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<document ID-CLB-Dataset="26095" ID-DOI="http://dx.doi.org/10.3897/zookeys.870.36970" ID-GBIF-Dataset="0c6fd07e-55da-42a2-85d1-4f1068abc419" ID-PMC="PMC6694067" ID-Pensoft-Pub="1313-2970-870-51" ID-Pensoft-UUID="B9AE5951AA745D6BAF3EEB40C2061AF4" ID-PubMed="31423078" ID-ZooBank="BEC530D3DD9A466AAAC705305B20E761" ModsDocID="1313-2970-870-51" checkinTime="1565240931249" checkinUser="pensoft" docAuthor="Sutcharit, Chirasak, Backeljau, Thierry &amp; Panha, Somsak" docDate="2019" docId="E4E7297D8C905B458E5741CCE3C20E43" docLanguage="en" docName="ZooKeys 870: 51-76" docOrigin="ZooKeys 870" docSource="http://dx.doi.org/10.3897/zookeys.870.36970" docTitle="Ganesella carinella Moellendorff 1902" docType="treatment" docVersion="6" id="B9AE5951AA745D6BAF3EEB40C2061AF4" lastPageNumber="51" masterDocId="B9AE5951AA745D6BAF3EEB40C2061AF4" masterDocTitle="Re-description of the type species of the genera Ganesella Blanford, 1863 and Globotrochus Haas, 1935; with description of a new Ganesella species from Thailand (Eupulmonata, Camaenidae)" masterLastPageNumber="76" masterPageNumber="51" pageNumber="51" updateTime="1732748597855" updateUser="ExternalLinkService">
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<mods:title id="8E760701C20FD68601AA3DE35719A7C7">Re-description of the type species of the genera Ganesella Blanford, 1863 and Globotrochus Haas, 1935; with description of a new Ganesella species from Thailand (Eupulmonata, Camaenidae)</mods:title>
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<mods:namePart id="011A24917C6BDD5C19A51EEDF2537E88">Sutcharit, Chirasak</mods:namePart>
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<mods:namePart id="A34022AD20F8E32118700E135142711A">Backeljau, Thierry</mods:namePart>
</mods:name>
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<mods:namePart id="8B754BC86893D942587DB595B8759FC6">Panha, Somsak</mods:namePart>
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<mods:typeOfResource id="FF380888BB281B4AD8132AFE1B7B6B2A">text</mods:typeOfResource>
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<mods:title id="6B9AE58791821226A99CF9E4B1CA84AB">ZooKeys</mods:title>
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<mods:date id="1E80E97B2BA34A23D5B087ADEDF7DCD5">2019</mods:date>
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<mods:number id="4E98E186217DA812F7CEAD11AC8FAE33">870</mods:number>
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<mods:start id="4690F73D891E7BA926257C81D59515C7">51</mods:start>
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<mods:classification id="0A2E471642EF1EB9BB45B76460DB1549">journal article</mods:classification>
<mods:identifier id="7E221590B4666FF667DBB3778A4859F4" type="DOI">http://dx.doi.org/10.3897/zookeys.870.36970</mods:identifier>
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<treatment id="E4E7297D8C905B458E5741CCE3C20E43" ID-GBIF-Taxon="159043263" LSID="urn:lsid:plazi:treatment:E4E7297D8C905B458E5741CCE3C20E43" httpUri="http://treatment.plazi.org/id/E4E7297D8C905B458E5741CCE3C20E43" lastPageNumber="51" pageId="0" pageNumber="51" scope_family="Camaenidae" scope_superOrder="Eupulmonata">
<subSubSection id="3CF439DC98F1C2651EC5976A28C8383B" pageId="0" pageNumber="51" type="nomenclature">
<paragraph id="3675F0A2486430FE1DF15643F93C96AD" pageId="0" pageNumber="51">
<taxonomicName id="6A461E4A0597F6BE6AD036E56D3727BE" ID-CoL="7RDBD" LSID="e4e7297d-8c90-5b45-8e57-41cce3c20e43" authority="Moellendorff, 1902" authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella carinella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="carinella">
Ganesella carinella (
<normalizedToken id="BA0641076E82AA53A0566E502789C751" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1902)
</taxonomicName>
<figureCitation id="0AC915288B17152D3F6A20E83DFC92B6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A, B Ganesella carinella from Keng-khoy, Saraburi (shell width about 15 mm) A live snail and B snails aestivated under loose tree bark, and with white epiphrams attached on substrate C Ganesella rhombostoma from Klong Had, Srakeo (shell height about 15 mm) D Globotrochus onestera from Cuc Phuong, Vietnam (shell width about 15 mm)." figureDoi="10.3897/zookeys.870.36970.figure1" httpUri="https://binary.pensoft.net/fig/323906" pageId="0" pageNumber="51">Figs 1A, B</figureCitation>
,
<figureCitation id="6A1E8920F3416E51DEF68976DA9737C3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A, B Ganesella capitium A lectotype UMZC I. 102385 / 1 and B paralectotype UMZC I. 102385 / 2 - 3 C-E Ganesella hariola C lectotype UMZC I. 104370 / 1 D paralectotype UMZC I. 104370 / 2 - 4 and E shell from Popa Mountain, Myanmar CUMZ 5134 F-H Ganesella carinella F lectotype SMF 27534 a G paralectotype SMF 27534 b and H shell from Takhli, Nakhonsawan CUMZ 5133." figureDoi="10.3897/zookeys.870.36970.figure2" httpUri="https://binary.pensoft.net/fig/323907" pageId="0" pageNumber="51">
2
<normalizedToken id="5D649399F94CBBE197C8B68AA1C5D174" originalValue="F–H">F-H</normalizedToken>
</figureCitation>
,
<figureCitation id="72464AD130B67F2910C15388BFC8FDC3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of &quot; harryleei Thach, 2017 &quot; from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">3A, B</figureCitation>
,
<figureCitation id="E0D932BFA74A7E6102D33CCDA04B5992" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">
5
<normalizedToken id="126A5D10C2BF85062B02F72116E726C6" originalValue="A–D">A-D</normalizedToken>
</figureCitation>
,
<figureCitation id="924B7D43A1227770A718D820394EE8D7" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Radula. A-C Ganesella carinella D-F Ganesella rhombostoma and G-I Globotrochus onestera A, D, G central tooth with the first to the second lateral teeth B, E, H lateral teeth with the tricuspid marginal teeth transition C, F, I outermost marginal teeth. Numbers indicate order of lateral and marginal teeth. Central tooth indicated by ' C'." figureDoi="10.3897/zookeys.870.36970.figure7" httpUri="https://binary.pensoft.net/fig/323912" pageId="0" pageNumber="51">
7
<normalizedToken id="DB01307CB4B106E0C40915A18F67A0B7" originalValue="A–C">A-C</normalizedToken>
</figureCitation>
,
<figureCitation id="FAE052552AB96D8E3E15BC8599742387" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic distribution of Ganesella spp. and Globotrochus spp." figureDoi="10.3897/zookeys.870.36970.figure8" httpUri="https://binary.pensoft.net/fig/323913" pageId="0" pageNumber="51">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CBD5162ECF435350D3363B4383199961" pageId="0" pageNumber="51" type="reference_group">
<paragraph id="6BA1AF23C2607D576F1E62D7FDE66377" pageId="0" pageNumber="51">
<taxonomicName id="B1DFC2D9622D5CCAD5E7CA8406945BE5" class="Gastropoda" family="Camaenidae" genus="Eulota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eulota (Ganesella) hariola" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="subspecies" species="hariola" subGenus="Ganesella" subspecies="carinella">Eulota (Ganesella) hariola carinella</taxonomicName>
<normalizedToken id="7E0B8CB01596D72B58890A78E30EA71E" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1902: 158, 159.
<bibRefCitation id="CA9CE97B1A557378E152458BBC5D997A" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="51" pagination="197 - 223" refId="B72" refString="Zilch, A, 1966. Die Typen und Typoide des Natur-Museums Senckenberg 34: Mollusca, Camaenidae (4). . Archiv fuer Molluskenkunde 95: 197 - 223" title="Die Typen und Typoide des Natur-Museums Senckenberg 34: Mollusca, Camaenidae (4)." volume="95" year="1966">Zilch 1966</bibRefCitation>
: 202.
</paragraph>
<paragraph id="5DBEC1B2F5DA62EB9764C443210DC1BD" pageId="0" pageNumber="51">
<taxonomicName id="A9E3EFC9FC3D68C6B07199F0DE2D591B" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella capitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="capitium">Ganesella capitium</taxonomicName>
[non
<bibRefCitation id="D886A8F61978E7C55F2E674E1A4B4F1B" DOI="https://doi.org/10.1080/03745485809494679" author="Benson, WH" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="51" pagination="158 - 164" refId="B7" refString="Benson, WH, 1848. Characters of seven new species of Helix, with amended descriptions of some species previously described, and notes on other requiring remark. . Proceedings of the Zoological Society of London 16: 158 - 164" title="Characters of seven new species of Helix, with amended descriptions of some species previously described, and notes on other requiring remark." url="https://doi.org/10.1080/03745485809494679" volume="16" year="1848">Benson 1848</bibRefCitation>
]:
<bibRefCitation id="7BD67B48494696DB3C48FF6E911E3880" author="Pilsbry, HA" journalOrPublisher="The Academy of Natural Sciences of Philadelphia, PA" pageId="0" pageNumber="51" refId="B48" refString="Pilsbry, HA, 1895. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Series 2 volume 9. . The Academy of Natural Sciences of Philadelphia, PA" title="Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Series 2 volume 9." year="1895">Pilsbry 1895</bibRefCitation>
: 170, 360, pl. 55, fig. 18.
<bibRefCitation id="071DBCA878A5383CC3850D42F4AAA833" DOI="https://doi.org/10.1093/oxfordjournals.mollus.a065982" author="Blanford, WT" journalOrPublisher="Proceeding of the Malacological Society of London" pageId="0" pageNumber="51" pagination="274 - 284" refId="B11" refString="Blanford, WT, 1903. Note on Mr. W.M. Daly's collections of land and freshwater Mollusca from Siam. . Proceeding of the Malacological Society of London 5: 274 - 284" title="Note on Mr. W. M. Daly's collections of land and freshwater Mollusca from Siam." url="https://doi.org/10.1093/oxfordjournals.mollus.a065982" volume="5" year="1903">Blanford 1903</bibRefCitation>
: 278.
<bibRefCitation id="6BFDDC580D43CCCE7CA5CB51DD3B4DC5" DOI="https://doi.org/10.5962/bhl.title.12891" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="51" refId="B22" refString="Gude, GK, 1914. Mollusca II (Trochomorphidae-Janellidae). In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, Including Ceylon and Burma. . Taylor and Francis, London" title="Mollusca II (Trochomorphidae-Janellidae). In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, Including Ceylon and Burma." url="https://doi.org/10.5962/bhl.title.12891" year="1914">Gude 1914</bibRefCitation>
: 196, 197.
<bibRefCitation id="D9FBCA003215BC9C20981E0B706F0F65" pageId="0" pageNumber="51" refId="B71">Zilch 1960</bibRefCitation>
: 610, fig. 2140.
<bibRefCitation id="F39203D4C776A7F6526F92A70D33AA1F" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="51" pagination="197 - 223" refId="B72" refString="Zilch, A, 1966. Die Typen und Typoide des Natur-Museums Senckenberg 34: Mollusca, Camaenidae (4). . Archiv fuer Molluskenkunde 95: 197 - 223" title="Die Typen und Typoide des Natur-Museums Senckenberg 34: Mollusca, Camaenidae (4)." volume="95" year="1966">Zilch 1966</bibRefCitation>
: 202.
<bibRefCitation id="526F37286859F1FEAEBA06CEBA2A0363" author="Richardson, L" journalOrPublisher="Tryonia" pageId="0" pageNumber="51" pagination="1 - 479" refId="B52" refString="Richardson, L, 1985. Camaenidae: Catalogue of Species. . Tryonia 12: 1 - 479" title="Camaenidae: Catalogue of Species." volume="12" year="1985">Richardson 1985</bibRefCitation>
: 132, 133.
<bibRefCitation id="78AF055746FB07D112CD96169571FF08" pageId="0" pageNumber="51" refId="B54">Schileyko 2003</bibRefCitation>
: fig. 1958.
</paragraph>
</subSubSection>
<subSubSection id="6E3D26FDC6B5E9A6F2AD56769C171353" pageId="0" pageNumber="51" type="type locality">
<paragraph id="52A697E503DF058FF18FA12EDEDEC2C3" pageId="0" pageNumber="51">Type locality.</paragraph>
<paragraph id="1CFCAE2DEDF305B1B63D21C4A0F299A3" pageId="0" pageNumber="51">Siam, Muoklek and Kanburi [Thailand: Muaklek District, Saraburi Province and Kanchanaburi Province].</paragraph>
<caption id="9CABFF86334BCE1C21B8A91FECBA4BA3" doi="10.3897/zookeys.870.36970.figure1" httpUri="https://binary.pensoft.net/fig/323906" pageId="0" pageNumber="51" start="Figure 1" startId="F1">
<paragraph id="8F4CBF8F013288ACA3F6CEDFE2061401" pageId="0" pageNumber="51">
Figure 1.
<emphasis id="48D9D2A5862D47836798BF6E601B88F6" bold="true" pageId="0" pageNumber="51">A, B</emphasis>
<taxonomicName id="307F1431174A22C0A97E61B4AE320A90" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella carinella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="carinella">
<emphasis id="FCDF28C437A0791172E63176B573A979" italics="true" pageId="0" pageNumber="51">Ganesella carinella</emphasis>
</taxonomicName>
from Keng-khoy, Saraburi (shell width about 15 mm)
<emphasis id="D7FAE4C9442777BE6E66ED94BAEF4057" bold="true" pageId="0" pageNumber="51">A</emphasis>
live snail and
<emphasis id="12BB030FB94C62DD66B74E61640E586B" bold="true" pageId="0" pageNumber="51">B</emphasis>
snails aestivated under loose tree bark, and with white epiphrams attached on substrate
<emphasis id="52289232F5E9D061F6B06A2810882F97" bold="true" pageId="0" pageNumber="51">C</emphasis>
<taxonomicName id="FCC8C7BC984FCD59AFBF5E1E8D7FF4D9" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella rhombostoma" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="rhombostoma">
<emphasis id="358096C72C27020F1EE30822EC9F8BF4" italics="true" pageId="0" pageNumber="51">Ganesella rhombostoma</emphasis>
</taxonomicName>
from Klong Had, Srakeo (shell height about 15 mm)
<emphasis id="5B13071C7BA8CD2F0D725EE0A4A6469F" bold="true" pageId="0" pageNumber="51">D</emphasis>
<taxonomicName id="8C5285D4C7B9DFBDD1B146A015F7DB89" class="Gastropoda" family="Camaenidae" genus="Globotrochus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Globotrochus onestera" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="onestera">
<emphasis id="B03CA093F14918AD0CBE2852511EEA07" italics="true" pageId="0" pageNumber="51">Globotrochus onestera</emphasis>
</taxonomicName>
from Cuc Phuong, Vietnam (shell width about 15 mm).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="B09BBA89741B37CDC0783CA4EDB39B98" pageId="0" pageNumber="51" type="materials_examined">
<paragraph id="D2BEDFF1D5E115960FA2E452268037BD" pageId="0" pageNumber="51">Material examined.</paragraph>
<paragraph id="8A8C67E542D800C48FAADC0DCC591CDC" pageId="0" pageNumber="51">
<emphasis id="FAD74A295B7CFAF0EF334B758E22E142" bold="true" pageId="0" pageNumber="51">Type specimens.</emphasis>
Lectotype SMF 27534a (
<figureCitation id="832E5728006CE7BB95093C7AD6C3245D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A, B Ganesella capitium A lectotype UMZC I. 102385 / 1 and B paralectotype UMZC I. 102385 / 2 - 3 C-E Ganesella hariola C lectotype UMZC I. 104370 / 1 D paralectotype UMZC I. 104370 / 2 - 4 and E shell from Popa Mountain, Myanmar CUMZ 5134 F-H Ganesella carinella F lectotype SMF 27534 a G paralectotype SMF 27534 b and H shell from Takhli, Nakhonsawan CUMZ 5133." figureDoi="10.3897/zookeys.870.36970.figure2" httpUri="https://binary.pensoft.net/fig/323907" pageId="0" pageNumber="51">Fig. 2F</figureCitation>
, height 14.8 mm, width 18.5 mm) and paralectotype SMF 27534b (1 shell;
<figureCitation id="B1D14DC09B98C117D1F87076415169D0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A, B Ganesella capitium A lectotype UMZC I. 102385 / 1 and B paralectotype UMZC I. 102385 / 2 - 3 C-E Ganesella hariola C lectotype UMZC I. 104370 / 1 D paralectotype UMZC I. 104370 / 2 - 4 and E shell from Popa Mountain, Myanmar CUMZ 5134 F-H Ganesella carinella F lectotype SMF 27534 a G paralectotype SMF 27534 b and H shell from Takhli, Nakhonsawan CUMZ 5133." figureDoi="10.3897/zookeys.870.36970.figure2" httpUri="https://binary.pensoft.net/fig/323907" pageId="0" pageNumber="51">Fig. 2G</figureCitation>
, height 17.6 mm, width 18.2 mm) from Siam [Thailand].
</paragraph>
<caption id="1286886B95EB762F5AC364ECE8556522" doi="10.3897/zookeys.870.36970.figure2" httpUri="https://binary.pensoft.net/fig/323907" pageId="0" pageNumber="51" start="Figure 2" startId="F2">
<paragraph id="D1C2D66BD4AEE22F897D7D65C3DD2CC5" pageId="0" pageNumber="51">
Figure 2.
<emphasis id="B3D4F4111A02BDEE7CA3381E283126BD" bold="true" pageId="0" pageNumber="51">A, B</emphasis>
<taxonomicName id="68C9DFC54A3C50E514F7D655BE8082BB" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella capitium" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="capitium">
<emphasis id="E4EAB95BE00378CBCB50ECAB1F67E4F9" italics="true" pageId="0" pageNumber="51">Ganesella capitium</emphasis>
</taxonomicName>
<emphasis id="5E819C55A900DC064CD0B0434BA20E6F" bold="true" pageId="0" pageNumber="51">A</emphasis>
lectotype UMZC I.102385/1 and
<emphasis id="EAD774CD894ED1F4E001BD2E83F9F13B" bold="true" pageId="0" pageNumber="51">B</emphasis>
paralectotype UMZC I.102385/2-3
<emphasis id="5056BB0AF637F150BA431D0B90409D03" bold="true" pageId="0" pageNumber="51">
<normalizedToken id="8D92193542186BF3EE728516E37F0AFE" originalValue="C–E">C-E</normalizedToken>
</emphasis>
<taxonomicName id="CB1AE36C9C48AEEC90BFDF560CA20563" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella hariola" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="hariola">
<emphasis id="2A383DEC9B8F5609F76E6513271A2A79" italics="true" pageId="0" pageNumber="51">Ganesella hariola</emphasis>
</taxonomicName>
<emphasis id="C4C00722CA04E16FC6DD98EE3101B1B6" bold="true" pageId="0" pageNumber="51">C</emphasis>
lectotype UMZC I.104370/1
<emphasis id="C8F81C4C05DE510119CECD8537C699C1" bold="true" pageId="0" pageNumber="51">D</emphasis>
paralectotype UMZC I.104370/2-4 and
<emphasis id="BCD2045A8CFC2D373AE6EDF043B90917" bold="true" pageId="0" pageNumber="51">E</emphasis>
shell from Popa Mountain, Myanmar CUMZ 5134
<emphasis id="8AC1ABB2677C0707F696CB4EEC12F6C3" bold="true" pageId="0" pageNumber="51">
<normalizedToken id="04E73CBB318C17E6EB778A8CD604FBFD" originalValue="F–H">F-H</normalizedToken>
</emphasis>
<taxonomicName id="DB47F0B2C4FBBF4E4AB91D755920A9AD" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella carinella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="carinella">
<emphasis id="4B1402A36A185B737EE7AFBF9DD1BE6D" italics="true" pageId="0" pageNumber="51">Ganesella carinella</emphasis>
</taxonomicName>
<emphasis id="80848493AB3AE4DE38E182DA85FD8456" bold="true" pageId="0" pageNumber="51">F</emphasis>
lectotype SMF 27534a
<emphasis id="C138987A09B211C2D4FC020061BED2FE" bold="true" pageId="0" pageNumber="51">G</emphasis>
paralectotype SMF 27534b and
<emphasis id="324EAD8A65D315E065A7520F06BA7052" bold="true" pageId="0" pageNumber="51">H</emphasis>
shell from Takhli, Nakhonsawan CUMZ 5133.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="57B2A476D3329BC0D87738E73AC4D79A" pageId="0" pageNumber="51" type="materials_examined">
<paragraph id="2C8057EE33FE42CEDB364A537A3535E9" pageId="0" pageNumber="51">Additional material.</paragraph>
<paragraph id="621BBCB52F9F51970C8F006842F70DC2" pageId="0" pageNumber="51">
<emphasis id="A407430126A51E826F18C4DCCA79E422" bold="true" pageId="0" pageNumber="51">CAMBODIA</emphasis>
: Cambodia: NHMUK Cuming coll. ex Mouhot (2 shells).
<emphasis id="8EF191762AFC5C871897F6B06A091CC5" bold="true" pageId="0" pageNumber="51">THAILAND</emphasis>
: Siam: NHMUK 1902.9.17.30-31 (2 shells). Tam Barijinda, Chom Thong District, Chiangmai Province: CUMZ 4153, 4193, 4165 (
<figureCitation id="FA4D3085976A9290E7B2C58B2E05C733" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of &quot; harryleei Thach, 2017 &quot; from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">Fig. 3B</figureCitation>
), 5123. Wat Tham Rakung, Sri Samrong District, Sukhothai Province: CUMZ 4937. Tam Lom-Tam Wang, Sri Samrong District, Sukhothai Province: CUMZ 4938. Tam Pha Thaphol, Nern Maprang District, Phitsanuloke Province: CUMZ 4195, 5127. Tam Wang Daeng, Nern Maprang District, Phitsanuloke Province: CUMZ 4932, 4939, 5113, 5126. Wat Chuek Charoentham, Ban Rai District, Uthaithani Province: CUMZ 4935. Wat Sri Uthumporn, Muang District, Nakhonsawan Province: CUMZ 4940. Tam Phet-Tam Thong, Takhi District, Nakhonsawan Province: CUMZ 4173, 4943, 5121, 5133 (
<figureCitation id="E78DB41EB888A69962C66C1785465C56" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. A, B Ganesella capitium A lectotype UMZC I. 102385 / 1 and B paralectotype UMZC I. 102385 / 2 - 3 C-E Ganesella hariola C lectotype UMZC I. 104370 / 1 D paralectotype UMZC I. 104370 / 2 - 4 and E shell from Popa Mountain, Myanmar CUMZ 5134 F-H Ganesella carinella F lectotype SMF 27534 a G paralectotype SMF 27534 b and H shell from Takhli, Nakhonsawan CUMZ 5133." figureDoi="10.3897/zookeys.870.36970.figure2" httpUri="https://binary.pensoft.net/fig/323907" pageId="0" pageNumber="51">Fig. 2H</figureCitation>
). Tam Poon Sawan, Srithep District, Phetchabun Province: CUMZ 4284. Tam Sombat Chomphol, Lomsak District, Phetchabun Province: CUMZ 4934, 5115, 5125. Khao Samokorn, Tha Wung District, Lopburi Province: CUMZ, 4218, 4279, 4282 (
<figureCitation id="F6576B3C1917DDE621317E7EF47D246A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of &quot; harryleei Thach, 2017 &quot; from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">Fig. 3A</figureCitation>
), 4933. Wat Bandai Samsaen, Banmee District, Lopburi Province: CUMZ 4280. Tam Santisuk, Kok Samrong District, Lopburi Province: CUMZ 4931. Tam Tam-bon, Chaibadan District, Lopburi Province: CUMZ 5116. Muak Lek Waterfall, Muak Lek District, Saraburi Province: CUMZ 4186, 4172, 4941. Tam Dao Khaokaeo, Muak Lek District, Saraburi Province: CUMZ 4197. Tam Singha Ratde-cho, Kaeng Khoi District, Saraburi Province: CUMZ 4164, 4178, 5122. Tam Sriwilai, Chaloem Phrakiat District, Saraburi Province: CUMZ 4187, 4930. Wat Thep Pitak, Pakchong District, Nakhon Ratchasrima Province: CUMZ 4199. Wang Takrai Waterfall, Muang District, Nakhon Nayok Province: CUMZ 4942. Wat Khao Chakan, Khao Chakan District, Srakaeo Province: CUMZ 4159, 4182, 4213, 5114. Ta Praya District, Srakaeo Province: CUMZ 4283. Tam Phet Phothong, Klonghad District, Srakaeo Province: CUMZ 5120. Tam Kaeo Sawanbandan, Pong Namron District, Chanthaburi Province: CUMZ 4165.
</paragraph>
<caption id="ACFCC211F5341F6A61DAB17F17DBA284" doi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51" start="Figure 3" startId="F3">
<paragraph id="CE9C48F501EE8E43CDE15B3495E61829" pageId="0" pageNumber="51">
Figure 3.
<emphasis id="D633BE345228EA003955100C74A65901" bold="true" pageId="0" pageNumber="51">A, B</emphasis>
<taxonomicName id="BF330C55843F1DD4E29077C5CFF33279" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella carinella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="carinella">
<emphasis id="5D590799B526F46A6EB46133BB35C52D" italics="true" pageId="0" pageNumber="51">Ganesella carinella</emphasis>
</taxonomicName>
<emphasis id="78859148F0ABCFA46C8E0B50AB7162F7" bold="true" pageId="0" pageNumber="51">A</emphasis>
shell from Lopburi CUMZ 4282 and
<emphasis id="C2D1C4268FBDE5B413EB81E86034F025" bold="true" pageId="0" pageNumber="51">B</emphasis>
shell from Tam Brichinda, Chiangmai CUMZ 4165
<emphasis id="66D16092941E01C8D10DEB3AA782589B" bold="true" pageId="0" pageNumber="51">
<normalizedToken id="3FDE35C9D1A2CCF25A0CEDF6867FFFEA" originalValue="C–I">C-I</normalizedToken>
</emphasis>
<taxonomicName id="4C3C853BE7482F1FC714474BF1892B9A" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella rhombostoma" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="rhombostoma">
<emphasis id="4B5B9966083C6593EB2578F7DE539E80" italics="true" pageId="0" pageNumber="51">Ganesella rhombostoma</emphasis>
</taxonomicName>
<emphasis id="820E42D0F6493C6D42BDA36D4466FC82" bold="true" pageId="0" pageNumber="51">C</emphasis>
lectotype NHMUK 20130215/1
<emphasis id="980BD2265A53F1F323DEDC628799D5F2" bold="true" pageId="0" pageNumber="51">D</emphasis>
paralectotype NHMUK 20130215/2-3
<emphasis id="7705CF82A0AF5251464A9E617B453F04" bold="true" pageId="0" pageNumber="51">E</emphasis>
shell of &quot;
<taxonomicName id="25D3F23B9878B66D9A5FC2BFCF081D4C" genus="Orthurethra" lsidName="Orthurethra harryleei" pageId="0" pageNumber="51" rank="species" species="harryleei">
<emphasis id="EB13738EFF92175C16BA13F373E9CF17" italics="true" pageId="0" pageNumber="51">harryleei</emphasis>
</taxonomicName>
Thach, 2017&quot; from Cambodia FMNH 381985
<emphasis id="49BDC11196FD045A0545249931C3A0A3" bold="true" pageId="0" pageNumber="51">F</emphasis>
shell from Klonghad, Srakeo CUMZ 5132
<emphasis id="4CAEE13C98E9635E586912F209616A0B" bold="true" pageId="0" pageNumber="51">G</emphasis>
shell from Chachoengsao CUMZ 5130
<emphasis id="EDAC5317E9AD1C8503C02E7F3195CAA7" bold="true" pageId="0" pageNumber="51">H</emphasis>
shell from Sirisophon, Cambodia CUMZ 5131 and
<emphasis id="AF2AEDC1E2F64C8DFFDE55325E6377F3" bold="true" pageId="0" pageNumber="51">I</emphasis>
shell from Srakaeo CUMZ 4286.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F04F7058D6CB4535BE4B88D6E459E3D1" pageId="0" pageNumber="51" type="description">
<paragraph id="DD1DAAE28B58B348C8076DE13F77F026" pageId="0" pageNumber="51">Description.</paragraph>
<paragraph id="0D8A7E9BF9A7DFFE04E5C6344DFCC0A6" pageId="0" pageNumber="51">
<emphasis id="60AA25DE32EDB8F6B0740717734D6A57" bold="true" pageId="0" pageNumber="51">Shell.</emphasis>
Shell small, dextral, thin and depressed trochoid. Apex acute; embryonic shell smooth with brownish colour. Whorls 5-6, increasing regularly, slightly convex and slightly shouldered near suture; suture wide and shallow. Shell surface smooth or with fine growth lines. Last whorl large, angular with strong keel; beneath convex. Shell brownish and translucent, with wide, whitish-opaque, irregular spiral band on upper and lower side. Aperture ovate; lip slightly expanded and whitish; parietal callus transparent. Umbilicus rimate with small hole; columella whitish and expanded overhanging umbilicus.
</paragraph>
<paragraph id="6A3F63E65B4F948C1E7B1DC0568FEE86" pageId="0" pageNumber="51">
<emphasis id="7A4634F5DC5FFA53F636B23C17B5252E" bold="true" pageId="0" pageNumber="51">Genital organs.</emphasis>
Atrium (at) short (
<emphasis id="DF6757CF4CA411DD78D8BC91A50EF924" italics="true" pageId="0" pageNumber="51">N</emphasis>
= 10) and proximally with blackish tissues. Penis (p) cylindrical, about as long as the vagina; proximally blackish; distally slightly enlarged at penial verge. Epiphallus (e) and flagellum (fl) very short, approximately one-third of penis length. Vas deferens (vd) long and narrow, extends from free oviduct and terminates at distal epiphallus. Penial retractor muscle (pr) long and thin. Flagellum short, as long as or longer than epiphallus (
<figureCitation id="59421BDC0373415230B11E043AF25FA8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="8184DC8546F8C6D78638771FF3D339A1" pageId="0" pageNumber="51">
Penial verge (pv) small, conical, and with smooth surface. Penial wall ribbed, forming a series of swollen longitudinal pilasters (pp); middle wall with very thin pilasters. Pilasters swollen in the distal portion of penial chamber (
<figureCitation id="DEFD2A51295D293A20A9BDB4A65B04AD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="216158A5580D33465563E3A3B3C7E048" pageId="0" pageNumber="51">
Vagina (v) long, cylindrical and proximally with blackish tissues. Gametolytic duct (gd) short; proximally wider; distally tapering to small tube and terminated with gametolytic sac (gc). Free oviduct (fo) short; oviduct (ov) enlarged with curled lobules. Albumen gland (ag) curved ligulate. Hermaphroditic duct (hd) convoluted and connected to hermaphrodic gland (hg) (
<figureCitation id="37EA130B3C991B29D497243B141F6A67" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="C3280AD2DE95B23DB1632692134129F0" pageId="0" pageNumber="51">
Internal wall of vagina with smooth, longitudinal, vaginal pilasters (vp); with slightly deep crenulated ridges throughout vaginal chamber (
<figureCitation id="180A2C292E00BEF72422EDD40C529BB4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="48D1500A9C21DEDAD5CBFA06BD98F205" pageId="0" pageNumber="51">
<emphasis id="77804AAFC18E4D8BBF94723A5ECAE7F5" bold="true" pageId="0" pageNumber="51">Radula.</emphasis>
Teeth arranged in nearly straight rows, each row containing 74 (37-(4-6)-1-(4-6)-36) teeth. Central tooth monocuspid with spatulate and truncate cusp. Lateral teeth larger than central tooth; teeth no. 1-3 monocuspid, no. 4-6 bicuspid, endocone spatulate and ectocone very small with pointed cusp (
<figureCitation id="F24427C798088C43D5B99735CB5E05A0" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Radula. A-C Ganesella carinella D-F Ganesella rhombostoma and G-I Globotrochus onestera A, D, G central tooth with the first to the second lateral teeth B, E, H lateral teeth with the tricuspid marginal teeth transition C, F, I outermost marginal teeth. Numbers indicate order of lateral and marginal teeth. Central tooth indicated by ' C'." figureDoi="10.3897/zookeys.870.36970.figure7" httpUri="https://binary.pensoft.net/fig/323912" pageId="0" pageNumber="51">Fig. 7A, B</figureCitation>
). Marginal teeth start from teeth no. 6-7. Inner marginal teeth tricuspid, endocone and ectocone very small with pointed cusp and mesocone large and spatulate. Outermost teeth (near radula edge) tricuspid, endocone and ectocone with two or more pointed cusps, and mesocone large with curved cusp (
<figureCitation id="49F8101819C01155C0C35A7A4F667143" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Radula. A-C Ganesella carinella D-F Ganesella rhombostoma and G-I Globotrochus onestera A, D, G central tooth with the first to the second lateral teeth B, E, H lateral teeth with the tricuspid marginal teeth transition C, F, I outermost marginal teeth. Numbers indicate order of lateral and marginal teeth. Central tooth indicated by ' C'." figureDoi="10.3897/zookeys.870.36970.figure7" httpUri="https://binary.pensoft.net/fig/323912" pageId="0" pageNumber="51">Fig. 7B, C</figureCitation>
).
</paragraph>
<paragraph id="6A6A90F3DA9DCB1A8A88791583AE40A0" pageId="0" pageNumber="51">
Jaw crescent, with anterior convex cutting margin. Vertical ribs prominent, and variable in number and size (
<figureCitation id="D7EDB641A740F464246F671BBE577A4D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">Fig. 5D</figureCitation>
).
</paragraph>
<paragraph id="6F9A12653EC5519EFF507682A1355E09" pageId="0" pageNumber="51">
<emphasis id="FDA7BB712016DE7100F0A513EEEA14DE" bold="true" pageId="0" pageNumber="51">Pallial system.</emphasis>
Typical sigmurethran; heart (au and ve) located left of kidney (on right in
<figureCitation id="FF8BCCF5148B103C80083DE5B9B4F32B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">Fig. 5C</figureCitation>
). Pulmonary cavity approximately 5
<normalizedToken id="C1F58B6E079BE70A4D80FD4134ADE797" originalValue="×">x</normalizedToken>
longer than wide. Pulmonary vein (puv) and venation on lung roof (l) distinct and well developed. Kidney (k) long, slender and extending from posterior side to approximately the middle of pulmonary cavity. Ureter (ur) is a sigmoid, closed tube arising from apex of kidney, extending along right side of kidney, recurving near rectum (r). Rectal opening adjacent to anus (a) and pneumostome (pn) (
<figureCitation id="517527DA5FE44D5079E839F47510E7D7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">Fig. 5C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="27338C1927502C3E534E5FA6307C661C" pageId="0" pageNumber="51" type="distribution">
<paragraph id="34518ABFCC079B98A7D313840B672B8B" pageId="0" pageNumber="51">Distribution.</paragraph>
<paragraph id="8F2900D6A83F48601E46989084D85314" pageId="0" pageNumber="51">
<taxonomicName id="66F53E7A7D14960520A8F08F46026CFD" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella carinella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="carinella">
<emphasis id="6A45E998A9F6DA2D9071B55B639B656F" italics="true" pageId="0" pageNumber="51">Ganesella carinella</emphasis>
</taxonomicName>
is widely distributed in Thailand: northern area in Chiangmai, Phitsanuloke; northeastern area in Loei, Phetchaboon, Nakhonratchasrima; central area in Saraburi, Lopburi; eastern area in Srakeow, Chanthaburi; western area in Kanchanaburi.
</paragraph>
</subSubSection>
<subSubSection id="8A5F31D184FB8F32C881433166D37D8E" pageId="0" pageNumber="51" type="remarks">
<paragraph id="3B3F5FC39F3879BCD7152172F3254532" pageId="0" pageNumber="51">Remarks.</paragraph>
<paragraph id="A44EE7B6C8412E5196D82528F0E8F295" pageId="0" pageNumber="51">
Hitherto,
<taxonomicName id="15673C54B0FD225773FF8C5D74B9B323" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella carinella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="carinella">
<emphasis id="C6D25C952D0ECE50ED19FE541B97B63F" italics="true" pageId="0" pageNumber="51">Ganesella carinella</emphasis>
</taxonomicName>
was regarded as a junior synonym of
<taxonomicName id="8C90A788E2298D7A47A720E33959DE9E" lsidName="G. capitium" pageId="0" pageNumber="51" rank="species" species="capitium">
<emphasis id="979C740CD695F42FC7990CFCD6074987" italics="true" pageId="0" pageNumber="51">G. capitium</emphasis>
</taxonomicName>
. However, after
<bibRefCitation id="0C8C85B29F983EB45D3E471FE2A4F634" DOI="https://doi.org/10.3897/zookeys.175.2755" author="Budha, PB" journalOrPublisher="ZooKeys" pageId="0" pageNumber="51" pagination="19 - 26" refId="B12" refString="Budha, PB, Mordan, PB, Naggs, F, Backeljau, T, 2012. Darwininitium - a new fully pseudosigmurethrous orthurethran genus from Nepal (Gastropoda: Pulmonata: Cerastidae). . ZooKeys 175: 19 - 26" title="Darwininitium - a new fully pseudosigmurethrous orthurethran genus from Nepal (Gastropoda: Pulmonata: Cerastidae)." url="https://doi.org/10.3897/zookeys.175.2755" volume="175" year="2012">Budha et al. (2012)</bibRefCitation>
described the genital apparatus of conchologically typical
<taxonomicName id="34B62570C8BB1060FE24AE365F0D78C8" lsidName="G. capitium" pageId="0" pageNumber="51" rank="species" species="capitium">
<emphasis id="B5CEB33A0FD864A7CEFF15E8BC903551" italics="true" pageId="0" pageNumber="51">G. capitium</emphasis>
</taxonomicName>
, it became clear that
<taxonomicName id="7C4EC0085D7F2F8AD1793A8618A3B934" lsidName="G. carinella" pageId="0" pageNumber="51" rank="species" species="carinella">
<emphasis id="203276ED417E51BA9C4A0F23D39F9D35" italics="true" pageId="0" pageNumber="51">G. carinella</emphasis>
</taxonomicName>
differs from the type species not only by having a more depressed shell with a strong peripheral keel but also by its longer penis and epiphallus, and its shorter vagina. In contrast,
<taxonomicName id="E466E0AD480FA6E94FC189C94ECA3F58" lsidName="G. capitium" pageId="0" pageNumber="51" rank="species" species="capitium">
<emphasis id="DD42A92B188D0901FD88E94F1FAB09AC" italics="true" pageId="0" pageNumber="51">G. capitium</emphasis>
</taxonomicName>
has a more elevated trochoid shell, an angular last whorl, a shorter penis and epiphallus, and a longer vagina.
</paragraph>
<paragraph id="EF5377DA976636DBB66E859E194F613F" pageId="0" pageNumber="51">
<taxonomicName id="879C28B88E020CEAC00E6B7C1F2AC6BF" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella carinella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="carinella">
<emphasis id="E96C13966E305F625834D9A5B93DD3AF" italics="true" pageId="0" pageNumber="51">Ganesella carinella</emphasis>
</taxonomicName>
shows considerable variation in shell shape and colour. For example, specimens from Lopburi (
<figureCitation id="186AD8AD80A01826A1FE8FB1A43E304D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of &quot; harryleei Thach, 2017 &quot; from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">Fig. 3A</figureCitation>
) tend to be paler and have a more descending aperture than specimens from Saraburi. Specimens from Chiangmai (
<figureCitation id="2D82232EBF5D962F1CE14AD99637C39B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of &quot; harryleei Thach, 2017 &quot; from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">Fig. 3B</figureCitation>
) tend to have more whitish spots on their shell than specimens from Lopburi and Saraburi. Yet, this conchological variation is not matched by consistent genital differences. Therefore, we conclude that the shell variation within this widely-distributed species only involves intraspecific polymorphism.
</paragraph>
</subSubSection>
</treatment>
</document>
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