241 lines
18 KiB
XML
241 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.738.22273" ID-GBIF-Dataset="f3cd9500-2125-4323-9e78-e42a6aa9b653" ID-PMC="PMC5904432" ID-Pensoft-Pub="1313-2970-738-67" ID-PubMed="29670422" ID-ZBK="D31A93AFB7024C44BB5379B377FFB9CD" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-738-67" ModsDocOrigin="ZooKeys 738" ModsDocTitle="Description of a new species of Coelosis Hope from Guajira Peninsula, northern Colombia (Coleoptera, Scarabaeidae, Dynastinae, Oryctini)" checkinTime="1519083490447" checkinUser="pensoft" docAuthor="Neita-Moreno, Jhon Cesar, Orozco, Jesus & Medina-Uribe, Claudia Alejandra" docDate="2018" docId="E48ED4B70BB92649339D3BB145350508" docLanguage="en" docName="ZooKeys 738: 67-80" docOrigin="ZooKeys 738" docSource="http://dx.doi.org/10.3897/zookeys.738.22273" docTitle="Coelosis wayuorum Neita-Moreno, Orozco & Medina-Uribe, 2018, sp. n." docType="treatment" docUuid="863A21DA-0365-4404-B1BF-C125EE00B609" docUuidSource="ZooBank" docVersion="5" lastPageNumber="73" masterDocId="FFBBFFF8FFEFFF92A50A35360576FF80" masterDocTitle="Description of a new species of Coelosis Hope from Guajira Peninsula, northern Colombia (Coleoptera, Scarabaeidae, Dynastinae, Oryctini)" masterLastPageNumber="80" masterPageNumber="67" pageNumber="69" updateTime="1668165458065" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Description of a new species of Coelosis Hope from Guajira Peninsula, northern Colombia (Coleoptera, Scarabaeidae, Dynastinae, Oryctini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neita-Moreno, Jhon Cesar</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Orozco, Jesus</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Medina-Uribe, Claudia Alejandra</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>738</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>67</mods:start>
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<mods:end>80</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.738.22273</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.738.22273</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-738-67</mods:identifier>
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<mods:identifier type="ZBK">D31A93AFB7024C44BB5379B377FFB9CD</mods:identifier>
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<mods:identifier type="ZooBank">D31A93AFB7024C44BB5379B377FFB9CD</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="141117776" LSID="urn:lsid:zoobank.org:act:863A21DA-0365-4404-B1BF-C125EE00B609" httpUri="http://treatment.plazi.org/id/E48ED4B70BB92649339D3BB145350508" lastPageId="6" lastPageNumber="73" pageId="2" pageNumber="69">
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<subSubSection pageId="2" pageNumber="69" type="nomenclature">
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<paragraph pageId="2" pageNumber="69">
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<taxonomicName LSID="http://zoobank.org/863A21DA-0365-4404-B1BF-C125EE00B609" class="Insecta" family="Dynastidae" genus="Coelosis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelosis wayuorum" order="Coleoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="wayuorum">Coelosis wayuorum</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="69">sp. n.</taxonomicNameLabel>
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Figs 3
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<normalizedToken originalValue="F–G">F-G</normalizedToken>
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; 4
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<normalizedToken originalValue="G–I">G-I</normalizedToken>
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; 5C, F, I, L; 6C, E, G, J, K, M; 7
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<normalizedToken originalValue="E–F">E-F</normalizedToken>
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; 8
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="69" type="type material">
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<paragraph pageId="2" pageNumber="69">Type material</paragraph>
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<paragraph pageId="2" pageNumber="69">
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(3). Holotype labeled "Colombia, La Guajira, Uribia, PNN La/Macuira Corregimiento Nazareth/
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<normalizedToken originalValue="Kajashiwoü">Kajashiwoue</normalizedToken>
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,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="12.193861">12°11'37.9"N</geoCoordinate>
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; 71°21'30.1"/W. WGS84, 70 m. Manual.19.ix.2014/C. Medina" [IAvH-E-195379]. Allotype [IAvH-E-195380] and one female paratype [IAvH-E-195381] with the same label data. Types deposited at the Instituto Alexander von Humboldt (IAvH) Villa de Leyva,
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<normalizedToken originalValue="Boyacá">Boyaca</normalizedToken>
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, Colombia.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="69" type="diagnosis">
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<paragraph pageId="2" pageNumber="69">Diagnosis.</paragraph>
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<paragraph pageId="2" pageNumber="69">
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<taxonomicName class="Insecta" family="Dynastidae" genus="Coelosis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelosis wayuorum" order="Coleoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="wayuorum">Coelosis wayuorum</taxonomicName>
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sp. n. can be separated from the other Colombian
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<taxonomicName class="Insecta" family="Dynastidae" genus="Coelosis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelosis" order="Coleoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="genus">Coelosis</taxonomicName>
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by the following characters: maxilla with lateral sclerite pronounced (Fig. 5C) [similar to
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<taxonomicName lsidName="C. bicornis" pageId="2" pageNumber="69" rank="species" species="bicornis">C. bicornis</taxonomicName>
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, but not as in
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<taxonomicName lsidName="C. biloba" pageId="2" pageNumber="69" rank="species" species="biloba">C. biloba</taxonomicName>
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]; galea without teeth (Fig. 5C) [two teeth in both
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<taxonomicName lsidName="C. bicornis" pageId="2" pageNumber="69" rank="species" species="bicornis">C. bicornis</taxonomicName>
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and
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<taxonomicName lsidName="C. biloba" pageId="2" pageNumber="69" rank="species" species="biloba">C. biloba</taxonomicName>
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]; ventral surface of the mandibles with keels 1 and 2 contiguous (Fig. 4H) [widely separate in
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<taxonomicName lsidName="C. bicornis" pageId="2" pageNumber="69" rank="species" species="bicornis">C. bicornis</taxonomicName>
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(Fig. 4B), slightly separate in
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<taxonomicName lsidName="C. biloba" pageId="2" pageNumber="69" rank="species" species="biloba">C. biloba</taxonomicName>
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(Figs 4E)]; pronotum and elytra strongly punctate, punctures with short, spine-like setae (Figs 6C, E) [spine-like setae absent in both
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<taxonomicName lsidName="C. bicornis" pageId="2" pageNumber="69" rank="species" species="bicornis">C. bicornis</taxonomicName>
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and
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<taxonomicName lsidName="C. biloba" pageId="2" pageNumber="69" rank="species" species="biloba">C. biloba</taxonomicName>
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(Fig. 6A, B, D)]; prohypomeron with spine-like setae [slender and long in
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<taxonomicName lsidName="C. bicornis" pageId="2" pageNumber="69" rank="species" species="bicornis">C. bicornis</taxonomicName>
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, slender and short in
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<taxonomicName lsidName="C. biloba" pageId="2" pageNumber="69" rank="species" species="biloba">C. biloba</taxonomicName>
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]; mesosternum convex as in
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<taxonomicName lsidName="C. biloba" pageId="2" pageNumber="69" rank="species" species="biloba">C. biloba</taxonomicName>
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[concave in
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<taxonomicName lsidName="C. bicornis" pageId="2" pageNumber="69" rank="species" species="bicornis">C. bicornis</taxonomicName>
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]; metasternum covered with short, spine-like setae (Fig. 6G) [setae long and slender in both
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<taxonomicName lsidName="C. bicornis" pageId="2" pageNumber="69" rank="species" species="bicornis">C. bicornis</taxonomicName>
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and
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<taxonomicName lsidName="C. biloba" pageId="2" pageNumber="69" rank="species" species="biloba">C. biloba</taxonomicName>
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(Fig. 6F)]; and meso- and metatibiae densely punctate (Fig. 6J, K) [scarcely punctate in both
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<taxonomicName lsidName="C. bicornis" pageId="2" pageNumber="69" rank="species" species="bicornis">C. bicornis</taxonomicName>
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and
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<taxonomicName lsidName="C. biloba" pageId="2" pageNumber="69" rank="species" species="biloba">C. biloba</taxonomicName>
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(Fig. 6H, I)]. The internal sac is different among the species: in
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<taxonomicName lsidName="C. biloba" pageId="2" pageNumber="69" rank="species" species="biloba">C. biloba</taxonomicName>
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, the accessorial lamella is short and simple (Fig. 5H), while in
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<taxonomicName lsidName="C. bicornis" pageId="2" pageNumber="69" rank="species" species="bicornis">C. bicornis</taxonomicName>
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and
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<taxonomicName lsidName="C. wayuorum" pageId="2" pageNumber="69" rank="species" species="wayuorum">C. wayuorum</taxonomicName>
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sp. n. the lamella is long and complex, although with differences between these two species (Fig. 5G, I).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="73" pageId="2" pageNumber="69" type="description">
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<paragraph pageId="2" pageNumber="69">Description.</paragraph>
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<paragraph lastPageId="6" lastPageNumber="73" pageId="2" pageNumber="69">
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Holotype male (Fig. 3F). Body length 20.2 mm; width 11.00 mm. Color dark reddish brown. Head: Frontoclypeal region with small horn, surface rugopunctate, punctures setose, setae spine-like (Fig. 3F). Clypeus strongly rugopunctate, slightly emarginate, apex broad with two reflexed teeth. Mandibles with three conical teeth, subapical notch slightly deep (Fig. 4G, H); labium rugopunctate, with sparse, short, spine-like setae, paraglossa undeveloped, narrow, apex truncate (Fig. 4I); maxilla with lateral projection, galea without teeth (Fig. 5C). Pronotum: Surface sparsely punctate; punctures moderately large, umbilicate, setose; setae spine-like (Fig. 6C). Disc with two small, widely separated horns, wide fovea between horns (Figs 3F, 6C).
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<pageBreakToken pageId="3" pageNumber="70" start="start">Elytra</pageBreakToken>
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: Surface with 10 distinct pairs of striae composed of ocellate punctures bearing short, spine-like setae, micropunctures densely intermixed between striae (Fig. 6E). Pygidium: Surface sparsely punctate, punctures small to moderate in size, weakly ocellate,
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<pageBreakToken pageId="4" pageNumber="71" start="start">with</pageBreakToken>
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short spine-like setae, becoming denser at basal angles. Surface regularly convex in lateral view. Legs: Protibia tridentate, basal tooth reduced. Meso- and metatibiae with one medial transverse carina, each with short, spine-like setae and one small inner
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<pageBreakToken pageId="5" pageNumber="72" start="start">tooth</pageBreakToken>
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(Fig. 6J, K). First metatarsomere apically expanded, apex subtruncate with acute outer projection (Fig. 6M). Venter: Prosternal process subtriangular, thick; apex short, parabolic, with process at middle, this process with short, stout setae (Fig. 5F). Mesosternum densely punctate, setose, slightly convex at middle. Metasternum densely
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<pageBreakToken pageId="6" pageNumber="73" start="start">punctate</pageBreakToken>
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; punctures ocellate, minutely setose; setae spine‒like; lateral edge rugopunctate with short, spine-like setae. Abdominal ventrite VIII depressed at middle. Genitalia: Parameres as in Fig. 7E, F, internal sac as in Fig. 5I.
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</paragraph>
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<caption pageId="6" pageNumber="73">
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<paragraph pageId="6" pageNumber="73">
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Figure 3.
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<taxonomicName class="Insecta" family="Dynastidae" genus="Coelosis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelosis" order="Coleoptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="genus">Coelosis</taxonomicName>
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species from Colombia (dorsal habitus): A (male major) B (female) C (male minor):
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<taxonomicName lsidName="C. bicornis" pageId="6" pageNumber="73" rank="species" species="bicornis">C. bicornis</taxonomicName>
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D (male major):
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<taxonomicName lsidName="C. biloba" pageId="6" pageNumber="73" rank="species" species="biloba">C. biloba</taxonomicName>
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(L.) E (female):
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<taxonomicName lsidName="C. biloba" pageId="6" pageNumber="73" rank="species" species="biloba">C. biloba</taxonomicName>
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F Holotype (male):
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<taxonomicName lsidName="C. wayuorum" pageId="6" pageNumber="73" rank="species" species="wayuorum">C. wayuorum</taxonomicName>
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sp. n. G Allotype (female):
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<taxonomicName lsidName="C. wayuorum" pageId="6" pageNumber="73" rank="species" species="wayuorum">C. wayuorum</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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<caption pageId="6" pageNumber="73">
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<paragraph pageId="6" pageNumber="73">
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Figure 4. A, B, C
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<taxonomicName class="Insecta" family="Dynastidae" genus="Coelosis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelosis bicornis" order="Coleoptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="species" species="bicornis">Coelosis bicornis</taxonomicName>
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D, E, F
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<taxonomicName lsidName="C. biloba" pageId="6" pageNumber="73" rank="species" species="biloba">C. biloba</taxonomicName>
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G, H, I
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<taxonomicName lsidName="C. wayuorum" pageId="6" pageNumber="73" rank="species" species="wayuorum">C. wayuorum</taxonomicName>
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sp. n. A, D, G mandibles (dorsal view) B, E, H mandibles (ventral view) C, F, I labium (ventral view).
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</paragraph>
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</caption>
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<caption pageId="6" pageNumber="73">
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<paragraph pageId="6" pageNumber="73">
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Figure 5. A, D, G, J
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<taxonomicName class="Insecta" family="Dynastidae" genus="Coelosis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelosis bicornis" order="Coleoptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="species" species="bicornis">Coelosis bicornis</taxonomicName>
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B, E, H, K
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<taxonomicName lsidName="C. biloba" pageId="6" pageNumber="73" rank="species" species="biloba">C. biloba</taxonomicName>
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C, F, I, L
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<taxonomicName lsidName="C. wayuorum" pageId="6" pageNumber="73" rank="species" species="wayuorum">C. wayuorum</taxonomicName>
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sp. n.
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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maxilla (ventral view)
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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prosternal process (antero-posterior view)
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<normalizedToken originalValue="G–I">G-I</normalizedToken>
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internal sac (copulatory lamellae)
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<normalizedToken originalValue="J–L">J-L</normalizedToken>
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genital plate (female).
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</paragraph>
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</caption>
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<caption pageId="6" pageNumber="73">
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<paragraph pageId="6" pageNumber="73">
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Figure 6. A, B, D, F, H, I, L
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<taxonomicName class="Insecta" family="Dynastidae" genus="Coelosis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelosis bicornis" order="Coleoptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="species" species="bicornis">Coelosis bicornis</taxonomicName>
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C, E, G, J, K, M
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<taxonomicName class="Insecta" family="Dynastidae" genus="Coelosis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelosis wayuorum" order="Coleoptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="species" species="wayuorum">Coelosis wayuorum</taxonomicName>
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sp. n.
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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pronotum (dorsal view) D, E elytra F, G metasternum H, J mesotibia (dorsal view) I, K metatibia (dorsal view) L, M first metatarsomere (dorsal view).
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</paragraph>
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</caption>
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<caption pageId="6" pageNumber="73">
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<paragraph pageId="6" pageNumber="73">
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Figure 7. A, B
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<taxonomicName class="Insecta" family="Dynastidae" genus="Coelosis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelosis bicornis" order="Coleoptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="species" species="bicornis">Coelosis bicornis</taxonomicName>
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C, D
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<taxonomicName lsidName="C. biloba" pageId="6" pageNumber="73" rank="species" species="biloba">C. biloba</taxonomicName>
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E, F
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<taxonomicName lsidName="C. wayuorum" pageId="6" pageNumber="73" rank="species" species="wayuorum">C. wayuorum</taxonomicName>
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sp. n. A, C, E parameres (frontral view) B, D, F phallobase (lateral view).
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</paragraph>
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</caption>
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<caption pageId="6" pageNumber="73">
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<paragraph pageId="6" pageNumber="73">
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Figure 8. Distribution of
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<taxonomicName class="Insecta" family="Dynastidae" genus="Coelosis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelosis" order="Coleoptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="genus">Coelosis</taxonomicName>
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in Colombia.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="73" type="allotype">
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<paragraph pageId="6" pageNumber="73">Allotype</paragraph>
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<paragraph pageId="6" pageNumber="73">(Fig. 3G). Female. Similar to holotype except for the following: body length 22.4 mm; width 12.0 mm, frons and disc of pronotum smooth, without fovea or horns, pygidium slightly concave in lateral view, and genital plate as Fig. 5L.</paragraph>
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<paragraph pageId="6" pageNumber="73">Paratype (1). Similar to allotype except for the following: body length 25.5 mm; width 13.0 mm.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="73" type="etymology">
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<paragraph pageId="6" pageNumber="73">Etymology.</paragraph>
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<paragraph pageId="6" pageNumber="73">
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The specific epithet
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<taxonomicName lsidName="wayuorum" pageId="6" pageNumber="73" rank="species" species="wayuorum">wayuorum</taxonomicName>
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refers to the Wayuu indigenous group inhabiting the Guajira Peninsula.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="73" type="distribution">
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<paragraph pageId="6" pageNumber="73">Distribution.</paragraph>
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<paragraph pageId="6" pageNumber="73">
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<taxonomicName class="Insecta" family="Dynastidae" genus="Coelosis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelosis wayuorum" order="Coleoptera" pageId="6" pageNumber="73" phylum="Arthropoda" rank="species" species="wayuorum">Coelosis wayuorum</taxonomicName>
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sp. n. is known only from one locality in Macuira National Park, Colombia (Fig. 8).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="73" type="distribution">
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<paragraph pageId="6" pageNumber="73">Temporal distribution.</paragraph>
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<paragraph pageId="6" pageNumber="73">Three specimens collected in September 2014, during the dry season.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="73" type="life history">
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<paragraph pageId="6" pageNumber="73">Life history.</paragraph>
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<paragraph pageId="6" pageNumber="73">The type material was collected at night with lights during the dry season.</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |