239 lines
30 KiB
XML
239 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.925.39691" ID-GBIF-Dataset="bd72aac6-9c05-4ca2-bfc8-dbf5dc462aa4" ID-PMC="PMC7160194" ID-Pensoft-Pub="1313-2970-925-1" ID-Pensoft-UUID="DCC1953545B9557EA73102AE6D25711F" ID-PubMed="32317852" ID-ZooBank="BB966609087849A1B13C138C2495E6B7" ModsDocID="1313-2970-925-1" checkinTime="1586365656080" checkinUser="pensoft" docAuthor="Maddison, Wayne P., Maddison, David R., Derkarabetian, Shahan & Hedin, Marshal" docDate="2020" docId="E4B7CD0426B9534B9A21FC95631A44C0" docLanguage="en" docName="ZooKeys 925: 1-54" docOrigin="ZooKeys 925" docSource="http://dx.doi.org/10.3897/zookeys.925.39691" docTitle="Jollas cupreus W. Maddison 2020, sp. nov." docType="treatment" docUuid="68F87DD6-8C31-4D0B-A349-245B9B201CF3" docUuidSource="ZooBank" docVersion="5" id="DCC1953545B9557EA73102AE6D25711F" lastPageNumber="1" masterDocId="DCC1953545B9557EA73102AE6D25711F" masterDocTitle="Sitticine jumping spiders: phylogeny, classification, and chromosomes (Araneae, Salticidae, Sitticini)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="1" updateTime="1732479024238" updateUser="ExternalLinkService">
|
||
<mods:mods id="CC6A9ECDF4F3CC6C700D9DB2ACF462AC" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="6732241563FE83EE99B2E9583711B1D9">
|
||
<mods:title id="3E5C56C83E4C2D379E09222567061CE4">Sitticine jumping spiders: phylogeny, classification, and chromosomes (Araneae, Salticidae, Sitticini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="91668699A1802D208C5C06CD944C630D" type="personal">
|
||
<mods:role id="503577096CFEE98822902A5AA5D7B145">
|
||
<mods:roleTerm id="DC428D44FD90FAE83B86BA161703F867">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9988DAB39503284AB0279FFD40C97ADE">Maddison, Wayne P.</mods:namePart>
|
||
<mods:affiliation id="420013AF39E7A404B8FE5DC2137DB5CD">Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada</mods:affiliation>
|
||
<mods:nameIdentifier id="B6C545087DCA9502EA92B3B1F39C731A" type="orcid">https://orcid.org/0000-0003-4953-4575</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="9A2A063E42A1F5983AFD85E97A5D62F0" type="email">wmaddisn@mail.ubc.ca</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="5658F9A04A2D360E1B6979D871141496" type="personal">
|
||
<mods:role id="5444A6B7148DDE63946074E07CAE87AD">
|
||
<mods:roleTerm id="0F111EA05557A00780367C8894B1D844">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="602D655010DCE57E72E577B38F9BB93A">Maddison, David R.</mods:namePart>
|
||
<mods:affiliation id="D8C64DDB457FF32B9B1CF7B3C49589EA">Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA</mods:affiliation>
|
||
<mods:nameIdentifier id="3D4E532FDF3519B7EC747F3E00BDD528" type="orcid">https://orcid.org/0000-0002-7152-3824</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="4048F9F095974FDC2EF6B488D48DF0EC" type="personal">
|
||
<mods:role id="6986735A3F2FC67A24648B128C1C4745">
|
||
<mods:roleTerm id="AF8D8C5316D2AF3DD002B902FB3F8A73">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A8DA6F1F6A327B5111C3BAF6A4CBB73E">Derkarabetian, Shahan</mods:namePart>
|
||
<mods:affiliation id="632761502B134CD3321240B4AA7651E1">Department of Biology, San Diego State University, San Diego, CA 92182, USA & Department of Organismic and Evolutionary Biology, Harvard University, Cambridge MA 02138, USA</mods:affiliation>
|
||
<mods:nameIdentifier id="DEF572F0E6B132FCAFAC1C2F6125BB2A" type="orcid">https://orcid.org/0000-0002-9163-9277</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="E5FE0700B9450E4B9D39B3BC5B97D49F" type="personal">
|
||
<mods:role id="735638B6EDD3A7142097876E3BBD6BD5">
|
||
<mods:roleTerm id="2C4964341C8C9646320690F7A13D0E37">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7958848CC47D1D488DB0248019E0F7F2">Hedin, Marshal</mods:namePart>
|
||
<mods:affiliation id="A6D2AC4F8F62EF42E63680610EEFC9CC">Department of Biology, San Diego State University, San Diego, CA 92182, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="14F198FFA8BD3B2FCB3568A348C49310">text</mods:typeOfResource>
|
||
<mods:relatedItem id="E778451BD39AFB9262FA02ADC80585DB" type="host">
|
||
<mods:titleInfo id="D53867B2349CADA90AD4A49A625D5D70">
|
||
<mods:title id="AFFA038D183C176F2DB2730BC6A9175E">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="FDB283835752B0D9250C77724AD421A9">
|
||
<mods:date id="8400647A5E539A14C435502D2C018933">2020</mods:date>
|
||
<mods:detail id="F75D191B45E1C5AE429BBAF6B79C051D" type="volume">
|
||
<mods:number id="1772A64DDD9222F8959C01722A7AF4E1">925</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="628672F804102207EEE5BEE2F132884C" unit="page">
|
||
<mods:start id="3203E8A129EAE01A008D8D525B3951B1">1</mods:start>
|
||
<mods:end id="825FCDE466CDA863DC62E7BCC08373EF">54</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="E113B6D91AF43A59C00EF5F68FAD7719">
|
||
<mods:url id="FB5682E8B0F4742AE417066FB8ED8FF1">http://dx.doi.org/10.3897/zookeys.925.39691</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="10C6A2A8B65294E96FDB3107704BCF76">journal article</mods:classification>
|
||
<mods:identifier id="1D39CBB02B74478B0909FD8E4FEB4135" type="DOI">http://dx.doi.org/10.3897/zookeys.925.39691</mods:identifier>
|
||
<mods:identifier id="E566D3939DE10CDA113DEFAF79657824" type="Pensoft-Pub">1313-2970-925-1</mods:identifier>
|
||
<mods:identifier id="3215A65F59DE67E788BA895473656B35" type="ZooBank">BB966609087849A1B13C138C2495E6B7</mods:identifier>
|
||
<mods:identifier id="6E08B05AC2DE65ADB96CABD0272DCBE4" type="Pensoft-UUID">DCC1953545B9557EA73102AE6D25711F</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="E4B7CD0426B9534B9A21FC95631A44C0" ID-GBIF-Taxon="163540029" LSID="urn:lsid:zoobank.org:act:68F87DD6-8C31-4D0B-A349-245B9B201CF3" httpUri="http://treatment.plazi.org/id/E4B7CD0426B9534B9A21FC95631A44C0" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Salticidae" scope_order="Araneae" scope_subTribe="Sitticina" scope_tribe="Sitticini">
|
||
<subSubSection id="33C74CC0A57994B24513A251A8F82237" pageId="0" pageNumber="1" type="nomenclature">
|
||
<paragraph id="52A38E8A0D9299A88F303041D93896B7" pageId="0" pageNumber="1">
|
||
<taxonomicName id="AB34946017FFDB108BC6527E992151D6" ID-CoL="943WJ" LSID="E4B7CD04-26B9-534B-9A21-FC95631A44C0" authority="W. Maddison" authorityName="W. Maddison" authorityYear="2020" class="Arachnida" family="Salticidae" genus="Jollas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Jollas cupreus" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cupreus" status="sp. nov.">Jollas cupreus W. Maddison</taxonomicName>
|
||
<taxonomicNameLabel id="4AFD7BF423794FB88E7945AB86896C9F" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
|
||
<figureCitation id="70D4538B2AB5565F5D83E67D60FD14F1" captionStart="Figures 1–14" captionStartId="F1" captionText="Figures 1 - 14. Subtribe Aillutticina (1 - 4) and the Jollas - Tomis clade of the subtribe Sitticina (5 - 14) 1 - 4 Aillutticus nitens, Uruguay (- 34.877, - 56.023): 1 - 3 male 4 female 5, 6 Tomis palpalis male and female, Ecuador (- 0.1996, - 77.7023) 7, 8 Jollas species: 7 J. cupreus male, Ecuador (- 0.675, - 76.397) 8 Jollas sp. female, Ecuador (- 0.7223, - 77.6408) 9 J. leucoproctus, Uruguay (- 34.94, - 54.95) 10 J. flabellatus, Uruguay (- 34.426, - 55.195) 11 - 14 Attinella dorsata male (11 - 13) and female (14), Canada (48.870, - 123.379). Also included in the Jollas - Tomis clade is Sittisax (Figs 99 - 103). Additional members of the Jollas - Tomis clade can be seen in Figs 108 - 128." figureDoi="10.3897/zookeys.925.39691.figures1-14" httpUri="https://binary.pensoft.net/fig/397983" pageId="0" pageNumber="1">Figures 7</figureCitation>
|
||
<figureCitation id="AFF500FB5863CACB2D8154CD8347CCB5" captionStart="Figures 108–119" captionStartId="F11" captionText="Figures 108 - 119. Jollas cupreus, sp. nov. (except 112, J. puntalara) 108, 109 Left palp of holotype 108 ventral view 109 retrolateral view 110 ventral view of epigyne of paratype 111 dorsal view of same, cleared 112 palp of holotype of J. puntalara Galiano 113 - 115 holotype male 116 male from Yasuni, Ecuador (- 0.675, - 76.397) 117 holotype male in alcohol 118, 119 paratype female." figureDoi="10.3897/zookeys.925.39691.figures108-119" httpUri="https://binary.pensoft.net/fig/397993" pageId="0" pageNumber="1">, 108-111</figureCitation>
|
||
<figureCitation id="146594F056E987D62F9C93A5ABD15314" captionStart="Figures 108–119" captionStartId="F11" captionText="Figures 108 - 119. Jollas cupreus, sp. nov. (except 112, J. puntalara) 108, 109 Left palp of holotype 108 ventral view 109 retrolateral view 110 ventral view of epigyne of paratype 111 dorsal view of same, cleared 112 palp of holotype of J. puntalara Galiano 113 - 115 holotype male 116 male from Yasuni, Ecuador (- 0.675, - 76.397) 117 holotype male in alcohol 118, 119 paratype female." figureDoi="10.3897/zookeys.925.39691.figures108-119" httpUri="https://binary.pensoft.net/fig/397993" pageId="0" pageNumber="1">, 113-119</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="318DA11511D337F247DA300D13DD89AC" pageId="0" pageNumber="1" type="type material">
|
||
<paragraph id="009D2EC3A0AA6CD611F2095F6C5FF74A" pageId="0" pageNumber="1">Type material.</paragraph>
|
||
<paragraph id="BF6F6DA21E1C5CFA9B1D9275DA1A53C6" pageId="0" pageNumber="1">
|
||
Male holotype and 2 male, 3 female paratypes from Ecuador: Orellana:
|
||
<normalizedToken id="78AC303EA9A95C0A9381953A8100BD64" originalValue="Río">Rio</normalizedToken>
|
||
Bigal Reserve, main camp area.
|
||
<geoCoordinate id="BF64FBA8163D09A302B94317C1E4746B" degrees="0.5251" direction="north" orientation="latitude" precision="5" value="0.5251">0.5251</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1680549F0EF357EE7B767F60AFB4675D" degrees="77.4177" direction="west" orientation="longitude" precision="5" value="-77.4177">-77.4177</geoCoordinate>
|
||
. 950 m elev. 1-5 November 2010. W & D Maddison, M Vega, M Reyes. WPM#10-041c. The holotype (specimen ECU2010-2060) pertains to the Museum of Zoology, Pontificia Universidad Católica, Quito, Ecuador (QCAZ), but is currently held in the Spencer Entomological Collection at the Beaty Biodiversity Museum, University of British Columbia (UBC-SEM).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="486BC4CF8533BDCB92985784586D980A" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="1D38EE702353A1F4A67CF9D188DE4D67" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="0CBE18B632DC816429B112F8B05C6AE7" pageId="0" pageNumber="1">Refers to the copper colour of males.</paragraph>
|
||
<paragraph id="9EB969CE81AA627F97F3910EFB208E16" pageId="0" pageNumber="1">
|
||
A species common in eastern Ecuador on disturbed open grassy ground. It was used in the molecular phylogenetic study of
|
||
<bibRefCitation id="5829FCF0D84DC4676E4D62442A9EA5AF" DOI="https://doi.org/10.1071/IS02044" author="Maddison, WP" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="1" pagination="529 - 549" refId="B39" refString="Maddison, WP, Hedin, MC, 2003. Jumping spider phylogeny (Araneae: Salticidae). Invertebrate Systematics 17: 529 - 549, DOI: https://doi.org/10.1071/IS02044" title="Jumping spider phylogeny (Araneae: Salticidae)." url="https://doi.org/10.1071/IS02044" volume="17" year="2003">Maddison and Hedin (2003)</bibRefCitation>
|
||
under the name "
|
||
<taxonomicName id="1CBBFF9EA622E636AC3CCCB939D993E3" authorityName="Simon" authorityYear="1901" class="Arachnida" family="Salticidae" genus="Jollas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Jollas" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9D7C6509F3DFBFC6DF3702184C2BE49D" italics="true" pageId="0" pageNumber="1">Jollas</emphasis>
|
||
</taxonomicName>
|
||
sp." (voucher S162) from Sucumbios, Ecuador.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="276E12E4BFC49DAA8AA8476FE902CD51" pageId="0" pageNumber="1" type="diagnosis">
|
||
<paragraph id="6E5F90F7078D91ED6CA0585EDFC7F0C3" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
||
<paragraph id="E77B8BD8892CE257A48AB1CEE6681A0F" pageId="0" pageNumber="1">
|
||
Differs from the very similar
|
||
<taxonomicName id="0180FBE3300AF76D651FDFE29C597D3E" authorityName="Galiano" authorityYear="1991" class="Arachnida" family="Salticidae" genus="Jollas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Jollas puntalara" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="puntalara">
|
||
<emphasis id="21DA5C1B04BDB27CC3AFEBB0925A43C9" italics="true" pageId="0" pageNumber="1">Jollas puntalara</emphasis>
|
||
</taxonomicName>
|
||
Galiano, 1991 in the thinner and straighter RTA and the angle at which the embolus arises. The RTA is more or less straight until a curl at the tip, but it narrows dramatically for its terminal three quarters (Fig.
|
||
<figureCitation id="7DF1581DBC0F58E763F96AC6925E30B1" captionStart="Figures 108–119" captionStartId="F11" captionText="Figures 108 - 119. Jollas cupreus, sp. nov. (except 112, J. puntalara) 108, 109 Left palp of holotype 108 ventral view 109 retrolateral view 110 ventral view of epigyne of paratype 111 dorsal view of same, cleared 112 palp of holotype of J. puntalara Galiano 113 - 115 holotype male 116 male from Yasuni, Ecuador (- 0.675, - 76.397) 117 holotype male in alcohol 118, 119 paratype female." figureDoi="10.3897/zookeys.925.39691.figures108-119" httpUri="https://binary.pensoft.net/fig/397993" pageId="0" pageNumber="1">109</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="1E584FB60B61ED7F122FF06C0BB07D3D" lsidName="J. puntalara" pageId="0" pageNumber="1" rank="species" species="puntalara">
|
||
<emphasis id="EDDCAE1ADC44EDBB27A0A3FD42943335" italics="true" pageId="0" pageNumber="1">J. puntalara</emphasis>
|
||
</taxonomicName>
|
||
(Galiano, 1991b: fig. 26) it bends at the midpoint and thins much less dramatically. The embolus of
|
||
<taxonomicName id="184D5AD38880DD587FE0DB79697D1796" lsidName="J. cupreus" pageId="0" pageNumber="1" rank="species" species="cupreus">
|
||
<emphasis id="18C3D2D0CE2AC20BC843166EC2D42D44" italics="true" pageId="0" pageNumber="1">J. cupreus</emphasis>
|
||
</taxonomicName>
|
||
, as it arises, proceeds directly to the prolateral, thus generating an angle in the retrolateral-basal corner of the bulb (like a chin pointing to the retrolateral), while the embolus of
|
||
<taxonomicName id="BB4223AFFFF95C59163CA822D01ED41D" lsidName="J. puntalara" pageId="0" pageNumber="1" rank="species" species="puntalara">
|
||
<emphasis id="ECD1D59A96EAB77AEA005BF90A718F9E" italics="true" pageId="0" pageNumber="1">J. puntalara</emphasis>
|
||
</taxonomicName>
|
||
emerges angled toward the basal, leaving the bulb more rounded (arrow in Fig.
|
||
<figureCitation id="5D8AD07B990D9402FE5B8045D2FA7F77" captionStart="Figures 108–119" captionStartId="F11" captionText="Figures 108 - 119. Jollas cupreus, sp. nov. (except 112, J. puntalara) 108, 109 Left palp of holotype 108 ventral view 109 retrolateral view 110 ventral view of epigyne of paratype 111 dorsal view of same, cleared 112 palp of holotype of J. puntalara Galiano 113 - 115 holotype male 116 male from Yasuni, Ecuador (- 0.675, - 76.397) 117 holotype male in alcohol 118, 119 paratype female." figureDoi="10.3897/zookeys.925.39691.figures108-119" httpUri="https://binary.pensoft.net/fig/397993" pageId="0" pageNumber="1">112</figureCitation>
|
||
). These differences are small but consistent, insofar as all Ecuadorian specimens show the distinct
|
||
<normalizedToken id="196D1407486F913D9D03DBD3EADFF246" originalValue="“chin”">"chin"</normalizedToken>
|
||
at the base of the embolus and the narrower RTA. It might usually be conservative to leave such close forms as a single species, but given that there is considerable data (molecular phylogenetic and chromosome) attached to the Ecuadorian form, it is safer to name it and thus provide an unambiguous anchor for these data. (Cristian Grismado kindly supplied photographs of
|
||
<bibRefCitation id="9D0C6BAAE1512F2CF4FC8CBEE41E049D" author="Galiano, ME" journalOrPublisher="Physis, Revista de la Sociedad Argentina de Ciencias Naturales (C)" pageId="0" pageNumber="1" pagination="15 - 29" refId="B15" refString="Galiano, ME, 1991b. Revision del genero Jollas (Araneae, Salticidae). Physis, Revista de la Sociedad Argentina de Ciencias Naturales (C) 47: 15 - 29" title="Revision del genero Jollas (Araneae, Salticidae)." volume="47" year="1991 b">
|
||
<normalizedToken id="F0B120F4BCA96A3EE4F7C0ACE14B8CBD" originalValue="Galiano’s">Galiano's</normalizedToken>
|
||
(1991b)
|
||
</bibRefCitation>
|
||
holotype of
|
||
<taxonomicName id="B0EE0471F9A1C657AABE1CFA49A35087" authorityName="Galiano" authorityYear="1991" class="Arachnida" family="Salticidae" genus="Jollas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Jollas puntalara" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="puntalara">
|
||
<emphasis id="39E91E527ACD52AD803D5BFD7193460A" italics="true" pageId="0" pageNumber="1">Jollas puntalara</emphasis>
|
||
</taxonomicName>
|
||
to facilitate our comparison, although these differences can be seen as well in her figures 26 and 29.)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7C48C408828F17E8D876CCC8E242A042" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="67864802C62F3C1649B5945D09BDF46B" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="A61712CB549833035EF36DDA0D1DF595" pageId="0" pageNumber="1">
|
||
<emphasis id="5773C102311E749735EF50E48A86A951" bold="true" pageId="0" pageNumber="1">Male</emphasis>
|
||
(holotype). Carapace length 1.37; abdomen length 1.16.
|
||
<emphasis id="6C267995B9D7A82CF7F0D98A670FB9D2" bold="true" italics="true" pageId="0" pageNumber="1">Carapace</emphasis>
|
||
orange with a black ocular area, mostly glabrous, with only a few scattered setae.
|
||
<emphasis id="5EAC59F7E0B8A0D95445B1F222F522EF" bold="true" italics="true" pageId="0" pageNumber="1">Clypeus</emphasis>
|
||
orange-brown.
|
||
<emphasis id="CDECBD0949A57A5BA9ED5C652E857F14" bold="true" italics="true" pageId="0" pageNumber="1">Chelicerae</emphasis>
|
||
vertical, orange.
|
||
<emphasis id="61947C0A8E70D1F798C30EFA2D3614A5" bold="true" italics="true" pageId="0" pageNumber="1">Palp</emphasis>
|
||
orange-brown except for dark brown cymbium, with dark setae except brilliant white patch of setae dorsally on patella.
|
||
<emphasis id="84DD188016270F9BEC3D90C841D0561A" bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
|
||
long, especially the first and fourth. Legs honey coloured to orange-brown except for a strong black line on prolateral-ventral face of first patella, tibia and metatarsus.
|
||
<taxonomicName id="5B2A7DD282223010E6763FBEDC3C2D04" class="Holothuroidea" family="Molpadiidae" genus="Embolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Embolus" order="Molpadida" pageId="0" pageNumber="1" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="607608B9963D3C1B832AC1D6979C06B5" bold="true" italics="true" pageId="0" pageNumber="1">Embolus</emphasis>
|
||
</taxonomicName>
|
||
arises at ca. 5
|
||
<normalizedToken id="D9BF7780AD237D58F4415FF02C20334B" originalValue="o’clock">o'clock</normalizedToken>
|
||
and curls half-way around bulb. Tibia somewhat bulbous, broad, with bases of setae on retrolateral side forming row of tubercles. RTA begins broad but then narrows abruptly at ca. one quarter its length, from which point it proceeds straight until just before the tip, where it curls.
|
||
<emphasis id="5A51E827CA3F683D57F2BC071B24F038" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
|
||
orange-brown, with black scalloped patch covering dorsum, covered with metallic scales. A patch of bright white setae sits above the anal tubercle.
|
||
</paragraph>
|
||
<paragraph id="3B03BF3B8B4F0C96621FD931ABB9BD59" pageId="0" pageNumber="1">
|
||
<emphasis id="F6E100027D84A817661A6AA9AF6E1068" bold="true" pageId="0" pageNumber="1">Female</emphasis>
|
||
(paratype).
|
||
<emphasis id="3D0C52118F20F6CD31FEA3752311513F" bold="true" italics="true" pageId="0" pageNumber="1">Carapace</emphasis>
|
||
length 1.36;
|
||
<emphasis id="58AAA500BFBCC57D43E4FA397A5C8469" bold="true" italics="true" pageId="0" pageNumber="1">abdomen</emphasis>
|
||
length 1.89. Much darker than the male in body and appendages (Figs
|
||
<figureCitation id="B809A1F3EDEF48E82C507F3ADAEBF5A9" captionStart="Figures 129–139" captionStartId="F13" captionText="Figures 129 - 139. Chromosomes of first meiotic division in males of the Jollas - Tomis clade 129, 130 Attinella concolor, with only seven pairs of autosomes, but each two-armed, 14 m + Xm 0, Florida (29.63 N, 82.37 W) 131 Tomis manabita, showing the two Xs off to one pole, and 13 acrocentric bivalents on the metaphase plate, Ecuador (0.9 S, 80.5 W) 132 - 136 Sittisax ranieri, whose distinctive XmXaYm appears as a rabbit head with a droopy ear. White triangles show points where two bivalents are apparently linked together 134 - 136 details of XXY of S. ranieri 137 - 139 Sittisax saxicola, with sex chromsomes, interpreted tentatively as XaXaXaYm, appearing as a rabbit head with three ears, Switzlerland (46.9 N, 9.2 E)." figureDoi="10.3897/zookeys.925.39691.figures129-139" httpUri="https://binary.pensoft.net/fig/397995" pageId="0" pageNumber="1">130</figureCitation>
|
||
,
|
||
<figureCitation id="C5BF1F80AD952F52FE90DA1603D5ECAB" captionStart="Figures 129–139" captionStartId="F13" captionText="Figures 129 - 139. Chromosomes of first meiotic division in males of the Jollas - Tomis clade 129, 130 Attinella concolor, with only seven pairs of autosomes, but each two-armed, 14 m + Xm 0, Florida (29.63 N, 82.37 W) 131 Tomis manabita, showing the two Xs off to one pole, and 13 acrocentric bivalents on the metaphase plate, Ecuador (0.9 S, 80.5 W) 132 - 136 Sittisax ranieri, whose distinctive XmXaYm appears as a rabbit head with a droopy ear. White triangles show points where two bivalents are apparently linked together 134 - 136 details of XXY of S. ranieri 137 - 139 Sittisax saxicola, with sex chromsomes, interpreted tentatively as XaXaXaYm, appearing as a rabbit head with three ears, Switzlerland (46.9 N, 9.2 E)." figureDoi="10.3897/zookeys.925.39691.figures129-139" httpUri="https://binary.pensoft.net/fig/397995" pageId="0" pageNumber="1">131</figureCitation>
|
||
).
|
||
<emphasis id="CD8520218C34D64185CAE066C0E2F09D" bold="true" italics="true" pageId="0" pageNumber="1">Carapace</emphasis>
|
||
dark brown, black in ocular area, sparsely covered with paler scales.
|
||
<emphasis id="BFB588393F074E954CBE916D65842D02" bold="true" italics="true" pageId="0" pageNumber="1">Clypeus</emphasis>
|
||
and
|
||
<emphasis id="243019FE639AE64232614A05A23143D8" bold="true" italics="true" pageId="0" pageNumber="1">chelicerae</emphasis>
|
||
brown, more or less glabrous.
|
||
<emphasis id="421D1862EDEDF9FFAF9CDDE6BCF7F75B" bold="true" italics="true" pageId="0" pageNumber="1">Chelicerae</emphasis>
|
||
brown, more or less.
|
||
<taxonomicName id="8C7F342FD637D7AE7477DE12BBACA720" genus="Palps" lsidName="Palps" pageId="0" pageNumber="1" rank="genus">
|
||
<emphasis id="558B61033F4E7E82E0F34371CFDAAACE" bold="true" italics="true" pageId="0" pageNumber="1">Palps</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B6D4324FF892245F32B86156D79C095D" bold="true" italics="true" pageId="0" pageNumber="1">legs</emphasis>
|
||
honey coloured but with strongly contrasting black markings: annulae at joints, black stripes or patches on front and back faces of femora, and black stripe on front face of first and second tibiae. Abdomen black but with reflective metallic scales.
|
||
<emphasis id="E1D9D3CBB9684BC1EBC0E1B3CE5977F9" bold="true" italics="true" pageId="0" pageNumber="1">Epigyne</emphasis>
|
||
(Figs
|
||
<figureCitation id="58B2E062FAE75948D109E80259E6FBEC" captionStart="Figures 108–119" captionStartId="F11" captionText="Figures 108 - 119. Jollas cupreus, sp. nov. (except 112, J. puntalara) 108, 109 Left palp of holotype 108 ventral view 109 retrolateral view 110 ventral view of epigyne of paratype 111 dorsal view of same, cleared 112 palp of holotype of J. puntalara Galiano 113 - 115 holotype male 116 male from Yasuni, Ecuador (- 0.675, - 76.397) 117 holotype male in alcohol 118, 119 paratype female." figureDoi="10.3897/zookeys.925.39691.figures108-119" httpUri="https://binary.pensoft.net/fig/397993" pageId="0" pageNumber="1">110</figureCitation>
|
||
,
|
||
<figureCitation id="40609CD40F220F68B97A625D67C46EC8" captionStart="Figures 108–119" captionStartId="F11" captionText="Figures 108 - 119. Jollas cupreus, sp. nov. (except 112, J. puntalara) 108, 109 Left palp of holotype 108 ventral view 109 retrolateral view 110 ventral view of epigyne of paratype 111 dorsal view of same, cleared 112 palp of holotype of J. puntalara Galiano 113 - 115 holotype male 116 male from Yasuni, Ecuador (- 0.675, - 76.397) 117 holotype male in alcohol 118, 119 paratype female." figureDoi="10.3897/zookeys.925.39691.figures108-119" httpUri="https://binary.pensoft.net/fig/397993" pageId="0" pageNumber="1">111</figureCitation>
|
||
) with distinctive dark inverted
|
||
<normalizedToken id="DA0764C7D7BFE3249CACF3E1366C3156" originalValue="“V”">"V"</normalizedToken>
|
||
in which are the narrow openings into the copulatory ducts, though lateral pockets may lead the embolus to the openings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C280AAF54DD8F42985819CB1CF059AA0" pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph id="46A9DFBE1F45127818C39FCD07F5C4AA" pageId="0" pageNumber="1">Additional material.</paragraph>
|
||
<paragraph id="574B16C6CF71FF7C6C8E5FA8A9568DBD" pageId="0" pageNumber="1">
|
||
22 males and 6 females from: Ecuador: Napo: Tarapoa. 23 June - 1 July 1988 W. Maddison WPM#88-002 (1 male); Ecuador: Napo: bridge over Rio Cuyabeno on road to Tipishca. 25-30 June 1988 W. Maddison WPM#88-004 (1 male 1 female); Ecuador: Napo: bridge over Rio Cuyabeno on road to Tipishca. 29-30 July 1988 W. Maddison WPM#88-018 (4 males 2 females); Ecuador: Napo: Reserva Faunistica de Cuyabeno, Laguna Grande, Sendero La Hormiga. 2-5 August 1988 W. Maddison WPM#88-023 (2 males); Ecuador: Napo: Reserva Faunistica de Cuyabeno, Laguna Grande, PUCE field station. 1-7 August 1988 W. Maddison WPM#88-025 (1 male); Ecuador: Napo: bridge over Rio Cuyabeno on road from Lago Agrio to Tipishca. 8-9 August 1988 W. Maddison WPM#88-027 (1 male); Ecuador: Sucumbios: Reserva Faunistica Cuyabeno, Laguna Grande, PUCE field station.
|
||
<geoCoordinate id="9360EA23E3F2C42CBD1864C46AADF233" degrees="0.002" direction="north" orientation="latitude" precision="55" value="0.002">0.002</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="98403CFA7AACA67C24D28F42C39A1201" degrees="76.172" direction="west" orientation="longitude" precision="55" value="-76.172">-76.172</geoCoordinate>
|
||
. 21-29 July 1989 W. Maddison WPM#89-032 (1 male); Ecuador: Sucumbios: bridge over Rio Cuyabeno on road between Tarapoa and Tipishca,
|
||
<geoCoordinate id="B57218A9031F77CD6326EA01CFC2AACE" degrees="0.025" direction="north" orientation="latitude" precision="55" value="0.025">0.025</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E5858B38C058504DB07215E5D09B61AE" degrees="77.308" direction="west" orientation="longitude" precision="55" value="-77.308">-77.308</geoCoordinate>
|
||
. 29 July 1989 W. Maddison WPM#89-036 (1 male); Ecuador: Sucumbios: Reserva Faunistica Cuyabeno, Nuevo Mundo cabins along Rio Cuyabeno at jcn with Lago Agrio-Tipishca HWY 19-29 April 1994 W. Maddison WPM#94-021 (3 males); Ecuador: Sucumbios: Reserva Faunistica Cuyabeno, Nuevo Mundo cabins, jcn Rio Cuyabeno & Lago Agrio-Tipishca HWY tree trunks 19-29 April 1994 W. Maddison WPM#94-023 (1 male); Ecuador: Morona Santiago: km 3 from
|
||
<normalizedToken id="7F0FCB17DFC6D3D10AEB77CB9FAE50F2" originalValue="Limón">Limon</normalizedToken>
|
||
towards Gualaceo.
|
||
<geoCoordinate id="CFB9BB9BBC6AA036BE19AE6C51AF5130" degrees="2.9663" direction="north" orientation="latitude" precision="5" value="2.9663">2.9663</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="0068CBD25ED64D377AF20294BFA8EC07" degrees="78.4209" direction="west" orientation="longitude" precision="5" value="-78.4209">-78.4209</geoCoordinate>
|
||
; 1250 m el. 12 July 2004 Maddison, Agnarsson, Iturralde, Salazar. WPM#04-030 (1 male 2 females); Ecuador: Morona Santiago: km 4 from
|
||
<normalizedToken id="DDD9BD81977454ABBA3AB0691756F2DF" originalValue="Limón">Limon</normalizedToken>
|
||
towards Gualaceo.
|
||
<geoCoordinate id="A717AF3D1674ABFC25F006DD59CF42F8" degrees="2.9808" direction="north" orientation="latitude" precision="5" value="2.9808">2.9808</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D263C0BE8A4A8282D6371CB28581AC78" degrees="78.4414" direction="west" orientation="longitude" precision="5" value="-78.4414">-78.4414</geoCoordinate>
|
||
; 1380 m el. 12 July 2004 Maddison, Agnarsson, Iturralde, Salazar. WPM#04-031 (2 males); Ecuador: Orellana:
|
||
<normalizedToken id="DC48B8DBED0F44E9B169B3EF64955959" originalValue="Yasuní">Yasuni</normalizedToken>
|
||
Res.Stn.area, Station area
|
||
<geoCoordinate id="51FCEFA30EEA5D3B9413CAB439567D84" degrees="0.675" direction="north" orientation="latitude" precision="55" value="0.675">0.675</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5F588D71704206375A73A8C96BFD1C96" degrees="76.397" direction="west" orientation="longitude" precision="55" value="-76.397">-76.397</geoCoordinate>
|
||
210-280 m elev. 26 July - 13 Aug 2011 Maddison/Piascik/Vega WPM#11-015 (2 males); Ecuador: Orellana:
|
||
<normalizedToken id="B865F67FB0A4F52A28398AD4BFC14EDB" originalValue="Yasuní">Yasuni</normalizedToken>
|
||
Res.Stn.area, Station area
|
||
<geoCoordinate id="EDE9220D79F9DACDDCF791E0311138D3" degrees="0.674" direction="north" orientation="latitude" precision="55" value="0.674">0.674</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="0BDD2ED80ABE1180BAA4A4C8746D58C5" degrees="76.397" direction="west" orientation="longitude" precision="55" value="-76.397">-76.397</geoCoordinate>
|
||
210-280 m elev. Clearings, forest edge 8-9.Aug.2011 Maddison/ Piascik/Vega. WPM#11-104 (1 male); Ecuador: Orellana:
|
||
<normalizedToken id="A98DB88729CBAA69E04DD99A48C59B22" originalValue="Río">Rio</normalizedToken>
|
||
Bigal Reserve, boundary along road.
|
||
<geoCoordinate id="0CC874BCE92ABF18D9C5BC9512B6453F" degrees="0.541" direction="north" orientation="latitude" precision="55" value="0.541">0.541</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="4FCA244603198F2305A3D06F642A0E44" degrees="77.424" direction="west" orientation="longitude" precision="55" value="-77.424">-77.424</geoCoordinate>
|
||
. 970 m elev. 5 November 2010. M Vega, D & W Maddison, M Reyes. WPM#10-048 (1 female). (Note: the province Sucumbios was established after 1988; the 1988 localities listed as Napo Province would now all be in Sucumbios.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |