273 lines
26 KiB
XML
273 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.692.14706" ID-GBIF-Dataset="81f065e7-a342-495c-9855-21bc26539311" ID-PMC="PMC5672734" ID-Pensoft-Pub="1313-2970-692-1" ID-PubMed="29133987" ID-ZBK="1B7C3F517CF543338EAB1CB1BD9D8A07" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-692-1" ModsDocOrigin="ZooKeys 692" ModsDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" checkinTime="1503352913593" checkinUser="pensoft" docAuthor="Foon, Junn Kitt & Liew, Thor-Seng" docDate="2017" docId="E4F3ED4DDF94DF97C4DBBBF4A6F331C7" docLanguage="en" docName="ZooKeys 692: 1-81" docOrigin="ZooKeys 692" docSource="http://dx.doi.org/10.3897/zookeys.692.14706" docTitle="Alycaeus thieroti Morgan 1885" docType="treatment" docVersion="5" lastPageNumber="47" masterDocId="FF8E6C15A17CDA48FFA7FFE36C7CFFA2" masterDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" masterLastPageNumber="81" masterPageNumber="1" pageNumber="44" updateTime="1668164728433" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Foon, Junn Kitt</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Liew, Thor-Seng</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2017</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>692</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>81</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.692.14706</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.692.14706</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-692-1</mods:identifier>
|
||
<mods:identifier type="ZBK">1B7C3F517CF543338EAB1CB1BD9D8A07</mods:identifier>
|
||
<mods:identifier type="ZooBank">1B7C3F517CF543338EAB1CB1BD9D8A07</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="132786242" LSID="urn:lsid:plazi:treatment:E4F3ED4DDF94DF97C4DBBBF4A6F331C7" httpUri="http://treatment.plazi.org/id/E4F3ED4DDF94DF97C4DBBBF4A6F331C7" lastPageId="46" lastPageNumber="47" pageId="43" pageNumber="44">
|
||
<subSubSection pageId="43" pageNumber="44" type="nomenclature">
|
||
<paragraph pageId="43" pageNumber="44">
|
||
<taxonomicName authority="Morgan, 1885 b" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus thieroti" order="Architaenioglossa" pageId="43" pageNumber="44" phylum="Mollusca" rank="species" species="thieroti">Alycaeus thieroti Morgan, 1885b</taxonomicName>
|
||
Figures 7D, 29, 31C
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="44" pageNumber="45" type="reference_group">
|
||
<paragraph pageId="44" pageNumber="45">
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus thieroti" order="Architaenioglossa" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="thieroti">
|
||
<pageBreakToken pageId="44" pageNumber="45" start="start">Alycaeus</pageBreakToken>
|
||
thieroti
|
||
</taxonomicName>
|
||
Morgan, 1885b: 403-404, plate 10;
|
||
<bibRefCitation author="Venmans, LAWC" journalOrPublisher="Proceedings of the Malacological Society" pageId="53" pageNumber="54" pagination="81 - 87" title="Notes on Alycaeus." volume="32" year="1956">Venmans 1956</bibRefCitation>
|
||
: 82, figure 3;
|
||
<bibRefCitation author="Benthem Jutting, van WSS" journalOrPublisher="Basteria" pageId="49" pageNumber="50" pagination="10 - 20" title="Some notes on land and freshwater mollusca of Malaya." url="http://natuurtijdschriften.nl/record/596402" volume="24" year="1960 a">Benthem Jutting 1960a</bibRefCitation>
|
||
: 13;
|
||
<bibRefCitation author="Sykes, ER" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="53" pageNumber="54" pagination="194 - 199" title="On the land operculate Mollusca collected during the " Skeat Expedition " to the Malay Peninsula in 1899 - 1900." url="http://www.biodiversitylibrary.org/part/145330/summary" volume="1903" year="1903">Sykes 1903</bibRefCitation>
|
||
: 197;
|
||
<bibRefCitation author="Chan, SY" journalOrPublisher="Club Conchylia Informationen" pageId="49" pageNumber="50" pagination="35 - 46" title="Non-marine mollusks from Selangor, West Malaysia, with a comparison note on introduced species." volume="29" year="1997">Chan 1997</bibRefCitation>
|
||
: 37;
|
||
<bibRefCitation pageId="44" pageNumber="45">Maassen 2001</bibRefCitation>
|
||
: 23;
|
||
<bibRefCitation author="Clements, R" journalOrPublisher="Biological Conservation" pageId="49" pageNumber="50" pagination="2751 - 2764" title="Using biogeographical patterns of endemic land snails to improve conservation planning for limestone karsts." url="https://doi.org/10.1016/j.biocon.2008.08.011" volume="141" year="2008">Clements et al. 2008</bibRefCitation>
|
||
: 2760;
|
||
<bibRefCitation author="Tarruella, A" journalOrPublisher="Spira" pageId="53" pageNumber="54" pagination="71 - 76" title="Listado taxonomico ilustrado de la familia Cyclophoridae J. E. Gray, 1847 (Mollusca: Gastropoda): Parte 2. La subfamilia Alycaeinae J. E. Gray, 1850." url="http://www.molluscat.com/assets/spira_4_12_7.pdf" volume="4" year="2011">
|
||
Tarruella and
|
||
<normalizedToken originalValue="Domènech">Domenech</normalizedToken>
|
||
2011
|
||
</bibRefCitation>
|
||
: 73;
|
||
<bibRefCitation author="Foon, JK" journalOrPublisher="ZooKeys" pageId="50" pageNumber="51" pagination="1 - 94" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. 2017</bibRefCitation>
|
||
: 13, figure 5A.
|
||
</paragraph>
|
||
<paragraph pageId="44" pageNumber="45">
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Orthalycaeus) thieroti" order="Architaenioglossa" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="thieroti" subGenus="Orthalycaeus">Alycaeus (Orthalycaeus) thieroti</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Moellendorff, von OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="52" pageNumber="53" pagination="330 - 348" title="On the land and freshwater shells of Perak." url="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" volume="1891" year="1891">
|
||
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
1891
|
||
</bibRefCitation>
|
||
: 342; Kobelt and
|
||
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
<bibRefCitation author="Kobelt, W" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="51" pageNumber="52" pagination="73 - 192" title="Catalog der gegenwaertig lebend bekannten Pneumonopomen." url="https://doi.org/10.5962/bhl.title.14877" volume="1" year="1897">1897</bibRefCitation>
|
||
: 151.
|
||
</paragraph>
|
||
<paragraph pageId="44" pageNumber="45">
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Alycaeus) thieroti" order="Architaenioglossa" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="thieroti" subGenus="Alycaeus">Alycaeus (Alycaeus) thieroti</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="51" pageNumber="52" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In: Kobelt W (Ed.) In Verbindung mit der deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." url="https://doi.org/10.5962/bhl.title.1227" year="1902">Kobelt 1902</bibRefCitation>
|
||
: 352;
|
||
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malayan Branch of the Royal Asiatic Society" pageId="51" pageNumber="52" pagination="25 - 37" title="A list of the Land and Fresh-water Mollusca of the Malay Peninsula with notes." url="http://www.jstor.org/stable/41559818" volume="6" year="1928">Laidlaw 1928</bibRefCitation>
|
||
: 35;
|
||
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="The Bulletin of the Raffles Museum" pageId="51" pageNumber="52" pagination="35 - 41" title="Notes on the land-mollusca of the Batu Caves, Selangor, with descriptions of two new species." url="http://lkcnhm.nus.edu.sg/nus/pdf/PUBLICATION/Raffles%20Bulletin%20of%20Zoology/Past%20Volumes/RBZ%207(1)/07brm035-041.pdf" volume="7" year="1932">Laidlaw 1932</bibRefCitation>
|
||
: 36.
|
||
</paragraph>
|
||
<paragraph pageId="44" pageNumber="45">
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus liratulus" order="Architaenioglossa" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="liratulus">Alycaeus liratulus</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Liew, TS" editor="Che, Omar I" journalOrPublisher="The Bulletin of the Raffles Museum" pageId="51" pageNumber="52" title="Penerbit Universiti Malaysia Kelantan, Kota Bharu" url="https://doi.org/10.13140/RG.2.1.4311.5364" year="2010">Liew 2010</bibRefCitation>
|
||
: 101. [not
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus liratulus" order="Architaenioglossa" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="liratulus">Alycaeus liratulus</taxonomicName>
|
||
(Preston, 1907)]
|
||
</paragraph>
|
||
<paragraph pageId="44" pageNumber="45">
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Pincerna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pincerna thieroti" order="Architaenioglossa" pageId="44" pageNumber="45" phylum="Mollusca" rank="species" species="thieroti">Pincerna thieroti</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Pall-Gergely, B" journalOrPublisher="Raffles Bulletin of Zoology" pageId="52" pageNumber="53" pagination="213 - 219" title="A new species of Alycaeidae, Pincernayanseni n. sp. from Sumatra, with the resurrection of the genus Pincerna Preston, 1907 (Gastropoda: Cyclophoroidea)." volume="65" year="2017">
|
||
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
||
2017
|
||
</bibRefCitation>
|
||
: 213-214.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="44" pageNumber="45" type="type locality">
|
||
<paragraph pageId="44" pageNumber="45">Type locality.</paragraph>
|
||
<paragraph pageId="44" pageNumber="45">
|
||
PRK 18 Gunung Lanno, Perak (
|
||
<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.5252776">4°31'31"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="east" orientation="longitude" precision="15" value="101.14667">101°08'48"E</geoCoordinate>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="44" pageNumber="45" type="type material">
|
||
<paragraph pageId="44" pageNumber="45">Type material.</paragraph>
|
||
<paragraph pageId="44" pageNumber="45">Syntype. Gunong Lano (= PRK 18 Gunung Lanno), Perak: MNHN IM-2000-31799/5 (1 specimen seen).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="materials_examined">
|
||
<paragraph pageId="44" pageNumber="45">Other examined materials.</paragraph>
|
||
<paragraph lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
|
||
Endau-Kluang, Johor: BOR/MOL 6466/1, BOR/MOL 6467/1. KTN 01 Gunung Reng, Kelantan: BOR/MOL 6835/1, BOR/MOL 8335/6. KTN 95 Gua Panjang, Kelantan: BOR/MOL 12962/5. Lojing, Kelantan: BOR/MOL 6528/2. PHG 01 Kota Gelanggi, Pahang: BOR/MOL 6886/2, BOR/MOL 6636/1. PHG 02 Gunung Senyum, Pahang: BOR/MOL 12961/4. PHG 03 Gunung Jebak Puyuh, Pahang: BOR/MOL 6885/3. PHG 05 Bukit Cintamanis, Pahang: BOR/MOL 6633/1. PHG 75 Bukit Tenggek, Pahang: ZRC 1975.2.22.8-11/2. PHG 77 Bukit Mengapur, Pahang: BOR/MOL 6889/1. Unnamed limestone hill north of Jalan Lipis-Gua Musang Lama, Felda Chegar Perah I and II, Pahang: BOR/MOL 6815/1. Gunung Tangkup, Kenong Rimba, Pahang: BOR/MOL 6237/1.
|
||
<pageBreakToken pageId="45" pageNumber="46" start="start">Gunung</pageBreakToken>
|
||
Kesong, Kenong Rimba, Pahang: BOR/MOL 6253/2. PRK 55 Gunung Pondok, Perak: BOR/MOL 8410/1. PRK 23 Gunung Rapat, Perak: BOR/MOL 10056/4, BOR/MOL 10258/2. PRK 62 Gunong Dayak, Perak: ZRC 1975.2.22.1/1. Mykarst-025, Perak: BOR/MOL 9415/1, BOR/MOL 12422/1. Lenggong, Perak: ZRC 1975.2.22.2-3/2. SGR 02 Bukit Takun, Selangor: ZRC 1997.27/5.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="45" pageNumber="46" type="description">
|
||
<paragraph pageId="45" pageNumber="46">Description.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Protoconch. Smooth.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="45" pageNumber="46" type="description">
|
||
<paragraph pageId="45" pageNumber="46">Shell shape and size.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Globose. Shell height: 4.55-5.68 mm. Shell width: 4.36-5.34 mm.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Spire. Spire height: 1.06-1.58 mm. Spire width: 1.73-2.30 mm. Number of whorls: up to 4. Spire shape: oblong conical. Whorl periphery rounded. Umbilicus open or partially closed.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Whorl constriction. At about 3 ⅝ whorls posterior of protoconch.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Breathing tube. Length: 0.40-0.53 mm.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Aperture and peristome. Aperture circular, moderately expanded. Aperture height: 2.28-2.76 mm. Aperture width: 2.42-2.98 mm. Peristome double, thickened, notched at suture. No interspace. Peristome orientation 15-18° oblique with respect to the coiling axis.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Spiral lines. Distinct. Regularly spaced. 20-37 lines per 1 mm.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Radial ribs running anterior of breathing tube. Radial ribs pronounced, evenly spaced. Approximately 8-15 ribs per 1 mm.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Radial ribs running perpendicular to breathing tube. Radial ribs pronounced, evenly spaced. Approximately 9-21 ribs per 1 mm.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Radial ribs running posterior of breathing tube. Ranges from absent to indistinct, unevenly spaced.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Operculum. Almost flat. Exterior covered by calcareous layer. Exterior has calcareous ridges radiating outwards of the nucleus. Interior covered by proteinaceous layer, smooth.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Shell colour. Variable - red, yellow or white.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Living animal. Body cream-white or maroon. Head pink. Tentacles red or brown tipped.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="45" pageNumber="46" type="habitat">
|
||
<paragraph pageId="45" pageNumber="46">Habitat and ecology.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Lives in crevices on wet limestone boulders and shrubs close to the ground. In thickly forested areas. Lives in both limestone and non-limestone areas.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="45" pageNumber="46" type="distribution">
|
||
<paragraph pageId="45" pageNumber="46">Distribution.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">Widespread in Perak, Selangor, Kelantan and Pahang.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="45" pageNumber="46" type="differential diagnosis">
|
||
<paragraph pageId="45" pageNumber="46">Differential diagnosis.</paragraph>
|
||
<paragraph pageId="45" pageNumber="46">
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus thieroti" order="Architaenioglossa" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="thieroti">Alycaeus thieroti</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus liratulus" order="Architaenioglossa" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="liratulus">Alycaeus liratulus</taxonomicName>
|
||
in shell shape but differs in having a smaller and more elongated shell (smaller by about 1 mm in shell height, 0.8 mm in shell width), narrowly-spaced radial ribs and spiral lines, shorter breathing tube (shorter by 0.65 mm) and flat operculum with calcareous ridges radiating out from the exterior nucleus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="discussion">
|
||
<paragraph pageId="45" pageNumber="46">Discussion.</paragraph>
|
||
<paragraph lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
|
||
<bibRefCitation author="Morgan, de J" journalOrPublisher="Bulletin de la Societe zoologique de France" pageId="52" pageNumber="53" pagination="353 - 429" title="Mollusques terrestes et fiuviatiles de Royaume de Perak et des pais voisins (presque'Ile Malaise)." url="https://doi.org/10.5962/bhl.part.14300" volume="10" year="1885 b">Morgan (1885b)</bibRefCitation>
|
||
first described
|
||
<taxonomicName lsidName="A. thieroti" pageId="45" pageNumber="46" rank="species" species="thieroti">A. thieroti</taxonomicName>
|
||
with simple shell characters and an erroneous description of the operculum as smooth and proteinaceous.
|
||
<bibRefCitation author="Moellendorff, von OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="52" pageNumber="53" pagination="330 - 348" title="On the land and freshwater shells of Perak." url="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" volume="1891" year="1891">
|
||
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
(1891)
|
||
</bibRefCitation>
|
||
subsequently redescribed the species with additional shell characters but did not point out the error. The operculum is a critical diagnostic feature that clearly distinguishes
|
||
<taxonomicName lsidName="A. thieroti" pageId="45" pageNumber="46" rank="species" species="thieroti">A. thieroti</taxonomicName>
|
||
from
|
||
<taxonomicName lsidName="A. liratulus" pageId="45" pageNumber="46" rank="species" species="liratulus">A. liratulus</taxonomicName>
|
||
. Thus, this uncorrected error has led to the misidentification of
|
||
<taxonomicName lsidName="A. thieroti" pageId="45" pageNumber="46" rank="species" species="thieroti">A. thieroti</taxonomicName>
|
||
as
|
||
<taxonomicName lsidName="A. liratulus" pageId="45" pageNumber="46" rank="species" species="liratulus">A. liratulus</taxonomicName>
|
||
and vice versa (e.g.
|
||
<bibRefCitation author="Liew, TS" editor="Che, Omar I" journalOrPublisher="The Bulletin of the Raffles Museum" pageId="51" pageNumber="52" title="Penerbit Universiti Malaysia Kelantan, Kota Bharu" url="https://doi.org/10.13140/RG.2.1.4311.5364" year="2010">Liew 2010</bibRefCitation>
|
||
, ZRC collection lots).
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus thieroti" order="Architaenioglossa" pageId="45" pageNumber="46" phylum="Mollusca" rank="species" species="thieroti">Alycaeus thieroti</taxonomicName>
|
||
was known from Perak, Selangor and Kelantan prior to this study (
|
||
<bibRefCitation author="Morgan, de J" journalOrPublisher="Bulletin de la Societe zoologique de France" pageId="52" pageNumber="53" pagination="353 - 429" title="Mollusques terrestes et fiuviatiles de Royaume de Perak et des pais voisins (presque'Ile Malaise)." url="https://doi.org/10.5962/bhl.part.14300" volume="10" year="1885 b">Morgan 1885b</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Moellendorff, von OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="52" pageNumber="53" pagination="330 - 348" title="On the land and freshwater shells of Perak." url="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" volume="1891" year="1891">
|
||
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
|
||
1891
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Sykes, ER" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="53" pageNumber="54" pagination="194 - 199" title="On the land operculate Mollusca collected during the " Skeat Expedition " to the Malay Peninsula in 1899 - 1900." url="http://www.biodiversitylibrary.org/part/145330/summary" volume="1903" year="1903">Sykes 1903</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malayan Branch of the Royal Asiatic Society" pageId="51" pageNumber="52" pagination="25 - 37" title="A list of the Land and Fresh-water Mollusca of the Malay Peninsula with notes." url="http://www.jstor.org/stable/41559818" volume="6" year="1928">Laidlaw 1928</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="The Bulletin of the Raffles Museum" pageId="51" pageNumber="52" pagination="35 - 41" title="Notes on the land-mollusca of the Batu Caves, Selangor, with descriptions of two new species." url="http://lkcnhm.nus.edu.sg/nus/pdf/PUBLICATION/Raffles%20Bulletin%20of%20Zoology/Past%20Volumes/RBZ%207(1)/07brm035-041.pdf" volume="7" year="1932">
|
||
<pageBreakToken pageId="46" pageNumber="47" start="start">Laidlaw</pageBreakToken>
|
||
1932
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Benthem Jutting, van WSS" journalOrPublisher="Basteria" pageId="49" pageNumber="50" pagination="10 - 20" title="Some notes on land and freshwater mollusca of Malaya." url="http://natuurtijdschriften.nl/record/596402" volume="24" year="1960 a">Benthem Jutting 1960a</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Venmans, LAWC" journalOrPublisher="Proceedings of the Malacological Society" pageId="53" pageNumber="54" pagination="81 - 87" title="Notes on Alycaeus." volume="32" year="1956">Venmans 1956</bibRefCitation>
|
||
).
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus thieroti" order="Architaenioglossa" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="thieroti">Alycaeus thieroti</taxonomicName>
|
||
in Pahang and Johor are new records.
|
||
<bibRefCitation author="Venmans, LAWC" journalOrPublisher="Proceedings of the Malacological Society" pageId="53" pageNumber="54" pagination="81 - 87" title="Notes on Alycaeus." volume="32" year="1956">Venmans (1956)</bibRefCitation>
|
||
examined the radular and jaw features of
|
||
<taxonomicName lsidName="A. thieroti" pageId="46" pageNumber="47" rank="species" species="thieroti">A. thieroti</taxonomicName>
|
||
, which is distinct from
|
||
<taxonomicName lsidName="A. selangoriensis" pageId="46" pageNumber="47" rank="species" species="selangoriensis">A. selangoriensis</taxonomicName>
|
||
sp. n. but similar to
|
||
<taxonomicName lsidName="A. conformis" pageId="46" pageNumber="47" rank="species" species="conformis">A. conformis</taxonomicName>
|
||
in teeth arrangement. Note that we maintained
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Pincerna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pincerna" order="Architaenioglossa" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">Pincerna</taxonomicName>
|
||
as a subgenus of
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
|
||
although
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Pincerna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pincerna" order="Architaenioglossa" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">Pincerna</taxonomicName>
|
||
was recently elevated to genus level (
|
||
<bibRefCitation author="Pall-Gergely, B" journalOrPublisher="Raffles Bulletin of Zoology" pageId="52" pageNumber="53" pagination="213 - 219" title="A new species of Alycaeidae, Pincernayanseni n. sp. from Sumatra, with the resurrection of the genus Pincerna Preston, 1907 (Gastropoda: Cyclophoroidea)." volume="65" year="2017">
|
||
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
||
2017
|
||
</bibRefCitation>
|
||
). See Remarks section under genus
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
|
||
for discussion.
|
||
</paragraph>
|
||
<caption pageId="46" pageNumber="47">
|
||
<paragraph pageId="46" pageNumber="47">
|
||
Figure 29.
|
||
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus thieroti" order="Architaenioglossa" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="thieroti">Alycaeus thieroti</taxonomicName>
|
||
Morgan, 1885b.
|
||
<normalizedToken originalValue="A–E">A-E</normalizedToken>
|
||
Shell of syntype, MNHN-IM-2000-31799
|
||
<normalizedToken originalValue="F–J">F-J</normalizedToken>
|
||
Shell of BOR/MOL 6835
|
||
<normalizedToken originalValue="K–M">K-M</normalizedToken>
|
||
Close up of shell of BOR/MOL 6835
|
||
<normalizedToken originalValue="N–Q">N-Q</normalizedToken>
|
||
Operculum of BOR/MOL 6835 R Shell of BOR/MOL 6528a
|
||
<normalizedToken originalValue="S–W">S-W</normalizedToken>
|
||
Shell of BOR/MOL 6528b. Scale bars:
|
||
<normalizedToken originalValue="A–J">A-J</normalizedToken>
|
||
,
|
||
<normalizedToken originalValue="R–W">R-W</normalizedToken>
|
||
1 mm; K, L 1 mm; M,
|
||
<normalizedToken originalValue="N–Q">N-Q</normalizedToken>
|
||
0.5 mm. Photographs
|
||
<normalizedToken originalValue="A–E">A-E</normalizedToken>
|
||
by Manuel Caballer (reproduced herein under CC-BY 4.0 license, Project RECOLNAT, MNHN). Photographs
|
||
<normalizedToken originalValue="F–W">F-W</normalizedToken>
|
||
by Junn Kitt Foon.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |