412 lines
49 KiB
XML
412 lines
49 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.231.100894" ID-Pensoft-Pub="1314-2003-231-1" ID-Pensoft-UUID="0D762CF99C605B04AF072C867EEB1488" ModsDocID="1314-2003-231-1" checkinTime="1693383686345" checkinUser="pensoft" docAuthor="Knapp, Sandra, Saerkinen, Tiina & Barboza, Gloria E." docDate="2023" docId="E4EDA60353E25204071F3E5A674FB06A" docLanguage="en" docName="PhytoKeys 231: 1-342" docOrigin="PhytoKeys 231" docPubDate="2023-08-29" docSource="http://dx.doi.org/10.3897/phytokeys.231.100894" docTitle="Solanum echegarayi Hieron., Bol. Acad. Nac. Cienc. (Cordoba) 9: 58. 1881." docType="treatment" docVersion="1" id="0D762CF99C605B04AF072C867EEB1488" lastPageNumber="1" masterDocId="0D762CF99C605B04AF072C867EEB1488" masterDocTitle="A revision of the South American species of the Morelloid clade (Solanum L., Solanaceae)" masterLastPageNumber="342" masterPageNumber="1" pageNumber="1" updateTime="1693383686345" updateUser="pensoft">
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<mods:title id="36A21F34797EFA2384739271B961DEFA">A revision of the South American species of the Morelloid clade (Solanum L., Solanaceae)</mods:title>
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<mods:name id="D758E0A787097535D98FE6E87EFFF8DB" type="personal">
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<mods:namePart id="49B22B97BE589555FEAE4F1556378FD2">Knapp, Sandra</mods:namePart>
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<mods:affiliation id="626DE8A30760DAB737EF49EE50420D5A">Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:nameIdentifier id="69E7D2F75A9E9477C5C3C714C17EDD9B" type="email">s.knapp@nhm.ac.uk</mods:nameIdentifier>
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<mods:namePart id="F3477DBF5FA0DC40454F53F3B68B1E69">Saerkinen, Tiina</mods:namePart>
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<mods:nameIdentifier id="AAAB66AE10DBADF8590A345268440281" type="ORCID">https://orcid.org/0000-0002-6956-3093</mods:nameIdentifier>
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<mods:affiliation id="A20C0E71082FBEB9711EFB331358B60D">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh EH 3 5 LR, UK</mods:affiliation>
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<mods:namePart id="659DBF4E46EF08558A500E523D5E0815">Barboza, Gloria E.</mods:namePart>
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<mods:nameIdentifier id="AE62D7E99F5AB885311C17B24017D21C" type="ORCID">https://orcid.org/0000-0003-1085-036X</mods:nameIdentifier>
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<mods:affiliation id="79DE644324D73CDF8E47976AD8BA4693">Instituto Multidisciplinario de Biologia Vegetal (CONICET-Universidad Nacional de Cordoba), Casilla de Correo 495, 5000 Cordoba, Argentina</mods:affiliation>
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<mods:title id="0CD13914780E3A3AF37A40ABDE2B7188">PhytoKeys</mods:title>
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<mods:date id="601EC445336A1009B35CD3DE68B30177">2023</mods:date>
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<mods:number id="BC0FBA9F637198B7D69DF9400A5115A2">231</mods:number>
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<mods:identifier id="30DCFBF645B7499F904E2B03044585A3" type="DOI">http://dx.doi.org/10.3897/phytokeys.231.100894</mods:identifier>
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<treatment id="E4EDA60353E25204071F3E5A674FB06A" LSID="urn:lsid:plazi:treatment:E4EDA60353E25204071F3E5A674FB06A" httpUri="http://treatment.plazi.org/id/E4EDA60353E25204071F3E5A674FB06A" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="89D21722BD7200A1DA34698C78CB5887" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="BB5723EFFDC40FAD2EAAE7BA680E7893" pageId="0" pageNumber="1">
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15.
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<taxonomicName id="505CD6AD13C4D788A1E7E36905B4E3E9" ID-CoL="4Y2BW" ID-ENA="1807823" authority="Hieron., Bol. Acad. Nac. Cienc. (Cordoba) 9: 58. 1881." authorityName="Hieron., Bol. Acad. Nac. Cienc. (Cordoba) 9: 58. 1881." class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum echegarayi" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="echegarayi">Solanum echegarayi Hieron., Bol. Acad. Nac. Cienc. (Cordoba) 9: 58. 1881.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="3512E6A0AA13ABB49A5584169EB793F8" pageId="0" pageNumber="1" type="description">
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||
<paragraph id="FF7D673F77F82C38F2D9FE68F9A00C4C" pageId="0" pageNumber="1">
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<figureCitation id="49A39FAA0FD281D9F35DB4DC00E01905" captionStart="Figure 47" captionStartId="F47" captionText="Figure 47. Solanum echegarayi A flowering and fruiting branch B flower C calyx D dissected flower E stamen, dorsal view F stamen, ventral view G gynoecium H fruit I seed (A-I Biurrun et al. 5038). Illustration by M. Moreno. Previously published in Barboza et al. (2013: 249), as S. hastatilobum." figureDoi="10.3897/phytokeys.231.100894.figure47" httpUri="https://binary.pensoft.net/fig/897508" pageId="0" pageNumber="1">Figs 47</figureCitation>
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<figureCitation id="E0E13ED4BE2084FFF8B9604D0BA654E0" captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Solanum echegarayi A habit B underground woody rhizomes C flowers at anthesis D immature and mature fruits (A, C, D Barboza et al. 4783 B Knapp et al. 10540). Photos by G. E. Barboza and S. Knapp." figureDoi="10.3897/phytokeys.231.100894.figure48" httpUri="https://binary.pensoft.net/fig/897509" pageId="0" pageNumber="1">, 48</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="64082EE033D3BBC97E07916C43506AF2" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="E8D140A9E3A5A8961958A865B7205447" pageId="0" pageNumber="1">
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<taxonomicName id="D8A0DF08CC0648765F8CA4AA993AEFF5" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum juncalense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="juncalense">Solanum juncalense</taxonomicName>
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Reiche, Anales Univ. Chile 124: 459. 1909. Type. Chile.
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<normalizedToken id="698004946C0E28D5608C4A47CB1EEC7F" originalValue="Región">Region</normalizedToken>
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VII (Valparaiso): [Los Andes] Juncal [protologue: "Cordilleras de la provincia de Aconcagua, Juncal"], 15 Jan,
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<emphasis id="C34D7A1AFEE699765F7995CE578240F6" italics="true" pageId="0" pageNumber="1">O. Buchtien 150</emphasis>
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(no herbaria or collector cited; neotype, designated here: SGO [SGO000004574]).
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</paragraph>
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<paragraph id="159B4E243DDB17758A4EEA65F0A2E397" pageId="0" pageNumber="1">
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<taxonomicName id="D5F6C856F3BF5DAF9F9771F06AEDB9E5" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum hastatilobum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="hastatilobum">Solanum hastatilobum</taxonomicName>
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Bitter, Repert. Spec. Nov. Regni Veg. 13: 246. 1912. Type. Argentina. San Luis: Quebrada del Salado, cerca de Bebida de las Varas, 9 Mar 1882,
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<emphasis id="78978E96BF2358EFC282B25B881C24F8" italics="true" pageId="0" pageNumber="1">C. Galander s.n.</emphasis>
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(holotype: B [destroyed]; lectotype, designated by Barboza et al. 2103, pg. 249: CORD [CORD00004221]).
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</paragraph>
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<paragraph id="263C84E87F3B964395CFEA35FF13BBB9" pageId="0" pageNumber="1">
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<taxonomicName id="3A4151006F6C59A6FA6A42F91F1EB2BB" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum juncalense subsp. aconcaguae" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="subSpecies" species="juncalense" subSpecies="aconcaguae">Solanum juncalense Reiche subsp. aconcaguae</taxonomicName>
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Bitter, Repert. Spec. Nov. Regni Veg. 12: 156. 1913. Type. Argentina. Mendoza: Dpto. Las Heras, "Puente del Inca, in viciniis montis Aconcagua", 23 Feb 1903,
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<emphasis id="CFF775DC220989577C01007A799F2A67" italics="true" pageId="0" pageNumber="1">G.A. Malme 2956</emphasis>
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(holotype: S [acc. # 10-15685]; isotypes: G [G00343486], MO [MO-256207, acc. # 2741560], US [00027638, acc. # 1572914]).
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</paragraph>
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<paragraph id="C5DF632DBC45EFC8AF6DFE50FD2EE6A2" pageId="0" pageNumber="1">
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<taxonomicName id="51EEF99DA9A66F695E17C0157422AD07" authorityName="Bitter" authorityYear="1914" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum hastatilobum subsp. brachyphyllum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="subSpecies" species="hastatilobum" subSpecies="brachyphyllum">Solanum hastatilobum Bitter subsp. brachyphyllum</taxonomicName>
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Bitter, Repert. Spec. Nov. Regni Veg. 13: 171. 1914. Type. Argentina. San Juan: Dpto. Angaco: Cumbre del Gato, Cerro Pico de Palo,
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<emphasis id="952256123EF28FADB3B15AB680A11E21" italics="true" pageId="0" pageNumber="1">T. Stuckert 7029</emphasis>
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(lectotype, designated by Barboza et al. 2103, pg. 249: G [G00343383]).
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</paragraph>
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<paragraph id="752BFAB2FE1EBC3CE31D1F1754DD2265" pageId="0" pageNumber="1">
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<taxonomicName id="26A6DB65CDC89DA2B0D558BCF0953E21" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum glaberrimum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="glaberrimum">Solanum glaberrimum</taxonomicName>
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C.V.Morton, Revis. Argentine Sp. Solanum 82. 1976. Type. Argentina. La Rioja: Quebrada de la Troya, 21 Feb 1941,
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<emphasis id="4D81E71A048001E6C32CA3C34BD8A138" italics="true" pageId="0" pageNumber="1">G. Covas 1235</emphasis>
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(holotype: GH [00062989]; isotypes: LP [LP010903, acc. # 048953], NY [00076825], US [00027581, acc. # 2639762, fragment of GH holotype]).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="78C29EE54CA367ABD5795BFE6F3B9F49" pageId="0" pageNumber="1" type="materials_examined">
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||
<paragraph id="66306685E224E9B1307FEA03A1A2ECEA" pageId="0" pageNumber="1">
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<typeStatus id="9FAB87629074CA1BE2EAA0D93B1C4B84">Type</typeStatus>
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.
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</paragraph>
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<paragraph id="C0E1C6BEB9C12B5E08FA8FCD31212BD2" pageId="0" pageNumber="1">
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<materialsCitation id="6A4F367EAABF4F94957C221DF2120EBA" collectingDate="1876-01" collectorName="S. Echegaray" country="Argentina" location="Salida de la Quebrada del Leoncito" specimenCount="1" stateProvince="San Juan" typeStatus="holotype">
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<collectingCountry id="A3B77D868BD803F56D36C793BCC194A2" name="Argentina">Argentina</collectingCountry>
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.
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<collectingRegion id="76912214691C7C193C51A2EDE07583C1" country="Argentina" name="San Juan">San Juan</collectingRegion>
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:
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<location id="86442E808DC4EBC0C12480EA1944DFB2" LSID="urn:lsid:plazi:treatment:E4EDA60353E25204071F3E5A674FB06A:86442E808DC4EBC0C12480EA1944DFB2" country="Argentina" name="Salida de la Quebrada del Leoncito" stateProvince="San Juan">Salida de la Quebrada del Leoncito</location>
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,
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<collectingDate id="6E64450F57C5876D3D183BAD3809A7F1" value="1876-01">Jan 1876</collectingDate>
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,
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<emphasis id="CB348870A09C1055CA5FF85D346905CB" italics="true" pageId="0" pageNumber="1">
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<collectorName id="DBA6FDEC043C66A2C4A02F1EC7BFD422">S. Echegaray</collectorName>
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s.n.
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</emphasis>
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(
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<typeStatus id="89221CC3DAB390187532B51951D16CCB">holotype</typeStatus>
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||
: CORD [CORD00004197]; isotype:
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</materialsCitation>
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<collectingCountry id="C9D3F7036B4D3B9CC11501D5D0EC7485" name="United States of America">US</collectingCountry>
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[00027559, acc. # 2678279]).
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</paragraph>
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||
<caption id="36B15B027A3A8229BA8011D043BA9151" doi="10.3897/phytokeys.231.100894.figure47" httpUri="https://binary.pensoft.net/fig/897508" pageId="0" pageNumber="1" start="Figure 47" startId="F47">
|
||
<paragraph id="800E518DF331B2C0D05B322EE4F10276" pageId="0" pageNumber="1">
|
||
<emphasis id="ADB5F10A349E13C9AE65F551D86D3CFA" bold="true" pageId="0" pageNumber="1">Figure 47.</emphasis>
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<taxonomicName id="00171684BBC3E676FAB0FE06A662F179" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum echegarayi" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="echegarayi">
|
||
<emphasis id="9B0B4FD6D0E3A77469AE857AC9E3DAF8" italics="true" pageId="0" pageNumber="1">Solanum echegarayi</emphasis>
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||
</taxonomicName>
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||
<emphasis id="609B73B750ACD17B0018C59151212935" bold="true" pageId="0" pageNumber="1">A</emphasis>
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||
flowering and fruiting branch
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||
<emphasis id="D0463C69D98059EEEB2E4090CA5B5C8C" bold="true" pageId="0" pageNumber="1">B</emphasis>
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||
flower
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||
<emphasis id="AF9B93954055CC266BD356028B2F2ED3" bold="true" pageId="0" pageNumber="1">C</emphasis>
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||
calyx
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||
<emphasis id="0B42E08C2636EE99D8862C6AF5CED921" bold="true" pageId="0" pageNumber="1">D</emphasis>
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||
dissected flower
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||
<emphasis id="A65F86DA16DA8D587263FB2617CA2519" bold="true" pageId="0" pageNumber="1">E</emphasis>
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||
stamen, dorsal view
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||
<emphasis id="F53CE4A2A251287F8B0763998850D70E" bold="true" pageId="0" pageNumber="1">F</emphasis>
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stamen, ventral view
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||
<emphasis id="74426D3E190F9A5A01DE133E5CEDF4FB" bold="true" pageId="0" pageNumber="1">G</emphasis>
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||
gynoecium
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||
<emphasis id="BED1A4A3C87E0911DC1CEA788BAF5A4F" bold="true" pageId="0" pageNumber="1">H</emphasis>
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||
fruit
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||
<emphasis id="E2D40072718C520827D3E801609E46DB" bold="true" pageId="0" pageNumber="1">I</emphasis>
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||
seed (
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||
<emphasis id="6F27BBAFB5CBFF512FE06528A3EA5D4A" bold="true" pageId="0" pageNumber="1">A-I</emphasis>
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||
<emphasis id="A42F7A8A6324C1CD43B6C361AD45D407" italics="true" pageId="0" pageNumber="1">Biurrun et al. 5038</emphasis>
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||
). Illustration by M. Moreno. Previously published in
|
||
<bibRefCitation id="FF3D33B8D9325CE6FB2D87BDE8447177" author="Barboza, GE" editor="Anton, AM" journalOrPublisher=") Flora Argentina (Vol. 13), Solanaceae. IOBDA- IMBIV, CONICET: Buenos Aires & Cordoba, Argentina" pageId="0" pageNumber="1" pagination="231 - 264" refId="B10" refString="Barboza, GE, Knapp, S, Saerkinen, T, 2013. Grupo VII. Moreloide. In: Anton, AM, Zuloaga, FO, Eds., Barboza GE (Coord.) Flora Argentina (Vol. 13), Solanaceae. IOBDA- IMBIV, CONICET: Buenos Aires & Cordoba, Argentina: 231 - 264" title="Grupo VII. Moreloide." volumeTitle="Barboza GE (Coord." year="2013">Barboza et al. (2013</bibRefCitation>
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||
: 249), as
|
||
<taxonomicName id="3D1F427CE11A64023ACE23EA658C2335" family="Solanaceae" kingdom="Plantae" lsidName="S. hastatilobum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="hastatilobum">
|
||
<emphasis id="129235333EC4EAFFEBE2055A3CBF4AD5" italics="true" pageId="0" pageNumber="1">S. hastatilobum</emphasis>
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||
</taxonomicName>
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||
.
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||
</paragraph>
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||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="67654162C2102D8734FFBD14D1FC9B09" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="158128B33FE7F447BC4FCCA59FE3EC35" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="BE77CBB2CBAEF6E1AA33A755FDF0384B" pageId="0" pageNumber="1">
|
||
Sprawling perennial herbs from woody rhizomes (underground rootstocks), prostrate to semi-erect, 0.1-0.5 m high, woody at the base, extremely variable in size depending on season of collection. Stems angled or slightly winged from the decurrent leaf bases, completely glabrous to sparsely and minutely pubescent with eglandular antrorse 1-2-celled simple uniseriate trichomes 0.1-0.2 mm long, these more like papillae, soon deciduous and the stems glabrescent; new growth glabrous to sparely papillate like the stems; bark of older stems pale tan or brown. Sympodial units difoliate, the leaves not geminate. Leaves simple and usually shallowly lobed, the blades (0.5)1.5-4.5 cm long, (0.3)0.5-2.2 cm wide, elliptic to ovate, widest at the middle or in the lower half, thick, fleshy and rubbery in texture in live plants, concolorous, extremely variable on individual plants and through the growing season; adaxial and abaxial surfaces glabrous, occasionally with a few scattered eglandular 1-2-celled simple uniseriate trichomes on the midrib; principal veins 3-5 pairs, often not visible in live or dried plants, if visible drying yellowish cream on herbarium specimens; base attenuate to truncate, always decurrent onto the petiole with a wing of leaf tissue; margins lobed, the lobes deltate, apically acute, often basiscopic (pointing towards stem), the sinuses reaching 1/4 to halfway to the midrib, revolute; apex acute; petiole 0.3-1.1 cm long, always winged with leaf tissue, glabrous or minutely puberulent with antrorse eglandular papillae. Inflorescences internodal or almost opposite the leaves, unbranched, (1)1.5-6.5 cm long, with 4-10 flowers, usually only 1-2 open at a time, glabrous or minutely puberulent with antrorse papillae like the rest of the plant; peduncle 0.5-2 cm long; pedicels 0.7-1.1 cm long, ca. 0.75 mm in diameter at the base, ca. 1.5 mm in diameter at the apex, tapering, glabrous or minutely papillate, articulated at the base; pedicel scars in pairs, each pair spaced ca. 2.5 mm apart. Buds ellipsoid, the corolla included in the calyx tube until just before anthesis due to rapid expansion of buds. Flowers 5-merous, cosexual (hermaphroditic). Calyx tube 1.5-2 mm long, conical, the lobes 2.5-4(5) mm long, 1-1.5 mm wide, long-triangular, rigid and fleshy, glabrous or minutely puberulent-papillate like the rest of the plant. Corolla 1.4-2 cm in diameter, white or pale violet, with a greenish yellow central eye edged with paler yellow, stellate, lobed ca. halfway to the base, the lobes 5-6 mm long, 2.5-4 mm wide, reflexed to spreading at anthesis, glabrous adaxially, glabrous or minutely puberulent abaxially with mixed eglandular simple uniseriate trichomes and papillae along the midvein, densely papillate at tips and margins. Stamens equal; filament tube less than 0.2 mm; free portion of the filaments ca. 1 mm long, glabrous or with a few eglandular tangled simple uniseriate trichomes to 0.5 mm long adaxially; anthers 4.5-5 mm long, 1-1.5 mm wide, ellipsoid, yellow, poricidal at the tips, the pores lengthening to slits with age. Ovary conical, glabrous; style 8.5-9 mm long, straight, exserted beyond the anther cone, minutely papillate in the lower half within the anther cone; stigma capitate, the surface minutely papillate, green in live plants. Fruit a globose berry, 0.7-1.2 cm in diameter, green or purplish green at maturity, the pericarp thin, shiny, opaque or slightly translucent, glabrous; fruiting pedicels 1-1.4 cm long, ca. 0.7 mm in diameter at the base, ca. 2 mm in diameter at the apex, spreading, not markedly woody, not persistent; fruiting calyx not accrescent, the lobes 2.5-4 mm long, 1-1.5 mm wide, spreading and slightly reflexed at the tips, fleshy and rubbery in live plants, somewhat woody in dried specimens. Seeds (5)10-20 per berry, ca. 2 mm long, 1.5-2 mm wide, reddish brown, teardrop shaped, the surfaces minutely pitted, the testal cells sinuate in outline in the seed centre, rectangular at the margins. Stone cells 10-12 per berry, 1-1.5 mm in diameter, pale creamy white. Chromosome number: n = 12 (
|
||
<bibRefCitation id="637658E491A6628642F01D0AFB9344F3" author="Moscone, EA" journalOrPublisher="Darwiniana" pageId="0" pageNumber="1" pagination="261 - 297" publicationUrl="https://www.jstor.org/stable/23222568" refId="B168" refString="Moscone, EA, 1992. Estudios en cromosomas meioticos en Solanaceae de Argentina. Darwiniana 31: 261 - 297, https://www.jstor.org/stable/23222568" title="Estudios en cromosomas meioticos en Solanaceae de Argentina." url="https://www.jstor.org/stable/23222568" volume="31" year="1992">Moscone 1992</bibRefCitation>
|
||
, voucher
|
||
<emphasis id="8C6963E797360725504323E58E1BA19E" italics="true" pageId="0" pageNumber="1">Barboza 62</emphasis>
|
||
;
|
||
<bibRefCitation id="5347380F1A471E919D43E8AD426B6BDB" DOI="https://doi.org/10.1080/00087114.2013.855389" author="Moyetta, NR" journalOrPublisher="Caryologia" pageId="0" pageNumber="1" pagination="333 - 345" refId="B169" refString="Moyetta, NR, Stiefkensa, LB, Bernardello, G, 2013. Karyotypes of South American species of the Morelloid and Dulcamaroid clades (Solanum, Solanaceae). Caryologia 66 (4): 333 - 345, DOI: https://doi.org/10.1080/00087114.2013.855389" title="Karyotypes of South American species of the Morelloid and Dulcamaroid clades (Solanum, Solanaceae)." url="https://doi.org/10.1080/00087114.2013.855389" volume="66" year="2013">Moyetta et al. 2013</bibRefCitation>
|
||
, voucher
|
||
<emphasis id="5B512E63BB9A1763B33C616317E09EB1" italics="true" pageId="0" pageNumber="1">Chiapella et al. 2630</emphasis>
|
||
, as
|
||
<taxonomicName id="FF234A6141DC64F8F231B48FC7218464" family="Solanaceae" kingdom="Plantae" lsidName="S. hastatilobum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="hastatilobum">
|
||
<emphasis id="6BCE8F84C6491B992EE0A1978F2800E0" italics="true" pageId="0" pageNumber="1">S. hastatilobum</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption id="FD9CCE35DA36741BAC688C10489FC959" doi="10.3897/phytokeys.231.100894.figure48" httpUri="https://binary.pensoft.net/fig/897509" pageId="0" pageNumber="1" start="Figure 48" startId="F48">
|
||
<paragraph id="AECAEAD858F09FDBB8BE9D45823D457A" pageId="0" pageNumber="1">
|
||
<emphasis id="A513F65A9E9CB15BF3D12C60585191E9" bold="true" pageId="0" pageNumber="1">Figure 48.</emphasis>
|
||
<taxonomicName id="14280DD9B2E9AC6259780B8A19C4A42E" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum echegarayi" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="echegarayi">
|
||
<emphasis id="D98BC49E418C15BF624A0EC7775861EB" italics="true" pageId="0" pageNumber="1">Solanum echegarayi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6A3B746B940D9B2771355881437D81B2" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
habit
|
||
<emphasis id="405D3B0D5AFD8612012842D6C6C5501E" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
underground woody rhizomes
|
||
<emphasis id="001273D2F27F8C0107359EE469B5ACA3" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
flowers at anthesis
|
||
<emphasis id="2CFC719A9AB90D693D50387FD6F38988" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
immature and mature fruits (
|
||
<emphasis id="6683CC85D2E8F7B4225C81DE268318E7" bold="true" pageId="0" pageNumber="1">A, C, D</emphasis>
|
||
<emphasis id="6072FC5F15D902C386910787A3625768" italics="true" pageId="0" pageNumber="1">Barboza et al. 4783</emphasis>
|
||
<emphasis id="C799F6EF31865060C6396B1FD9BBBE5D" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
<emphasis id="387B6733DE9445FDE08A794DF30AD9B5" italics="true" pageId="0" pageNumber="1">Knapp et al. 10540</emphasis>
|
||
). Photos by G.E. Barboza and S. Knapp.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="5773E2EA67A36369D59A832904B7CC8E" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="EADD1BEB8380CB615B99FFDA3CEAA561" pageId="0" pageNumber="1">Distribution</paragraph>
|
||
<paragraph id="91256B2F00C02D0C08D49F63C8ABC110" pageId="0" pageNumber="1">
|
||
<emphasis id="4DD53B8E8457BFDC854563D0E592625E" bold="true" pageId="0" pageNumber="1">
|
||
(Fig.
|
||
<figureCitation id="B1F2CA8F682A0D65338F0A25D09C1181" captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Distribution map of Solanum echegarayi." figureDoi="10.3897/phytokeys.231.100894.figure49" httpUri="https://binary.pensoft.net/fig/897510" pageId="0" pageNumber="1">49</figureCitation>
|
||
).
|
||
</emphasis>
|
||
<taxonomicName id="AFD40BEB740E7FD0E732ADC9FC1F9A31" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum echegarayi" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="echegarayi">
|
||
<emphasis id="D5586DD0B9AC570F06BF08DC2ADB4DDD" italics="true" pageId="0" pageNumber="1">Solanum echegarayi</emphasis>
|
||
</taxonomicName>
|
||
occurs in the Andes of central Argentina (Provs. Catamarca,
|
||
<normalizedToken id="E381FF57FEA0116B8C65769F8D97861E" originalValue="Córdoba">Cordoba</normalizedToken>
|
||
, La Rioja, Mendoza, Salta, San Juan, San Luis) and adjacent Chile (
|
||
<normalizedToken id="3C66CA6E03331DBA4B49E70F241AD4F4" originalValue="Región">Region</normalizedToken>
|
||
V [
|
||
<normalizedToken id="1201AC99BA79944D67C8CAF490C7CDFA" originalValue="Valparaíso">Valparaiso</normalizedToken>
|
||
]).
|
||
</paragraph>
|
||
<caption id="04E0F3B0A0D8B5C2E1F5A8CE6FF88769" doi="10.3897/phytokeys.231.100894.figure49" httpUri="https://binary.pensoft.net/fig/897510" pageId="0" pageNumber="1" start="Figure 49" startId="F49">
|
||
<paragraph id="BFC0EB095B3C1F48E19AEC60F2EFA170" pageId="0" pageNumber="1">
|
||
<emphasis id="EE089DB4C78E363373239099E4428FD0" bold="true" pageId="0" pageNumber="1">Figure 49.</emphasis>
|
||
Distribution map of
|
||
<taxonomicName id="93ED9D35AC9F8D029C51F7EBBA268AAD" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum echegarayi" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="echegarayi">
|
||
<emphasis id="D1119ABAA851E0568A5656E46C56DED9" italics="true" pageId="0" pageNumber="1">Solanum echegarayi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="A2227E28204D20E6944DC7E244607328" pageId="0" pageNumber="1" type="ecology">
|
||
<paragraph id="1A88DB70253D1ED10613F4AD42D2B46B" pageId="0" pageNumber="1">Ecology and habitat.</paragraph>
|
||
<paragraph id="3940C7F76F1B413C730CE4DF367A4BEF" pageId="0" pageNumber="1">
|
||
<taxonomicName id="C1DAD2CD3E20FFB046B9BD6FA519A997" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum echegarayi" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="echegarayi">
|
||
<emphasis id="D2DAF38AB40D4285198AEAE7ED237141" italics="true" pageId="0" pageNumber="1">Solanum echegarayi</emphasis>
|
||
</taxonomicName>
|
||
grows in dry, scrubby habitats, usually at high elevation, and in open rocky areas, often where little other vegetation occurs, from 650 to 4,200 m elevation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9438F601D7FF07913D42DE50F5F05547" pageId="0" pageNumber="1" type="common names and uses">
|
||
<paragraph id="2C5E7F078036F77E9C7B2E2D43F10443" pageId="0" pageNumber="1">Common names and uses.</paragraph>
|
||
<paragraph id="EBFE7FDA3C1988D25486C77BC3929A9C" pageId="0" pageNumber="1">None recorded.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0970B019CD932363B394021098B62696" pageId="0" pageNumber="1" type="preliminary conservation status">
|
||
<paragraph id="3BA944D0AA62DFC20587E6C995A71A52" pageId="0" pageNumber="1">Preliminary conservation status</paragraph>
|
||
<paragraph id="2E4964996D5E0C7B6B95BF698448C029" pageId="0" pageNumber="1">
|
||
<emphasis id="C400A90ABABD3F6B45139FAA7731C45D" bold="true" pageId="0" pageNumber="1">
|
||
(
|
||
<bibRefCitation id="F43DC24120CDB13FA9DC376F98971267" author="Ibarrola, DA" journalOrPublisher="Nueva Serie" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org/resources/redlistguidelines" refId="B116" refString="2022. . https://www.iucnredlist.org/resources/redlistguidelines" url="https://www.iucnredlist.org/resources/redlistguidelines" year="2022">IUCN 2022</bibRefCitation>
|
||
).
|
||
</emphasis>
|
||
Least Concern [LC]. EOO = 352,787 km2 [LC]; AOO = 408 km2 [EN].
|
||
<taxonomicName id="3442213DBEF67C165465BCC8DA3F6127" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum echegarayi" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="echegarayi">
|
||
<emphasis id="122559A881D7838C4C1C8A49E46F0A5A" italics="true" pageId="0" pageNumber="1">Solanum echegarayi</emphasis>
|
||
</taxonomicName>
|
||
is widespread along the Andes and occurs in disturbed habitats. It is found in several protected areas throughout its range in Argentina (e.g., Parque Nacional Talampaya, Parque Provincial
|
||
<normalizedToken id="94A9505085B41372E1EF24018B020A63" originalValue="Volcán">Volcan</normalizedToken>
|
||
Tupungato, Parque Nacional El Leoncito).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BD2D1FC5893435F78DD47E94EFCC5B23" pageId="0" pageNumber="1" type="discussion">
|
||
<paragraph id="6F9A8035E81505E3D5DDD0FD5A7B6702" pageId="0" pageNumber="1">Discussion.</paragraph>
|
||
<paragraph id="CAF4C15D2BFCD2AE6ED5F2C30BBDF0C1" pageId="0" pageNumber="1">
|
||
<taxonomicName id="47275E42B06E9443DE8926A3902A1600" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum echegarayi" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="echegarayi">
|
||
<emphasis id="AAD783FAB5837E156A44200C7D26A2AB" italics="true" pageId="0" pageNumber="1">Solanum echegarayi</emphasis>
|
||
</taxonomicName>
|
||
is a fleshy, almost succulent plant with deep woody rhizomes from which new shoots arise every growing season. It is a member of the
|
||
<taxonomicName id="5CB6B3685A85DE8A6219B120CB640891" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Episarcophyllum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Episarcophyllum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="4F6F4D29FF390103BF76B6DC36383A32" italics="true" pageId="0" pageNumber="1">Episarcophyllum</emphasis>
|
||
</taxonomicName>
|
||
clade (
|
||
<bibRefCitation id="DD0323E95307DDB6599F44ED58333306" DOI="https://doi.org/10.12705/645.5" author="Saerkinen, T" journalOrPublisher="Taxon" pageId="0" pageNumber="1" pagination="945 - 958" refId="B206" refString="Saerkinen, T, Barboza, GE, Knapp, S, 2015b. True Black nightshades: Phylogeny and delimitation of the Morelloid clade of Solanum. Taxon 64 (5): 945 - 958, DOI: https://doi.org/10.12705/645.5" title="True Black nightshades: Phylogeny and delimitation of the Morelloid clade of Solanum." url="https://doi.org/10.12705/645.5" volume="64" year="2015 b">
|
||
<normalizedToken id="BF37E58133254C41635DFB189FEB7487" originalValue="Särkinen">Saerkinen</normalizedToken>
|
||
et al. 2015b
|
||
</bibRefCitation>
|
||
) together with
|
||
<taxonomicName id="3F14A105486C3961CA5E57D044503E8D" family="Solanaceae" kingdom="Plantae" lsidName="S. riojense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="riojense">
|
||
<emphasis id="D8BFE1FB8A63B0E056982C306EAD2F58" italics="true" pageId="0" pageNumber="1">S. riojense</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="EB6D29B940F5687A5A2A400A374B8479" family="Solanaceae" kingdom="Plantae" lsidName="S. sinuatirecurvum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="sinuatirecurvum">
|
||
<emphasis id="FACF79DFE2379EAB0E1FEB54CA0C1FE1" italics="true" pageId="0" pageNumber="1">S. sinuatirecurvum</emphasis>
|
||
</taxonomicName>
|
||
, but not related to
|
||
<taxonomicName id="F7397F4BF5B6824AED407C2783BE7375" family="Solanaceae" kingdom="Plantae" lsidName="S. caesium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="caesium">
|
||
<emphasis id="86E60F3A72C012E2CFFC9A1EE852B75D" italics="true" pageId="0" pageNumber="1">S. caesium</emphasis>
|
||
</taxonomicName>
|
||
which has previously (
|
||
<bibRefCitation id="53A18E81BF888FE3A154E7261D3835A9" author="Del Vitto, LA" journalOrPublisher="Kurtziana" pageId="0" pageNumber="1" pagination="319 - 326" refId="B65" refString="Del Vitto, LA, Petenatti, FM, 1999. Notas en Solanum (Solanaceae) de Argentina II. Aportes al conocimiento de la Sect. Episarcophyllum. Kurtziana 27: 319 - 326" title="Notas en Solanum (Solanaceae) de Argentina II. Aportes al conocimiento de la Sect. Episarcophyllum." volume="27" year="1999">Del Vitto and Petenatti 1999</bibRefCitation>
|
||
) been thought to have affinities to these taxa (
|
||
<bibRefCitation id="A19541F9A4FB1CA4854BADF442AF85F5" DOI="https://doi.org/10.1002/ajb2.1827" author="Gagnon, E" journalOrPublisher="American Journal of Botany" pageId="0" pageNumber="1" pagination="580 - 601" refId="B92" refString="Gagnon, E, Hilgenhof, R, Orejuela, A, McDonnell, A, Sablok, G, Aubriot, X, Giacomin, LL, Gouvea, Y, Bohs, L, Dodsworth, S, Martine, C, Poczai, P, Knapp, S, Saerkinen, T, 2022. Phylogenomic discordance suggests polytomies along the backbone of the large genus Solanum. American Journal of Botany 109 (4): 580 - 601, DOI: https://doi.org/10.1002/ajb2.1827" title="Phylogenomic discordance suggests polytomies along the backbone of the large genus Solanum." url="https://doi.org/10.1002/ajb2.1827" volume="109" year="2022">Gagnon et al. 2022</bibRefCitation>
|
||
). All species of the
|
||
<taxonomicName id="61E026DD130AF2439E930F22DCD2121D" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Episarcophyllum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Episarcophyllum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="364F245205993ABB6631669D1B9AE95C" italics="true" pageId="0" pageNumber="1">Episarcophyllum</emphasis>
|
||
</taxonomicName>
|
||
clade are perennial herbs with woody underground rhizomes and occur in dry habitats in Argentina and neighbouring Chile, generally above 2,000 m elevation. The species all have slightly thick and fleshy leaves that appear succulent when compared to other species of the Morelloid clade.
|
||
</paragraph>
|
||
<paragraph id="999FD0CBF8AA725FDBDECA246A43B589" pageId="0" pageNumber="1">
|
||
<taxonomicName id="3E920709679B63C2C1FE959FB46C3C43" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum echegarayi" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="echegarayi">
|
||
<emphasis id="3F7FB76550AE0C245D66C685DC21CB35" italics="true" pageId="0" pageNumber="1">Solanum echegarayi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8F6E5FAB317117E7FF39843B06C0A6E4" family="Solanaceae" kingdom="Plantae" lsidName="S. riojense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="riojense">
|
||
<emphasis id="6A6314816790A5B65F0A4E1A67E02F15" italics="true" pageId="0" pageNumber="1">S. riojense</emphasis>
|
||
</taxonomicName>
|
||
have long been confused due to a mix-up of type specimens (see below).
|
||
<taxonomicName id="4A2AE6BBC6060FC70208498CD299E18F" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum echegarayi" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="echegarayi">
|
||
<emphasis id="8A452C5E78324717A4815BCD3FC32E20" italics="true" pageId="0" pageNumber="1">Solanum echegarayi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="D41627F9CB035611AC5FCEFDAEA6B1F8" family="Solanaceae" kingdom="Plantae" lsidName="S. riojense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="riojense">
|
||
<emphasis id="681C93F1ACE069E903095B0DCE20BE75" italics="true" pageId="0" pageNumber="1">S. riojense</emphasis>
|
||
</taxonomicName>
|
||
in its lack of cobwebby, tangled trichomes and in its sharply pointed rather than rounded calyx lobe apices.
|
||
<taxonomicName id="E6F7D1C497ACB5342EDA80C09786CA33" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum sinuatirecurvum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sinuatirecurvum">
|
||
<emphasis id="8E269100B2709DA796F4C800E5852B2C" italics="true" pageId="0" pageNumber="1">Solanum sinuatirecurvum</emphasis>
|
||
</taxonomicName>
|
||
also has cobwebby trichomes and differs from
|
||
<taxonomicName id="EADFD2E39345267BE63C36C18AD4C7FA" family="Solanaceae" kingdom="Plantae" lsidName="S. echegarayi" order="Solanales" pageId="0" pageNumber="1" rank="species" species="echegarayi">
|
||
<emphasis id="E87FD7AE9B66FC8D93E1F42D6C7EB558" italics="true" pageId="0" pageNumber="1">S. echegarayi</emphasis>
|
||
</taxonomicName>
|
||
in its much larger berries (more than 1 cm in diameter versus usually less than 1 cm in diameter) with a yellow, leathery pericarp rather than a green to greenish purple, somewhat translucent pericarp.
|
||
</paragraph>
|
||
<paragraph id="BABF7912A020B975B9F60C34B661DE0B" pageId="0" pageNumber="1">
|
||
<taxonomicName id="210E5A1D131360297F8A4CE496E4B2D4" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum echegarayi" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="echegarayi">
|
||
<emphasis id="450AC95F107F7A327DE37E966F2DFCEC" italics="true" pageId="0" pageNumber="1">Solanum echegarayi</emphasis>
|
||
</taxonomicName>
|
||
is very variable depending upon when in the growing season the plant is collected; plants from early in the season are quite small and can look markedly different from those collected later in the season. In addition, specimens are often collected without the deep rhizomes, and so have the appearance of ephemeral annuals. Plants arise from deep underground stems (see Figs
|
||
<figureCitation id="1BB8425225E5B8C8D746C673C93C7FD4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Representative habits and leaves of South American morelloids A prostrate annual herbs (S. weddellii) B large lax shrubs (S. aloysiifolium) C roots forming along a creeping stem (S. juninense) D spinose processes on stems of many species (S. huayavillense) E glandular trichomes found in some members of the clade (S. glandulosipilosum) F dendritic / branched trichomes found in S. pallidum G regularly 3 - lobed leaves in S. palitans H highly variable leaves along a single stem in S. salicifolium (A Saerkinen et al. 4038 B Barboza et al. 3505 C Saerkinen et al. 4754 D Barboza et al. 3536 E Barboza et al. 3520 F Saerkinen et al. 4010 G Atchison & Gagnon 25 H Barboza et al. 3473). Photos by S. Knapp, G. Atchison, and T. Saerkinen." figureDoi="10.3897/phytokeys.231.100894.figure2" httpUri="https://binary.pensoft.net/fig/897463" pageId="0" pageNumber="1">2C</figureCitation>
|
||
,
|
||
<figureCitation id="BBA13F103FFAEF85780F37F78C0F1451" captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Solanum echegarayi A habit B underground woody rhizomes C flowers at anthesis D immature and mature fruits (A, C, D Barboza et al. 4783 B Knapp et al. 10540). Photos by G. E. Barboza and S. Knapp." figureDoi="10.3897/phytokeys.231.100894.figure48" httpUri="https://binary.pensoft.net/fig/897509" pageId="0" pageNumber="1">48B</figureCitation>
|
||
) and the junction of vegetative shoots and rhizomes is quite fragile. Characters used to distinguish the species here recognised as synonyms of
|
||
<taxonomicName id="9F4FCF81279A9626832FE2EBCFE3AA5A" family="Solanaceae" kingdom="Plantae" lsidName="S. echegarayi" order="Solanales" pageId="0" pageNumber="1" rank="species" species="echegarayi">
|
||
<emphasis id="A5A1EA999763471686550F24E5F27FD4" italics="true" pageId="0" pageNumber="1">S. echegarayi</emphasis>
|
||
</taxonomicName>
|
||
that were regarded as distinct taxa by
|
||
<bibRefCitation id="4B9098F71D03F9CD9A342B2C9A6AC825" author="Del Vitto, LA" journalOrPublisher="Kurtziana" pageId="0" pageNumber="1" pagination="319 - 326" refId="B65" refString="Del Vitto, LA, Petenatti, FM, 1999. Notas en Solanum (Solanaceae) de Argentina II. Aportes al conocimiento de la Sect. Episarcophyllum. Kurtziana 27: 319 - 326" title="Notas en Solanum (Solanaceae) de Argentina II. Aportes al conocimiento de la Sect. Episarcophyllum." volume="27" year="1999">Del Vitto and Petenatti (1999)</bibRefCitation>
|
||
overlap broadly across the species range; plants from northern part of the range from lower elevations (described as
|
||
<taxonomicName id="66BCD14343FD86C6336D9AD7E0AD0B04" family="Solanaceae" kingdom="Plantae" lsidName="S. hastatilobatum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="hastatilobatum">
|
||
<emphasis id="36998898AFD5F1B78A4F038DFB386824" italics="true" pageId="0" pageNumber="1">S. hastatilobatum</emphasis>
|
||
</taxonomicName>
|
||
) tend to have smaller fruits that are green or purple and more hastate leaves, but all intermediate variations occur (for example, the collections
|
||
<emphasis id="8DF4D8A2EA1DEDA1F000A6510D732BD9" italics="true" pageId="0" pageNumber="1">Barboza et al. 3447, 3450</emphasis>
|
||
from high elevation in La Rioja Province match these plants to some degree). The highest degree of variation occurs in the Province of San Juan, where intensive study of climatic and environmental conditions will prove useful in untangling patterns of vegetative variation. Molecular sequence data suggest there is some regional variation in
|
||
<taxonomicName id="B3329137B1CD55C694F72D16C7493455" family="Solanaceae" kingdom="Plantae" lsidName="S. echegarayi" order="Solanales" pageId="0" pageNumber="1" rank="species" species="echegarayi">
|
||
<emphasis id="3546592BBEAEC7ECF0E037585105B3FC" italics="true" pageId="0" pageNumber="1">S. echegarayi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF6D7E01515528AF176170C0A2160DD0" DOI="https://doi.org/10.1002/ajb2.1827" author="Gagnon, E" journalOrPublisher="American Journal of Botany" pageId="0" pageNumber="1" pagination="580 - 601" refId="B92" refString="Gagnon, E, Hilgenhof, R, Orejuela, A, McDonnell, A, Sablok, G, Aubriot, X, Giacomin, LL, Gouvea, Y, Bohs, L, Dodsworth, S, Martine, C, Poczai, P, Knapp, S, Saerkinen, T, 2022. Phylogenomic discordance suggests polytomies along the backbone of the large genus Solanum. American Journal of Botany 109 (4): 580 - 601, DOI: https://doi.org/10.1002/ajb2.1827" title="Phylogenomic discordance suggests polytomies along the backbone of the large genus Solanum." url="https://doi.org/10.1002/ajb2.1827" volume="109" year="2022">Gagnon et al. 2022</bibRefCitation>
|
||
) but further sampling of all three species in this small clade (e.g.,
|
||
<taxonomicName id="A7187032C5C5CA86CC3E9C711BD82FD6" family="Solanaceae" kingdom="Plantae" lsidName="S. echegarayi" order="Solanales" pageId="0" pageNumber="1" rank="species" species="echegarayi">
|
||
<emphasis id="44D540AEAF6FB894880137562B3B090A" italics="true" pageId="0" pageNumber="1">S. echegarayi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="233F30ED5CE0302B479B1D2101A791AF" family="Solanaceae" kingdom="Plantae" lsidName="S. riojense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="riojense">
|
||
<emphasis id="F94F92706DE5A5105DE15C2040FE7FB8" italics="true" pageId="0" pageNumber="1">S. riojense</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="115BF07FC1B475DEE3343FD4136C9C74" family="Solanaceae" kingdom="Plantae" lsidName="S. sinuatirecurvum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="sinuatirecurvum">
|
||
<emphasis id="8B87DA0E78DF87BD5AFB1CBA57257DD8" italics="true" pageId="0" pageNumber="1">S. sinuatirecurvum</emphasis>
|
||
</taxonomicName>
|
||
) will be necessary to confirm these results.
|
||
</paragraph>
|
||
<paragraph id="655D564246287519828B6E46CCA2AD4D" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="72B10A3971F78D9FD4331A2C6D0A9729" author="Morton, CV" journalOrPublisher="Biodiversity and Conservation" pageId="0" pageNumber="1" refId="B166" refString="Morton, CV, 1976. A revision of the Argentine species of Solanum. Academia Nacional de Ciencias, Cordoba." title="A revision of the Argentine species of Solanum. Academia Nacional de Ciencias, Cordoba." year="1976">Morton (1976)</bibRefCitation>
|
||
lectotypified
|
||
<taxonomicName id="3C6E1823A5AE9AB5F09D4A6050A71AF0" family="Solanaceae" kingdom="Plantae" lsidName="S. echegarayi" order="Solanales" pageId="0" pageNumber="1" rank="species" species="echegarayi">
|
||
<emphasis id="384B7F50343E570B3C775A49E0981B40" italics="true" pageId="0" pageNumber="1">S. echegarayi</emphasis>
|
||
</taxonomicName>
|
||
with a specimen in CORD (CORD00012856) labelled "Echegaray 472", indicating the type in B was destroyed. Hieronymus used only specimens at CORD in his description (as clearly noted in the introduction to his catalogue of
|
||
<normalizedToken id="0657F0EAE3D407430300CAD70B109C5F" originalValue="Echegaray’s">Echegaray's</normalizedToken>
|
||
collections;
|
||
<bibRefCitation id="9B143BEF71CC1EEE4D9140C1924A047A" DOI="https://doi.org/10.5962/bhl.title.9016" author="Hieronymus, G" journalOrPublisher="Boletin de la Academia Nacional de Ciencias en Cordoba" pageId="0" pageNumber="1" pagination="1 - 108" refId="B106" refString="Hieronymus, G, 1881. Sertum Sanjuaninum o descripciones y determinaciones de plantas faner o gamas y cript o gamas vasculares recolectadas por el Dr. D. Saile Echegaray en la Provincia de San Juan. Boletin de la Academia Nacional de Ciencias en Cordoba 4: 1 - 108, DOI: https://doi.org/10.5962/bhl.title.9016" title="Sertum Sanjuaninum o descripciones y determinaciones de plantas faner o gamas y cript o gamas vasculares recolectadas por el Dr. D. Saile Echegaray en la Provincia de San Juan." url="https://doi.org/10.5962/bhl.title.9016" volume="4" year="1881">Hieronymus 1881</bibRefCitation>
|
||
);
|
||
<bibRefCitation id="F8D5A8BF23E524B710BE67FB28994DDC" author="Morton, CV" journalOrPublisher="Biodiversity and Conservation" pageId="0" pageNumber="1" refId="B166" refString="Morton, CV, 1976. A revision of the Argentine species of Solanum. Academia Nacional de Ciencias, Cordoba." title="A revision of the Argentine species of Solanum. Academia Nacional de Ciencias, Cordoba." year="1976">
|
||
<normalizedToken id="68C4C8D6B4DD8FA05EB813D2C0AD3ECE" originalValue="Morton’s">Morton's</normalizedToken>
|
||
(1976)
|
||
</bibRefCitation>
|
||
lectotypification is thus superfluous. In addition, the specimen selected by
|
||
<bibRefCitation id="65CB10A08477E7379975984E49F8040A" author="Morton, CV" journalOrPublisher="Biodiversity and Conservation" pageId="0" pageNumber="1" refId="B166" refString="Morton, CV, 1976. A revision of the Argentine species of Solanum. Academia Nacional de Ciencias, Cordoba." title="A revision of the Argentine species of Solanum. Academia Nacional de Ciencias, Cordoba." year="1976">Morton (1976)</bibRefCitation>
|
||
is densely pubescent with cobwebby hairs on the new growth and does not match the protologue, where the plant is described as completely glabrous; this specimen is likely a label mix-up for
|
||
<emphasis id="D39EA63DE2B38D986FA21ADF5CB45F54" italics="true" pageId="0" pageNumber="1">Hieronymous & Neiderlein 472</emphasis>
|
||
, a syntype of
|
||
<taxonomicName id="3E132EEFC8D4299784356868BD8BC61D" family="Solanaceae" kingdom="Plantae" lsidName="S. riojense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="riojense">
|
||
<emphasis id="29C8A5E34D5C9102A8257FA5CE3096DB" italics="true" pageId="0" pageNumber="1">S. riojense</emphasis>
|
||
</taxonomicName>
|
||
(see discussion under
|
||
<taxonomicName id="31B8EFAD06905405C2F4510191569442" family="Solanaceae" kingdom="Plantae" lsidName="S. riojense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="riojense">
|
||
<emphasis id="72A4061C0E544E7F765D30E28A9CABFE" italics="true" pageId="0" pageNumber="1">S. riojense</emphasis>
|
||
</taxonomicName>
|
||
). The sheets labelled "Echegaray 472" at CORD and SI (CORD [CORD00012856], SI [003309]) are therefore excluded from consideration as type material of
|
||
<taxonomicName id="8E8CCAD94DFF726C5ABA9917B65A9B1C" family="Solanaceae" kingdom="Plantae" lsidName="S. echegarayi" order="Solanales" pageId="0" pageNumber="1" rank="species" species="echegarayi">
|
||
<emphasis id="8B300AE9F315A8B889DD2E85ABECB298" italics="true" pageId="0" pageNumber="1">S. echegarayi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2BDAE48587270CFC56FB266E5355B675" pageId="0" pageNumber="1">
|
||
<taxonomicName id="C62F0BE1EC00C8F20DDA6A9115D0552B" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum juncalense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="juncalense">
|
||
<emphasis id="D96E9810403EBD13579341DE6D86D5FC" italics="true" pageId="0" pageNumber="1">Solanum juncalense</emphasis>
|
||
</taxonomicName>
|
||
was described from material from "Cordilleras de la provincia de Aconcagua (Juncal, 2,200 m)", with no collector or herbarium cited. A specimen in SGO (SGO000004574) from [Nevado] Juncal and the same elevation (
|
||
<emphasis id="0B2B547DDD0ED540714DCF689161E7FC" italics="true" pageId="0" pageNumber="1">Buchtien 150</emphasis>
|
||
) and annotated "S. juncalense R" is almost certainly original material and is here selected as the neotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |