246 lines
23 KiB
XML
246 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.589.7933" ID-GBIF-Dataset="6d2188dd-976a-47a7-b857-5a14d5eb2081" ID-PMC="PMC4926661" ID-Pensoft-Pub="1313-2970-589-23" ID-PubMed="27408533" ID-ZBK="EECEC70CA98343C486A208CD536293EB" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-589-23" ModsDocOrigin="ZooKeys 589" ModsDocTitle="The first revision of the carnivorous land snail family Streptaxidae in Laos, with description of three new species (Pulmonata, Stylommatophora, Streptaxidae)" checkinTime="1463539066408" checkinUser="pensoft" docAuthor="Inkhavilay, Khamla, Siriboon, Thanit, Sutcharit, Chirasak, Rowson, Ben & Panha, Somsak" docDate="2016" docId="E4E68B4AED1C33387648A45A44CB31C0" docLanguage="en" docName="ZooKeys 589: 23-53" docOrigin="ZooKeys 589" docSource="http://dx.doi.org/10.3897/zookeys.589.7933" docTitle="Perrottetia unidentata Inkhavilay & Panha, sp. n." docType="treatment" docUuid="B47C107D-B7A5-4D70-8640-47F10AE13AC7" docUuidSource="ZooBank" docVersion="5" lastPageNumber="37" masterDocId="FFF6FFEEFFBEFFA73169C413B161FFDC" masterDocTitle="The first revision of the carnivorous land snail family Streptaxidae in Laos, with description of three new species (Pulmonata, Stylommatophora, Streptaxidae)" masterLastPageNumber="53" masterPageNumber="23" pageNumber="36" updateTime="1668163105615" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The first revision of the carnivorous land snail family Streptaxidae in Laos, with description of three new species (Pulmonata, Stylommatophora, Streptaxidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Inkhavilay, Khamla</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Siriboon, Thanit</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rowson, Ben</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="volume">
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<mods:number>589</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>23</mods:start>
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<mods:end>53</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.589.7933</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.589.7933</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-589-23</mods:identifier>
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<mods:identifier type="ZBK">EECEC70CA98343C486A208CD536293EB</mods:identifier>
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<mods:identifier type="ZooBank">EECEC70CA98343C486A208CD536293EB</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127892683" LSID="urn:lsid:zoobank.org:act:B47C107D-B7A5-4D70-8640-47F10AE13AC7" httpUri="http://treatment.plazi.org/id/E4E68B4AED1C33387648A45A44CB31C0" lastPageId="14" lastPageNumber="37" pageId="13" pageNumber="36">
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<subSubSection pageId="13" pageNumber="36" type="multiple">
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<paragraph pageId="13" pageNumber="36">Taxon classification Animalia Pulmonata Streptaxidae</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="36" type="nomenclature">
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<paragraph pageId="13" pageNumber="36">
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<taxonomicName LSID="http://zoobank.org/B47C107D-B7A5-4D70-8640-47F10AE13AC7" authority="Inkhavilay & Panha" class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia unidentata" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="unidentata">Perrottetia unidentata Inkhavilay & Panha</taxonomicName>
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<taxonomicNameLabel pageId="13" pageNumber="36">sp. n.</taxonomicNameLabel>
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Figs 1, 5
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<normalizedToken originalValue="F–I">F-I</normalizedToken>
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, 7E, F, 10
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<normalizedToken originalValue="A–F">A-F</normalizedToken>
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, I; Table 1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="36" type="type material">
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<paragraph pageId="13" pageNumber="36">Type material.</paragraph>
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<paragraph pageId="13" pageNumber="36">
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Holotype CUMZ 6281 (Fig. 5F). Measurement: shell height 5.3 mm, shell width 9.7 mm and
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<normalizedToken originalValue="6½">61/2</normalizedToken>
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whorls. Paratypes CUMZ 6282 (4 shells; Fig. 5G), CUMZ 6283 (1 specimen in ethanol; Figs 7E, F, 9
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<normalizedToken originalValue="A–F">A-F</normalizedToken>
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, I), NHMUK 20160250 (2 shells).
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</paragraph>
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<caption pageId="13" pageNumber="36">
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<paragraph pageId="13" pageNumber="36">
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Figure 6. Shells of
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="genus">Perrottetia</taxonomicName>
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and
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Indoartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Indoartemon" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="genus">Indoartemon</taxonomicName>
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spp. A, B
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia megadentata" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="megadentata">Perrottetia megadentata</taxonomicName>
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sp. n. A holotype CUMZ 6286 with apertural dentition, and B paratype CUMZ 6287. C
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Indoartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Indoartemon tridens" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="tridens">Indoartemon tridens</taxonomicName>
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holotype SMF 108507/1 with apertural dentition
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Indoartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Indoartemon diodonta" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="diodonta">Indoartemon diodonta</taxonomicName>
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sp. n. D holotype CUMZ 6289 with apertural dentition E paratypes CUMZ 6290, and F specimen from Tam Nang Ann, Tha Khek, Khammouan CUMZ 2691.
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</paragraph>
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</caption>
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<caption pageId="13" pageNumber="36">
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<paragraph pageId="13" pageNumber="36">
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Figure 7. Genitalia of
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="genus">Haploptychius</taxonomicName>
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and
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="genus">Perrottetia</taxonomicName>
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species. A, B
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius pellucens" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="pellucens">Haploptychius pellucens</taxonomicName>
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CUMZ 2670 A reproductive system, and B insertion of vas deferens into penial sheath C, D
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius porrectus" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="porrectus">Haploptychius porrectus</taxonomicName>
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CUMZ 6274 C reproductive system, and D insertion of vas deferens into penial sheath E, F
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia unidentata" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="unidentata">Perrottetia unidentata</taxonomicName>
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sp. n. CUMZ 6283 E reproductive system, and F insertion of vas deferens into penial sheath.
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</paragraph>
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</caption>
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<caption pageId="13" pageNumber="36">
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<paragraph pageId="13" pageNumber="36">
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Figure 8. Genitalia of
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius pellucens" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="pellucens">Haploptychius pellucens</taxonomicName>
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A completed reproductive system CUMZ 2670 B, C completed reproductive system with "vaginal caecum" CUMZ 6265, and D aphallic reproductive system CUMZ 6265.
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</paragraph>
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</caption>
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<caption pageId="13" pageNumber="36">
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<paragraph pageId="13" pageNumber="36">
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Figure 9. Internal sculpture of genitalia of
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="genus">Haploptychius</taxonomicName>
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spp.
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<normalizedToken originalValue="A–F">A-F</normalizedToken>
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius pellucens" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="pellucens">Haploptychius pellucens</taxonomicName>
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, CUMZ 2670 A details of atrium surface B arrangement of penial hooks C top view of penial hook (from white square in B) D arrangement of penial hooks E lateral view of penial hook, and F arrangement of undulated parallel vaginal folds.
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<normalizedToken originalValue="G–M">G-M</normalizedToken>
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius porrectus" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="porrectus">Haploptychius porrectus</taxonomicName>
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specimen CUMZ 6275 G details of atrium surface H arrangement of penial hooks in distal area I lateral view of penial hook in distal area J arrangement of penial hooks in proximal area K lateral view of penial hook in proximal area L arrangement of papillae and vaginal folds, and M arrangement of vaginal folds.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="36" type="materials_examined">
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<paragraph pageId="13" pageNumber="36">Other material examined.</paragraph>
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<paragraph pageId="13" pageNumber="36">Tam Than Kaisone, Viengxay, Houaphanh, Laos: CUMZ 6284 (5 shell; Fig. 5I), CUMZ 6285 (2 shells; Fig. 5H).</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="36" type="type locality">
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<paragraph pageId="13" pageNumber="36">Type locality.</paragraph>
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<paragraph pageId="13" pageNumber="36">
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The limestone outcrop at Ban Nawit, Viengxay, Houaphanh, Laos (
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="20.377028">20°22'37.3"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="104.27867">104°16'43.2"E</geoCoordinate>
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) about 700 meters above mean sea level.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="36" type="diagnosis">
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<paragraph pageId="13" pageNumber="36">Diagnosis.</paragraph>
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<paragraph pageId="13" pageNumber="36">
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This new species differs from
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia daedaleus" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="daedaleus">Perrottetia daedaleus</taxonomicName>
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,
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<taxonomicName class="Magnoliopsida" family="Celastraceae" genus="Perrotettia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Perrotettia aquilonaria" order="Celastrales" pageId="13" pageNumber="36" phylum="Tracheophyta" rank="species" species="aquilonaria">Perrotettia aquilonaria</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia dugasti" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="dugasti">Perrottetia dugasti</taxonomicName>
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and
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia messageri" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="messageri">Perrottetia messageri</taxonomicName>
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from Vietnam in having an oblique shell, a single parietal lamella, widely expanded lip, the last whorl strongly axially deflected, the left side of penultimate whorl well extended beyond the diameter of last whorl, and the distal end of penis with a wing-like structure. The other four species have two parietal lamellae, the last whorl little axially deflected and the left side of penultimate whorl not extended beyond the diameter of the last whorl. For further comparison,
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia daedaleus" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="daedaleus">Perrottetia daedaleus</taxonomicName>
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has an elevated spire, transverse ridges over the entire shell and a basal lamella located deep inside aperture (Fig. 5C);
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<taxonomicName class="Magnoliopsida" family="Celastraceae" genus="Perrotettia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Perrotettia aquilonaria" order="Celastrales" pageId="13" pageNumber="36" phylum="Tracheophyta" rank="species" species="aquilonaria">Perrotettia aquilonaria</taxonomicName>
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has a smaller shell, elevated spire, bifid columellar lamella, and genitalia with atrial pores and vaginal hooks absent (Fig. 5E);
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia dugasti" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="dugasti">Perrottetia dugasti</taxonomicName>
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and
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia messageri" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="messageri">Perrottetia messageri</taxonomicName>
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have a smooth shell surface, a bifid collumella lamella and a supracolumellar lamella (Fig. 5A, D).
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia gudei" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="gudei">Perrottetia gudei</taxonomicName>
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from north Vietnam differs from the new species in having an elevated spire, in being less deviated from the vertical axis, and in having thin transverse ridges (see
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<bibRefCitation author="Siriboon, T" journalOrPublisher="ZooKeys" pageId="24" pageNumber="47" pagination="41 - 57" title="Three new species of the carnivorous snail genus Perrottetia Kobelt, 1905 from Thailand (Pulmonata, Streptaxidae)." url="10.3897/zookeys.287.4572" volume="287" year="2013">Siriboon et al. 2013</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="14" lastPageNumber="37" pageId="13" pageNumber="36" type="description">
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<paragraph pageId="13" pageNumber="36">Description.</paragraph>
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<paragraph pageId="13" pageNumber="36">
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Shell. Shell oblique-heliciform, semi-transparent; whorls
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<normalizedToken originalValue="6½">61/2</normalizedToken>
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, spire weakly convex with distinct suture. Shell surface glossy with strong transverse ridges on upper shell surface. Embryonic shell large, about
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<normalizedToken originalValue="2½">21/2</normalizedToken>
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whorls, with a smooth surface; following whorls regularly coiled. Shell periphery shouldered; last whorl axially deflected; two deep longitudinal furrows present. Aperture semi-ovate; peristome discontinuous; parietal callus thin; lip thickened, broadly expanded and slightly reflected. Apertural dentition with one large, strong and sinuous parietal lamella, one small upper palatal lamella, one palatal lamella, one large basal lamella, one strong columellar lamella, and one small supracolumellar lamella. Umbilicus widely open and shallow (Fig. 5
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<normalizedToken originalValue="F–I">F-I</normalizedToken>
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).
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</paragraph>
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<paragraph pageId="13" pageNumber="36">Radula. Each row consists of 26-38 teeth with formula (13-19)-1-(13-19). Central tooth small and triangular, with pointed cusp. Lateral and marginal teeth undifferentiated, lanceolate, unicuspid. Latero-marginal teeth gradually reduce in size, with outermost teeth much smaller and shorter than inner teeth (Fig. 10I).</paragraph>
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<caption pageId="13" pageNumber="36">
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<paragraph pageId="13" pageNumber="36">
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Figure 10. Internal sculpture of genitalia of
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<normalizedToken originalValue="A–F">A-F</normalizedToken>
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia unidentata" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="unidentata">Perrottetia unidentata</taxonomicName>
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sp. n. paratype CUMZ 6283 A details of atrium surface B arrangement of penial hooks C top view of penial hook D lateral view of penial hooks E arrangement of vaginal hooks, and F top view of vaginal hook. Radula morphology of G
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius pellucens" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="pellucens">Haploptychius pellucens</taxonomicName>
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specimen CUMZ 2670 H
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haploptychius porrectus" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="porrectus">Haploptychius porrectus</taxonomicName>
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specimen CUMZ 6275, and I
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<taxonomicName class="Gastropoda" family="Streptaxidae" genus="Perrottetia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Perrottetia unidentata" order="Stylommatophora" pageId="13" pageNumber="36" phylum="Mollusca" rank="species" species="unidentata">Perrottetia unidentata</taxonomicName>
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sp. n. paratype CUMZ 6283.
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</paragraph>
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</caption>
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<paragraph pageId="13" pageNumber="36">Genital organs. Atrium (at) short. Proximal penis (p) long and slender; distal part near retractor muscle with an expanded wing-like structure (a flat blade on either side of the penis, each about one-tenth of penis length). Penial sheath (ps) thin and extending about one-third of penis length; penial sheath retractor muscle (psr) very thin, originating at atrium and inserting distally on penial sheath (Fig. 7E). Vas deferens (vd) passes through about one-third of penial sheath length before entering into penis apically (Fig. 7F). Penial retractor muscle (pr) thin and long, inserted at penis and vas deferens junction.</paragraph>
|
||
<paragraph pageId="13" pageNumber="36">
|
||
Internal wall of atrium generally smooth (Fig. 10A). Penial wall densely covered with light brown penial hooks, about 20 hooks/200
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
2; hooks located on low elliptical penial papillae. Penial hooks small (<0.1 mm in length), slender, expanded at base, tips pointed and curved towards genital orifice (Fig. 10
|
||
<normalizedToken originalValue="B–D">B-D</normalizedToken>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="14" pageNumber="37">
|
||
<pageBreakToken pageId="14" pageNumber="37" start="start">Vagina</pageBreakToken>
|
||
(v) short, about one-tenth of penis length. Gametolytic duct (gd) a long tube extending as far as albumin gland; gametolytic sac (gs) ovate. Free oviduct (fo) long and cylindrical with equivalent diameter to vagina, tapering distally. Oviduct (ov) enlarged and folded; prostate gland inconspicuous and bound to oviduct. Talon (ta) very small, short and club shape. Hermaphroditic duct (hd) bearing very short and thin seminal vesicle (sv) about one and half times longer than the length from talon to branching point of seminal vesicle (Fig. 7E).
|
||
</paragraph>
|
||
<paragraph pageId="14" pageNumber="37">
|
||
Vaginal wall with transparent vaginal hooks (about 10 hooks/200
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
2). Hooks located on low conical vaginal papillae. Vaginal hooks small (<0.1 mm in length), short and expanded at base; tips pointed and straight to slightly curving away from genital orifice (Fig. 10E, F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="37" type="etymology">
|
||
<paragraph pageId="14" pageNumber="37">Etymology.</paragraph>
|
||
<paragraph pageId="14" pageNumber="37">
|
||
The specific epithet
|
||
<normalizedToken originalValue="“unidentata”">"unidentata"</normalizedToken>
|
||
derived from the Latin words
|
||
<normalizedToken originalValue="“unus”">"unus"</normalizedToken>
|
||
meaning
|
||
<normalizedToken originalValue="“one”">"one"</normalizedToken>
|
||
and
|
||
<normalizedToken originalValue="“dens”">"dens"</normalizedToken>
|
||
meaning
|
||
<normalizedToken originalValue="“tooth”">"tooth"</normalizedToken>
|
||
. It referred to a single parietal lamella (or teeth) of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="37" type="distribution">
|
||
<paragraph pageId="14" pageNumber="37">Distribution.</paragraph>
|
||
<paragraph pageId="14" pageNumber="37">This species is known only from the type locality, Houaphanh, a limestone karst area.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="14" pageNumber="37" type="remarks">
|
||
<paragraph pageId="14" pageNumber="37">Remarks.</paragraph>
|
||
<paragraph pageId="14" pageNumber="37">Shell variation is evident from specimens from Tam Than Kaisone, about 20 km west of the type locality (Fig. 5H, I; CUMZ 6284, 6285). They are smaller, with a sinuous parietal lamella, and sometimes lack the upper palatal lamella (Table 1). However, only five shells and no living specimens were collected, so we provisionally identifying them as the same species.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |