, is in honor of Efrem J. G. Ferreira of the Instituto Nacional de Pesquisas da
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5418579" ID-Zenodo-Dep="5418579" httpUri="https://zenodo.org/record/5418579/files/figure.png" pageId="9" pageNumber="136" startId="9.[151,190,583,606]" targetBox="[245,1379,219,557]" targetPageId="9">
<paragraph blockId="9.[151,1474,582,638]" pageId="9" pageNumber="136">
<emphasis bold="true" box="[151,215,583,606]" pageId="9" pageNumber="136">Fig. 3</emphasis>
.
<taxonomicName authority=", ANSP" authorityName="ANSP" box="[229,546,582,606]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="136" phylum="Chordata" rank="species" species="minutum">
<emphasis box="[229,466,582,606]" italics="true" pageId="9" pageNumber="136">Cetopsidium minutum</emphasis>
, ANSP
</taxonomicName>
175839, 21 mm SL; Guyana, Siparuni VIII-2, Essequibo River at Essequibo campsite (4°4541&quot;N, 58°4553&quot;W).
</paragraph>
</caption>
<paragraph blockId="9.[151,794,705,792]" pageId="9" pageNumber="136">Amazônia (INPA), the collector of all the known specimens of the species, in recognition to his contributions to our knowledge of the fishes of the Brazilian Amazon.</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="136" type="discussion">
<paragraph blockId="9.[151,794,833,1336]" pageId="9" pageNumber="136">
<emphasis bold="true" box="[151,267,833,856]" pageId="9" pageNumber="136">Remarks.</emphasis>
In his report on the ichthyofauna of the rio Trombetas basin
<bibRefCitation author="Ferreira, E" box="[331,531,865,888]" pageId="9" pageNumber="136" pagination="1 - 4" refId="ref108846" refString="Ferreira, E. 1995. Composicao, distribuicao e aspectos ecologicos da ictiofauna de um trecho do rio Trombetas, na area de influencia da futura UHE Cachoeira Porteira, Estado do Para, Brasil. ActaAmazonica, 23 (1 - 4, Suplemento): 1 - 88. [issue for 1993, actual publication date 1995]" type="journal article" year="1995">Ferreira (1995: 52)</bibRefCitation>
reported
<taxonomicName baseAuthorityName="Eigenmann" baseAuthorityYear="1912" class="Actinopterygii" family="Cetopsidae" genus="Pseudocetopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="136" phylum="Chordata" rank="species" species="minuta">
<emphasis italics="true" pageId="9" pageNumber="136">Pseudocetopsis minuta</emphasis>
</taxonomicName>
(the
<taxonomicName box="[285,524,896,920]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="136" phylum="Chordata" rank="species" species="minutum">
<emphasis box="[285,524,896,920]" italics="true" pageId="9" pageNumber="136">Cetopsidium minutum</emphasis>
</taxonomicName>
of this study) from four regions within that drainage system. According to our results,
<taxonomicName box="[151,394,960,984]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="136" phylum="Chordata" rank="species" species="minutum">
<emphasis box="[151,394,960,984]" italics="true" pageId="9" pageNumber="136">Cetopsidium minutum</emphasis>
</taxonomicName>
is only known from the
<collectingCountry box="[682,794,961,984]" name="Guyana" pageId="9" pageNumber="136">Essequibo</collectingCountry>
River basin of
<collectingCountry box="[323,410,993,1016]" name="Guyana" pageId="9" pageNumber="136">Guyana</collectingCountry>
, a considerable distance from the localities from which Ferreiras material originated. We have examined specimens from Ferreiras “Região Cachoeira Porteira” (CPT of
<bibRefCitation author="Ferreira, E" box="[366,532,1089,1112]" pageId="9" pageNumber="136" pagination="1 - 4" refId="ref108846" refString="Ferreira, E. 1995. Composicao, distribuicao e aspectos ecologicos da ictiofauna de um trecho do rio Trombetas, na area de influencia da futura UHE Cachoeira Porteira, Estado do Para, Brasil. ActaAmazonica, 23 (1 - 4, Suplemento): 1 - 88. [issue for 1993, actual publication date 1995]" type="journal article" year="1995">Ferreira, 1995</bibRefCitation>
:
<tableCitation box="[546,632,1089,1112]" captionStart="Table 3" captionStartId="22.[158,221,567,590]" captionText="Table 3. Number of specimens of the species of Cetopsidium with cited number of branched anal-fin rays. Bold indicates value for holotype when examined." pageId="9" pageNumber="136">Table 3</tableCitation>
) and “Região Mapuera” (MAP of
<bibRefCitation author="Ferreira, E" box="[357,509,1121,1144]" pageId="9" pageNumber="136" pagination="1 - 4" refId="ref108846" refString="Ferreira, E. 1995. Composicao, distribuicao e aspectos ecologicos da ictiofauna de um trecho do rio Trombetas, na area de influencia da futura UHE Cachoeira Porteira, Estado do Para, Brasil. ActaAmazonica, 23 (1 - 4, Suplemento): 1 - 88. [issue for 1993, actual publication date 1995]" type="journal article" year="1995">Ferreira, 1995</bibRefCitation>
:
<tableCitation box="[516,592,1121,1144]" captionStart="Table 3" captionStartId="22.[158,221,567,590]" captionText="Table 3. Number of specimens of the species of Cetopsidium with cited number of branched anal-fin rays. Bold indicates value for holotype when examined." pageId="9" pageNumber="136">Table 3</tableCitation>
) and find that these population samples represent a previously undescribed species that we describe herein as
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[546,786,1184,1208]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="136" phylum="Chordata" rank="species" species="ferreirai">
<emphasis box="[546,786,1184,1208]" italics="true" pageId="9" pageNumber="136">Cetopsidium ferreirai</emphasis>
</taxonomicName>
. Given those findings we assume that Ferreiras references of
<taxonomicName baseAuthorityName="Eigenmann" baseAuthorityYear="1912" box="[151,402,1248,1272]" class="Actinopterygii" family="Cetopsidae" genus="Pseudocetopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="136" phylum="Chordata" rank="species" species="minuta">
<emphasis box="[151,402,1248,1272]" italics="true" pageId="9" pageNumber="136">Pseudocetopsis minuta</emphasis>
</taxonomicName>
from “Região Oriximiná” (ORI of Ferreira,1993:
<tableCitation box="[297,368,1281,1304]" captionStart="Table 3" captionStartId="22.[158,221,567,590]" captionText="Table 3. Number of specimens of the species of Cetopsidium with cited number of branched anal-fin rays. Bold indicates value for holotype when examined." pageId="9" pageNumber="136">Table 3</tableCitation>
) and“Região Caxiaporé” (CAX of Ferreira, 1993:
<tableCitation box="[215,291,1313,1336]" captionStart="Table 3" captionStartId="22.[158,221,567,590]" captionText="Table 3. Number of specimens of the species of Cetopsidium with cited number of branched anal-fin rays. Bold indicates value for holotype when examined." pageId="9" pageNumber="136">Table 3</tableCitation>
) likely also refer to
<emphasis box="[495,728,1312,1336]" italics="true" pageId="9" pageNumber="136">
<taxonomicName authorityName="Vari &amp; Ferraris Jr &amp; de Pinna" authorityYear="2005" box="[495,724,1312,1336]" class="Actinopterygii" family="Cetopsidae" genus="Cetopsidium" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="136" phylum="Chordata" rank="species" species="ferreirai">Cetopsidium ferreirai</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="136" type="materials_examined">
<paragraph blockId="9.[151,794,1377,1657]" pageId="9" pageNumber="136">
<emphasis bold="true" box="[151,356,1377,1398]" pageId="9" pageNumber="136">Material examined.</emphasis>
<materialsCitation collectingDate="1985-04-19" collectionCode="INPA" collectorName="E. J. G. Ferreira" country="Brazil" county="rio Trombetas" latitude="-1.0833334" location="Para" longLatPrecision="1308" longitude="-57.016666" municipality="rio Mapuera" pageId="9" pageNumber="136" specimenCode="INPA 10194" specimenCount="7" typeStatus="holotype">
<specimenCount box="[365,486,1377,1398]" pageId="9" pageNumber="136" type="generic">7 specimens</specimenCount>
(
<quantity box="[503,633,1377,1398]" metricMagnitude="-2" metricUnit="m" metricValue="2.2125" metricValueMax="3.7" metricValueMin="0.725" pageId="9" pageNumber="136" unit="mm" value="22.125" valueMax="37.0" valueMin="7.25">7, 25-37 mm</quantity>
SL).
<emphasis bold="true" box="[692,787,1377,1398]" pageId="9" pageNumber="136">
<typeStatus box="[692,787,1377,1398]" pageId="9" pageNumber="136">Holotype</typeStatus>
</emphasis>
.
<collectingCountry box="[151,214,1405,1426]" name="Brazil" pageId="9" pageNumber="136">Brazil</collectingCountry>
.
<location LSID="urn:lsid:plazi:treatment:E56BC71F0E31FFA93C85FBD5FF00E42C:681D20D20E3EFFA93F93FAD5FEC0E7C7" box="[223,270,1405,1426]" country="Brazil" county="rio Trombetas" latitude="-1.0833334" longLatPrecision="1308" longitude="-57.016666" municipality="rio Mapuera" name="Para" pageId="9" pageNumber="136">Pará</location>
:
<location LSID="urn:lsid:plazi:treatment:E56BC71F0E31FFA93C85FBD5FF00E42C:681D20D20E3EFFA93E54FAD5FE6CE7C7" box="[280,418,1405,1426]" country="Brazil" county="rio Trombetas" latitude="-1.0833334" longLatPrecision="1308" longitude="-57.016666" municipality="rio Mapuera" name="rio Trombetas" pageId="9" pageNumber="136">rio Trombetas</location>
basin,
<collectingMunicipality box="[489,611,1405,1426]" pageId="9" pageNumber="136">rio Mapuera</collectingMunicipality>
, last cachoeira before
<collectingCounty box="[194,327,1434,1455]" pageId="9" pageNumber="136">rio Trombetas</collectingCounty>
(approximately
<geoCoordinate box="[484,553,1434,1455]" degrees="1" direction="south" minutes="05" orientation="latitude" pageId="9" pageNumber="136" precision="925" value="-1.0833334">1°05S</geoCoordinate>
,
<geoCoordinate box="[562,654,1434,1455]" degrees="57" direction="west" minutes="01" orientation="longitude" pageId="9" pageNumber="136" precision="925" value="-57.016666">57°01W</geoCoordinate>
),
<specimenCode box="[670,790,1434,1455]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="9" pageNumber="136">INPA 10194</specimenCode>
, male,
<quantity box="[213,283,1463,1484]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="136" unit="mm" value="25.0">25 mm</quantity>
SL;
<collectorName box="[335,496,1463,1484]" pageId="9" pageNumber="136">E. J. G. Ferreira</collectorName>
,
<date box="[508,651,1463,1484]" pageId="9" pageNumber="136" value="1985-04-19">
<collectingDate box="[508,651,1463,1484]" pageId="9" pageNumber="136" value="1985-04-19">19 April 1985</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation collectingDate="1985-04-19" collectionCode="INPA" collectorName="E. J. G. Ferreira" country="Brazil" county="Cachoeira Porteira" latitude="-1.0833334" location="Para" longLatPrecision="1308" longitude="-57.016666" municipality="rio Trombetas" pageId="9" pageNumber="136" specimenCode="INPA 10193, 3" specimenCount="6" typeStatus="paratype">
<emphasis bold="true" box="[662,766,1463,1484]" pageId="9" pageNumber="136">
<typeStatus box="[662,766,1463,1484]" pageId="9" pageNumber="136">Paratypes</typeStatus>
</emphasis>
.
<specimenCount pageId="9" pageNumber="136" type="generic">6 specimens</specimenCount>
(
<quantity box="[269,398,1492,1513]" metricMagnitude="-2" metricUnit="m" metricValue="2.1625" metricValueMax="3.7" metricValueMin="0.625" pageId="9" pageNumber="136" unit="mm" value="21.625" valueMax="37.0" valueMin="6.25">6, 25-37 mm</quantity>
SL)
<collectingCountry box="[450,513,1492,1513]" name="Brazil" pageId="9" pageNumber="136">Brazil</collectingCountry>
.
<location LSID="urn:lsid:plazi:treatment:E56BC71F0E31FFA93C85FBD5FF00E42C:681D20D20E3EFFA93D40FA7CFDF5E7BC" box="[524,571,1492,1513]" country="Brazil" county="Cachoeira Porteira" latitude="-1.0833334" longLatPrecision="1308" longitude="-57.016666" municipality="rio Trombetas" name="Para" pageId="9" pageNumber="136">Pará</location>
:
<collectingMunicipality box="[582,724,1492,1513]" pageId="9" pageNumber="136">rio Trombetas</collectingMunicipality>
, above
<collectingCounty box="[151,329,1521,1542]" pageId="9" pageNumber="136">Cachoeira Porteira</collectingCounty>
(approximately
<geoCoordinate box="[485,554,1521,1542]" degrees="1" direction="south" minutes="05" orientation="latitude" pageId="9" pageNumber="136" precision="925" value="-1.0833334">1°05S</geoCoordinate>
,
<geoCoordinate box="[563,654,1521,1542]" degrees="57" direction="west" minutes="01" orientation="longitude" pageId="9" pageNumber="136" precision="925" value="-57.016666">57°01W</geoCoordinate>
),
<specimenCode box="[670,789,1521,1542]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="9" pageNumber="136">INPA 10193</specimenCode>
,
<specimenCode box="[151,163,1549,1570]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="9" pageNumber="136">3</specimenCode>
(26-33),
<collectorName box="[251,401,1549,1570]" pageId="9" pageNumber="136">E. J. G. Ferreira</collectorName>
,
<date box="[409,543,1549,1570]" pageId="9" pageNumber="136" value="1985-04-19">
<collectingDate box="[409,543,1549,1570]" pageId="9" pageNumber="136" value="1985-04-19">19 April 1985</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation collectingDate="1985-04-19" collectionCode="INPA" collectorName="E. J. G. Ferreira" country="Brazil" latitude="-1.0833334" location="rio Trombetas" longLatPrecision="1308" longitude="-57.016666" municipality="Cachoeira Porteira" pageId="9" pageNumber="136" specimenCode="INPA 6501, 3" specimenCount="1" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:E56BC71F0E31FFA93C85FBD5FF00E42C:681D20D20E3EFFA93D6BF9A5FD61E477" box="[551,687,1549,1570]" country="Brazil" latitude="-1.0833334" longLatPrecision="1308" longitude="-57.016666" municipality="Cachoeira Porteira" name="rio Trombetas" pageId="9" pageNumber="136">rio Trombetas</location>
,
<collectingMunicipality pageId="9" pageNumber="136">Cachoeira Porteira</collectingMunicipality>
(approximately
<geoCoordinate box="[391,460,1578,1599]" degrees="1" direction="south" minutes="05" orientation="latitude" pageId="9" pageNumber="136" precision="925" value="-1.0833334">1°05S</geoCoordinate>
,
<geoCoordinate box="[471,563,1578,1599]" degrees="57" direction="west" minutes="01" orientation="longitude" pageId="9" pageNumber="136" precision="925" value="-57.016666">57°01W</geoCoordinate>
),
<specimenCode box="[582,693,1578,1599]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="9" pageNumber="136">INPA 6501</specimenCode>
,
<specimenCode box="[703,715,1578,1599]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="9" pageNumber="136">3</specimenCode>
(25-37;
<specimenCount box="[151,263,1607,1628]" pageId="9" pageNumber="136" type="generic">1 specimen</specimenCount>
,
<quantity box="[271,340,1607,1628]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="136" unit="mm" value="37.0">37 mm</quantity>
, cleared and stained),
<collectorName box="[553,705,1607,1628]" pageId="9" pageNumber="136">E. J. G. Ferreira</collectorName>
,
<date pageId="9" pageNumber="136" value="1985-04-19">
<collectingDate pageId="9" pageNumber="136" value="1985-04-19">19 April 1985</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>