393 lines
50 KiB
XML
393 lines
50 KiB
XML
<document id="BBCC859B03AF8A4268F9F0A01F1CB5D4" ID-CLB-Dataset="298186" ID-DOI="10.3897/zookeys.1204.114932" ID-GBIF-Dataset="33ca18a1-99c6-4b56-9a8e-92fe6f6f9209" ID-ZooBank="F5134CC6-A83C-48B6-A1EA-235E6858D5CB" ID-publisher-id="114932" URI-arpha="EAF89A3A-8354-506D-AF9B-9E1C53605541" URI-zoobank="http://zoobank.org/F5134CC6-A83C-48B6-A1EA-235E6858D5CB" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1717680124246" checkinUser="pensoft" docAuthor="Nascimento, Elynton Alves & Bocakova, Milada" docDate="2024" docId="E48A6CB9FB205286B7586BEEA6FB4F65" docLanguage="en" docName="ZooKeys 1204: 241-259" docOrigin="ZooKeys 1204" docSource="https://zookeys.pensoft.net/article/114932/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Gorhamium Nascimento & Bocakova 2024, gen. nov." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="259" masterDocId="EAF89A3A8354506DAF9B9E1C53605541" masterDocTitle="Phylogenetic analysis reveals a new net-winged beetle genus of Eurrhacini (Coleoptera, Lycidae) from the Pacific slopes of Central America and Ecuador" masterLastPageNumber="259" masterPageNumber="241" pageNumber="241" updateTime="1717709099581" updateUser="ExternalLinkService">
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<mods:mods id="35B7CF7BBA258C518959F7D093557F69" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4429E21E4A7352EE66E34332CE03640B">
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<mods:title id="6D3A35831DC5098EDAA263F26D2DE05A">Phylogenetic analysis reveals a new net-winged beetle genus of Eurrhacini (Coleoptera, Lycidae) from the Pacific slopes of Central America and Ecuador</mods:title>
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<mods:name id="9671579DDF5A49521B60EEB2EDEC9347" type="personal">
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<mods:roleTerm id="E8D02216CBF535567631B5161BACF9F5">Author</mods:roleTerm>
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<mods:namePart id="6311391885CAE6DA16A8EA5E2974C0EF">Nascimento, Elynton Alves</mods:namePart>
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<mods:nameIdentifier id="5E8751D8280659AB3BB2D003F3A7384B" type="ORCID">0000-0002-9071-2823</mods:nameIdentifier>
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<mods:affiliation id="AF298F930DC8EFB26A7255CF36FCA56A">Departamento de Engenharia Ambiental, Universidade Estadual do Centro-Oeste, Rua Professora Maria Roza Zanon de Almeida, s / n, Engenheiro Gutierrez, Irati – PR, CEP 84505 - 677, Brazil</mods:affiliation>
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<mods:namePart id="D4AE33FE6C467B2EB3EE4A7CCD39A2A1">Bocakova, Milada</mods:namePart>
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<mods:nameIdentifier id="17113D6C2FF46A001B4887A4757EB058" type="ORCID">0000-0002-2507-0887</mods:nameIdentifier>
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<mods:affiliation id="68133E34583D9A18683E71F38F17935A">Department of Biology, Faculty of Education, Palacky University, Purkrabska 2, CZ- 77140 Olomouc, Czech Republic</mods:affiliation>
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<mods:typeOfResource id="DE9A82BA1AA28725EF0657F2215FBB51">text</mods:typeOfResource>
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<mods:title id="1C0F667445BF7288F4CFB8BCA469A393">ZooKeys</mods:title>
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<mods:part id="39CDC4E84F883EF9C29F8FB7199FC1EC">
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<mods:date id="BF6543E0D69C2D47D14CEA112819ECEE">2024</mods:date>
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<mods:number id="CDA5FCAB34BE81209E3705685710A0FD">2024-06-06</mods:number>
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<mods:detail id="F1BE360024BC5A6DCC94B8B50E60E9DC" type="volume">
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<mods:number id="F2F9B7F930219589B15FC2E1FA22E7EA">1204</mods:number>
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<mods:classification id="09DF5002EB41F465A63C077C0941598E">journal article</mods:classification>
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<mods:identifier id="B0C6EF99F5E9E44D36C13CE18F10BBCB" type="DOI">10.3897/zookeys.1204.114932</mods:identifier>
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<treatment id="E48A6CB9FB205286B7586BEEA6FB4F65" ID-DOI="http://doi.org/10.5281/zenodo.11505165" ID-GBIF-Taxon="232157810" ID-Zenodo-Dep="11505165" ID-arpha="E48A6CB9-FB20-5286-B758-6BEEA6FB4F65" ID-zoobank="https://zoobank.org/C31BE6D3-296C-45B1-B685-AD85CB34EA65" LSID="urn:lsid:plazi:treatment:E48A6CB9FB205286B7586BEEA6FB4F65" LSID-ZBK="urn:lsid:zoobank.org:act:C31BE6D3-296C-45B1-B685-AD85CB34EA65" httpUri="http://treatment.plazi.org/id/E48A6CB9FB205286B7586BEEA6FB4F65">
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<subSubSection id="59754AB1D7D086569F0364644C4E786F" type="nomenclature">
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<paragraph id="515C331977EDC32034BF8B19DBA47FE0">
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<taxonomicName id="41D0EDAFD8F1BE08DA829643F1D7B03D" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="C81C4566E0B8ACF0693375B96484CD2D" italics="true">Gorhamium</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F35000D74010B5ECDB26EDB82838AD73" rank="genus">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0EIKAG" type="type_taxon">
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<paragraph id="78347DF3D3DCE8CDC50E99DF29A29C81">
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<heading id="D2D82D74D6EC7889C0691DD38A2B7123" reason="title">
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<typeStatus id="86B53464A2DF85ACF0F4C8119DC9BEC3">Type</typeStatus>
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species.
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</heading>
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</paragraph>
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<paragraph id="D46D29601BCDCED4452D9DB36EF0EAB7">
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<taxonomicName id="16931F7317C220E51FFD3E6274A86E7E" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bidentatum" status="sp. nov.">
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<emphasis id="6CC036A8F1ECB2EAF4598EF64CB1C5D0" italics="true">Gorhamium bidentatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D2B3721C247783AE51A70A63822A1B9C" rank="species">sp. nov.</taxonomicNameLabel>
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(by present designation).
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0EZKAG" type="diagnosis">
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<paragraph id="9D3F36ABD2BF096E4ECE8BFC5AC231E3">
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<heading id="77CC49F3FDF263F3142850EEB2E5EF50" reason="title">Diagnosis.</heading>
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</paragraph>
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<paragraph id="A5C285EAA17822160205A932192E7919">
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<taxonomicName id="1C12A7DE7D279E77545F69D0F53B0A9B" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="3C784A2F8106DB1C8A651816BEA464D9" italics="true">Gorhamium</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5F72E8A5590DEDA9FA0166D7695D7EEC" rank="genus">gen. nov.</taxonomicNameLabel>
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can be distinguished from other
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<taxonomicName id="D0856DC217095343C75EFB61DF351B69" class="Insecta" family="Lycidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="tribe" tribe="Eurrhacini">Eurrhacini</taxonomicName>
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by the combination of the following characters: a) elytra (Fig.
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<figureCitation id="D05DC6E89C64B91E1B63211E09DCF232" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal view A, B Gorhamium bidentatum sp. nov. A male B female C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure2" httpUri="https://binary.pensoft.net/fig/1063776">2 A – C</figureCitation>
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) with nine longitudinal costae (4 costae in
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<taxonomicName id="31B2AE0D1E194457B6C02B8868CACEF9" authorityName="Bourgeois" authorityYear="1879" class="Insecta" family="Lycidae" genus="Haplobothris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
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<emphasis id="4FEEF3BB147912FDF77A13DB8612ED72" italics="true">Haplobothris</emphasis>
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</taxonomicName>
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); b) pronotum (Fig.
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<figureCitation id="FEDDADCCA687D887F9869486709DBCD9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pronotum A Gorhamium bidentatum sp. nov., male B Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure3" httpUri="https://binary.pensoft.net/fig/1063777">3 A, B</figureCitation>
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) wider than long (elongated in
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<taxonomicName id="821DCE8BE0370F9421E56D64BEE6AE32" authorityName="Gorham" authorityYear="1881" class="Insecta" family="Lycidae" genus="Calocladon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
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<emphasis id="05BC51154079876B1E57ED677CAEC49C" italics="true">Calocladon</emphasis>
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</taxonomicName>
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); c) median areola on pronotum slenderly lenticular (slightly wider in
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<taxonomicName id="4A582ADD0347F2597BD9FB78BB484AD9" authorityName="Nascimento & Bocakova" authorityYear="2022" class="Insecta" family="Lycidae" genus="Cladocalon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
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<emphasis id="02D9DEABFAA662240C247ABD9844A0A9" italics="true">Cladocalon</emphasis>
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</taxonomicName>
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and
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<emphasis id="756C0D420F39446B2049031074FC9005" italics="true">Atlanticolycus</emphasis>
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); d) male antennomere 3–10 flabellate (Fig.
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<figureCitation id="B864E0D4CDB037BECAAADAA56BD8742F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A head ventrally B, C antenna dorsally A, B Gorhamium bidentatum sp. nov., male C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure4" httpUri="https://binary.pensoft.net/fig/1063778">4 B, C</figureCitation>
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); e) aedeagus with each paramere projected ventrobasally into a slender, medially curved process (d
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<subScript id="CC5686A6F2ACE72FA41CD13D5C48FF22">1</subScript>
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, Fig.
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<figureCitation id="403715F458E4507898C2797AE57DCFE3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 C</figureCitation>
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), sometimes joining at midline forming an annular bridge (d
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<subScript id="6580E751C495E3C5B1F911CD8D184385">3</subScript>
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, Fig.
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<figureCitation id="4130430A715A071F6B7FC6A3B25ADB02" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 G</figureCitation>
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) (also present in
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<taxonomicName id="605A4C52F1CC387A5F3E155C6B64F03D" authorityName="Gorham" authorityYear="1881" class="Insecta" family="Lycidae" genus="Calocladon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
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<emphasis id="F3FB35D27EF0F50CDA97FCBF98067A97" italics="true">Calocladon</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="615BB1C1C701B92E17C238F65FA1ADFB" authorityName="Nascimento & Bocakova" authorityYear="2022" class="Insecta" family="Lycidae" genus="Cladocalon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
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<emphasis id="EB5515CDA989AA948836376E7DEF209E" italics="true">Cladocalon</emphasis>
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</taxonomicName>
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, and
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<emphasis id="A1A7E844AE31CD7460BA100AE65DF2A8" italics="true">Atlanticolycus</emphasis>
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). Among unique features of
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<taxonomicName id="0E6DDDC53CBDA32AAA9155ED7FA52105" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="65C55B474DBCA1A08C8B4A91571E3670" italics="true">Gorhamium</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BD4D5C022109A96C44D32941E8185387" rank="genus">gen. nov.</taxonomicNameLabel>
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belong: a) base of phallus pointed anchored-shaped (inverted mushroom-shaped), with arcuate arms and a pointed tip (a
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<subScript id="FDA1B11D6F671C848FB1EC1BA5BC04E8">1</subScript>
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, Fig.
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<figureCitation id="C6389A2E1C30B7C3834FC86AFC84854D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 E, F</figureCitation>
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), while the base of phallus of
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<taxonomicName id="8B11A925D9D32D7C6586620A3B0ED39B" authorityName="Nascimento & Bocakova" authorityYear="2022" class="Insecta" family="Lycidae" genus="Cladocalon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
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<emphasis id="BA552870ADC5032A15746B498A4F184E" italics="true">Cladocalon</emphasis>
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</taxonomicName>
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and
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<emphasis id="47A56918EE6007824FAA5DE7DEF674F9" italics="true">Atlanticolycus</emphasis>
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is flat, or rounded (a
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<subScript id="C70601E90948FF79E28B4FC218B0B701">2</subScript>
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, Fig.
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<figureCitation id="ED16641537042D0B3B1DCA780AED89D9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 A</figureCitation>
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); b) median portion of phallus extending ventrally into oval opening (b, Fig.
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<figureCitation id="DAC18F6834F36FE2E5BFB4D643EA7466" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 C</figureCitation>
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); c) dorsal edge of phallus hooked (c, Fig.
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<figureCitation id="8AD5C575FEA1A61A7A065800645CAB9B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 D, G</figureCitation>
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); d) internal sac membranous with minute spines distally (e, Fig.
|
||
<figureCitation id="3ED8CFAA822460F72EAFB0748D53CCCE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 D, G</figureCitation>
|
||
); e) parameres shorter than 2 / 3 of phallus (while the parameres are almost as long as phallus in
|
||
<emphasis id="937A55AB88A209DBD3D8CC9291CBDB95" italics="true">Atlanticolycus</emphasis>
|
||
); f) base of parameres semicircular in cross-section (flattened / ribbon-like in
|
||
<taxonomicName id="8F325EC90EC5BA2179F61D273387CCCD" authorityName="Nascimento & Bocakova" authorityYear="2022" class="Insecta" family="Lycidae" genus="Cladocalon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02DDAD691414CEE787294E76678C3086" italics="true">Cladocalon</emphasis>
|
||
</taxonomicName>
|
||
); g) apex of parameres denticulate, provided with one or two coarse teeth; h) female genitalia with valvifers as long as coxites and styli combined (Fig.
|
||
<figureCitation id="BCD6CE0C6A06F54957525D86B8998E7C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="2D568ED7F4DD149CCBEB3855CFB004C4" ID-DOI="10.3897/zookeys.1204.114932.figure2" ID-arpha="594F0428-E84B-59EE-869B-3964BDEB36DE" httpUri="https://binary.pensoft.net/fig/1063776" startId="F2">
|
||
<paragraph id="42E8DFB50301EA62C2BA0A80559023DD">
|
||
<label id="8A2138DCA4963701CFA797480CF6E7AE">Figure 2.</label>
|
||
</paragraph>
|
||
<paragraph id="FF0968D5F51E882CFF6CC58B5D4D6097">
|
||
Habitus, dorsal view
|
||
<emphasis id="A17D06735CA5CA8914218EB7C0F30AC2" bold="true">A, B</emphasis>
|
||
<taxonomicName id="04AD9065F473ED822A9D437717D832E9" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bidentatum" status="sp. nov.">
|
||
<emphasis id="0C1D6366FEAD467CD49C562FBC61222D" italics="true">Gorhamium bidentatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9C8262A106DDB18EE57E279FE7F2CA6B" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="A5E21776F2DF515863CA4558A865F463" bold="true">A</emphasis>
|
||
male
|
||
<emphasis id="47817D26552A9AA5A37F48824313830B" bold="true">B</emphasis>
|
||
female
|
||
<emphasis id="244ECEBF79F44D05520B007ED4AE024D" bold="true">C</emphasis>
|
||
<taxonomicName id="70271428D9082F9B68F2530DE4363819" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="unidentatum" status="sp. nov.">
|
||
<emphasis id="C86A3622A94D885091578D42EFF43C7D" italics="true">Gorhamium unidentatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="135CDE10245B869E1CBC4435B344256A" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="28E31BBA3FA57C805F7222F1CF8DB1E1" ID-DOI="10.3897/zookeys.1204.114932.figure3" ID-arpha="FEA3FAD5-3CD2-59BA-8369-04D6240FDE97" httpUri="https://binary.pensoft.net/fig/1063777" startId="F3">
|
||
<paragraph id="FBC29D57A48C286FDA3E6E403E8F6FAC">
|
||
<label id="842C0644F6E8E9BE34AA31D83ECCACBD">Figure 3.</label>
|
||
</paragraph>
|
||
<paragraph id="7015251C59C6D5A1AB49E345F1D080CC">
|
||
Pronotum
|
||
<emphasis id="D2AE1A73C5EBC88DC370F478732F0BAA" bold="true">A</emphasis>
|
||
<taxonomicName id="DD5B24745DAF486A1BCD8DABD1869320" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bidentatum" status="sp. nov.">
|
||
<emphasis id="FD240C02918544CFA812C12A158DCF31" italics="true">Gorhamium bidentatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CF638F89A09372D74243D056CB2E3464" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male
|
||
<emphasis id="83D5340F4F8A2993313D6663DE765373" bold="true">B</emphasis>
|
||
<taxonomicName id="3528050C7810CC0AAD210257C57862C0" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="unidentatum" status="sp. nov.">
|
||
<emphasis id="1551320D28D4CABECB8BC2E97DE3F7C2" italics="true">Gorhamium unidentatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="582F7544A8AB5AAC874918A689B10715" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C4D64B768F0E5DBFD2B638B65A14B053" ID-DOI="10.3897/zookeys.1204.114932.figure4" ID-arpha="FDE2E42B-686E-576A-8EA9-B5C43E722699" httpUri="https://binary.pensoft.net/fig/1063778" startId="F4">
|
||
<paragraph id="20EEE2488346367AD06364A6C3A6B9B9">
|
||
<label id="F5ACF88F7AA48E00F26678ECF289CA8B">Figure 4.</label>
|
||
</paragraph>
|
||
<paragraph id="56BC7E69402DF7A33C8F58B8FA90FE06">
|
||
<emphasis id="DAF3CD64399F0BC93D4FC5339A1EF15B" bold="true">A</emphasis>
|
||
head ventrally
|
||
<emphasis id="D01A71141086888B97583FC203AE9A1D" bold="true">B, C</emphasis>
|
||
antenna dorsally
|
||
<emphasis id="06B38B751F6A3E5F96CC6DC4DF1E4916" bold="true">A, B</emphasis>
|
||
<taxonomicName id="486E67A4530B173BE84961090B738A81" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bidentatum" status="sp. nov.">
|
||
<emphasis id="96DDB8B00E365F419899ABB3B5B6399A" italics="true">Gorhamium bidentatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8FD1F3264918D91236F590FBE7F0DA44" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male
|
||
<emphasis id="9EF50CE5089929E47E1BE0EDCAABB5A6" bold="true">C</emphasis>
|
||
<taxonomicName id="18798F833C1B64AD05ECA8DB62CDF0A7" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="unidentatum" status="sp. nov.">
|
||
<emphasis id="D723B2E677FE596BA1A8E2BCA7C972E3" italics="true">Gorhamium unidentatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="976533B69D6368AC6E84F8139A779F65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EQTAG" type="description">
|
||
<paragraph id="47CA725DC9AFB4DC8605029D72DDB246">
|
||
<heading id="90522D1A317ED79F052FB6756F2E71AF" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="85EC25890C4B0BFA9BDD42A873BB627F">
|
||
Body length:
|
||
<quantity id="D850592B82DB6B145BBE8106924B4813" metricMagnitude="-3" metricUnit="m" metricValue="5.95" metricValueMax="6.4" metricValueMin="5.5" unit="mm" value="5.95" valueMax="6.4" valueMin="5.5">5.5–6.4 mm</quantity>
|
||
, width across the humeri:
|
||
<quantity id="E4EB2E94191C446A4237D0BEB1FA344E" metricMagnitude="-3" metricUnit="m" metricValue="1.2" unit="mm" value="1.2">1.2 mm</quantity>
|
||
. Head partly covered by pronotum from above. Labrum small, mandibles slender, arcuate (Fig.
|
||
<figureCitation id="7DEADA2143896B105C03525C0139D2CC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A head ventrally B, C antenna dorsally A, B Gorhamium bidentatum sp. nov., male C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure4" httpUri="https://binary.pensoft.net/fig/1063778">4 A</figureCitation>
|
||
). Maxillary palps 4 - segmented, gradually widened distally, palpomere 1 (= P 1) at least 3 × shorter than P 2, P 2 longest of all, ~ 2 × longer than P 4, P 3 1.5 × shorter than P 4, terminal palpomere securiform, apex obliquely rounded (Fig.
|
||
<figureCitation id="9DB7ABFF8D073C44713F502E425E8CFE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A head ventrally B, C antenna dorsally A, B Gorhamium bidentatum sp. nov., male C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure4" httpUri="https://binary.pensoft.net/fig/1063778">4 A</figureCitation>
|
||
). Terminal palpomere of labial palps securiform. Pronotum somewhat trapezoidal, with anterior margin produced forward, posterior margin 1.4 × wider than median length; lateral margins divergent posterad, with anterior 2 / 3 almost straight, convergent anteriorly, posterior angles acute; posterior margin bisinuate, medioposterior process almost triangular (Fig.
|
||
<figureCitation id="5FCD2E81A96242D2C9218AB671EFC049" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pronotum A Gorhamium bidentatum sp. nov., male B Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure3" httpUri="https://binary.pensoft.net/fig/1063777">3 A, B</figureCitation>
|
||
); median longitudinal carina on pronotum bifurcating in anterior third, forming very slender, lenticular areola. Scutellum square, apex minutely emarginate medially (Fig.
|
||
<figureCitation id="BBE10B1C951BF7B47DDB7DB7C12E18F9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pronotum A Gorhamium bidentatum sp. nov., male B Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure3" httpUri="https://binary.pensoft.net/fig/1063777">3 A</figureCitation>
|
||
). Elytra subparallel-sided, slender, 4 × longer than humeral width (Fig.
|
||
<figureCitation id="899018E91892FE5EFE882932F0D63C16" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal view A, B Gorhamium bidentatum sp. nov. A male B female C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure2" httpUri="https://binary.pensoft.net/fig/1063776">2 A – C</figureCitation>
|
||
). Each elytron with nine longitudinal costae (4 primary costae and 5 less elevated secondary costae), primary costae 2 and 4 strongly elevated; intercostal intervals with a row of irregular reticulate cells, secondary costae 3 and 4 absent posteriorly. Anterior thoracic spiracles small, tubulate. Legs compressed, trochanters almost triangular (Fig.
|
||
<figureCitation id="DCBD291CBF72588B891DE7BFFAA2399A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A head ventrally B, C antenna dorsally A, B Gorhamium bidentatum sp. nov., male C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure4" httpUri="https://binary.pensoft.net/fig/1063778">4 A</figureCitation>
|
||
), as long as third of femur, tibiae straight, their spurs small, covered by pubescence, tarsomeres 1–4 lobed.
|
||
</paragraph>
|
||
<paragraph id="0D2FF2AE2E7A0C8A2982E96BFB04F67C">
|
||
<emphasis id="ABF1926B68EC8800C716E3F321F1CE81" bold="true">Male.</emphasis>
|
||
Eyes medium-sized to large, eye diameter 1.3–1.7 × longer than interocular distance. Antennae reaching beyond elytral midlength, antennomeres 3–10 flabellate, antennal branches flattened, antennomere 1 (= A 1) stout, A 2 small, transverse, A 3 slightly (1.15–1.3 ×) shorter than A 4, A 4 – A 10 subequal in length. Lamellae arise basally, lamella of A 3 slightly longer than antennomere body, remaining lamellae considerably longer. Abdominal sternum VIII widely emarginated distally (Fig.
|
||
<figureCitation id="9CCEE22AE10116E48584FDC8B0682923" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Terminal abdominal segments A – D Gorhamium bidentatum sp. nov. E – H Gorhamium unidentatum sp. nov., male G Cladocalon chiriquense (Gorham, 1884) A, B, E, G male terminalia (sternum IX and tergum IX – X), A – ventral view; B, E, G – dorsal view C Male sternum and tergum VIII, ventral view D female terminal sternum, ventral view. F, Male sternum VIII, ventral view H Male tergum VIII, dorsal view." figureDoi="10.3897/zookeys.1204.114932.figure5" httpUri="https://binary.pensoft.net/fig/1063779">5 C, F</figureCitation>
|
||
), emargination shallow, as deep as ¼ of sternum length. Sternum IX elongate, 3.5 × longer than wide (Fig.
|
||
<figureCitation id="6FCF5E152E487FFC0144C79901CBD07D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Terminal abdominal segments A – D Gorhamium bidentatum sp. nov. E – H Gorhamium unidentatum sp. nov., male G Cladocalon chiriquense (Gorham, 1884) A, B, E, G male terminalia (sternum IX and tergum IX – X), A – ventral view; B, E, G – dorsal view C Male sternum and tergum VIII, ventral view D female terminal sternum, ventral view. F, Male sternum VIII, ventral view H Male tergum VIII, dorsal view." figureDoi="10.3897/zookeys.1204.114932.figure5" httpUri="https://binary.pensoft.net/fig/1063779">5 A, E</figureCitation>
|
||
), widest in distal quarter, proximal half narrow with lateral margins convergent. Phallus with ventromedial oval opening (b, Fig.
|
||
<figureCitation id="9F68A631A54D38BB6519F323EF461483" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 C, G</figureCitation>
|
||
), base of phallus pointed anchored-shaped, or inverted mushroom-shaped (a
|
||
<subScript id="BE7A622ECADB1376200CAB691AFC100E">1</subScript>
|
||
, Fig.
|
||
<figureCitation id="FF4D95D3682BD16B8ED494F836AC4931" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 E, F</figureCitation>
|
||
); distal portion of phallus rod-like, apex clavate, dorsal margin hooked (c, Fig.
|
||
<figureCitation id="829CD5E6792AE6640C1F1270277E4F38" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 D, G</figureCitation>
|
||
); internal sac membranous with minute spines distally (e, Fig.
|
||
<figureCitation id="BC49B362C13597B969051533F8F59093" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 G</figureCitation>
|
||
), sometimes also medially. Parameres at most as long as 2 / 3 of phallus, base of parameres almost semicircular in cross-section; each paramere projected basally in a thin ventral, medially arched, process (d
|
||
<subScript id="E1EACF0496D65EF77C4377FD6EF8D463">1</subScript>
|
||
, Fig.
|
||
<figureCitation id="123F93B13952F647DBE33237CDD34F3D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 C</figureCitation>
|
||
), sometimes joining medially in a ring-like bridge (d
|
||
<subScript id="FF345B8B15E1C317F70E822620DDAAFC">3</subScript>
|
||
, Fig.
|
||
<figureCitation id="9395A9FA36B27BCF221DF5AC135538DE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 G</figureCitation>
|
||
); parameral apex denticulate, provided with one or two coarse teeth. Phallobase slightly asymmetrical, distorted, moderately arched ventrally.
|
||
</paragraph>
|
||
<caption id="9112F10374B19134816558DAA494C1A6" ID-DOI="10.3897/zookeys.1204.114932.figure5" ID-arpha="68B8C686-662B-5B58-BB3A-26C6E1D81D06" httpUri="https://binary.pensoft.net/fig/1063779" startId="F5">
|
||
<paragraph id="18C7F91699B865EED3D2AB868B605517">
|
||
<label id="C927C25EF44C91846D15CEF9DAE533BA">Figure 5.</label>
|
||
</paragraph>
|
||
<paragraph id="800F0D092D26D0EA7892EFBDB79113F8">
|
||
Terminal abdominal segments
|
||
<emphasis id="912A791CC8692AFA0EBE8C01AE92C926" bold="true">A – D</emphasis>
|
||
<taxonomicName id="F6EFF0D0D5EC5523852F5CB87A2E352B" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bidentatum" status="sp. nov.">
|
||
<emphasis id="AE390AECD9FD4996D58BA46758181AEB" italics="true">Gorhamium bidentatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="4C09D400F903A8F3653458697AF5B9A6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="FC280BE4F77EAD08ABD297238B3791E7" bold="true">E – H</emphasis>
|
||
<taxonomicName id="F2E6DAF54F61A68E5AD49A125CCBA086" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="unidentatum" status="sp. nov.">
|
||
<emphasis id="F32C2F77EE4445ECDBE890F9B672FE76" italics="true">Gorhamium unidentatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="33E9C8F928021F9218DE36836E351088" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male
|
||
<emphasis id="D3D01AE625C56CBABBB9EA7954AFE987" bold="true">G</emphasis>
|
||
<taxonomicName id="945DBE94EAEA7875E9163A301DB06915" authority="(Gorham, 1884)" baseAuthorityName="Gorham" baseAuthorityYear="1884" class="Insecta" family="Lycidae" genus="Cladocalon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="chiriquense">
|
||
<emphasis id="D74EAA992818BE7AA45348EF153A1404" italics="true">Cladocalon chiriquense</emphasis>
|
||
(
|
||
<bibRefCitation id="C26BB7042F2DC6F42A93A7EA893736C7" author="Gorham HS" journalOrPublisher="Coleoptera" pagination="225-272" refId="B12" refString="Gorham HS (1884) Malacodermata. Suplement. Fam. Lycidae. Biologia Centrali-Americana. Coleoptera 3: 225 - 272." title="Malacodermata. Suplement. Fam. Lycidae. Biologia Centrali-Americana." volume="3" year="1884">Gorham, 1884</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="DEBBAE3821F0EED3CEC770F6D130B40D" bold="true">A, B, E, G</emphasis>
|
||
male terminalia (sternum IX and tergum IX – X),
|
||
<emphasis id="25EA5C1127C1B498521477F92BE000C0" bold="true">A</emphasis>
|
||
– ventral view;
|
||
<emphasis id="6546AE30DCD295689DA5FDB6E57CB873" bold="true">B, E, G</emphasis>
|
||
– dorsal view
|
||
<emphasis id="C70514D203C2D724914B41E79E0CB6CD" bold="true">C</emphasis>
|
||
Male sternum and tergum VIII, ventral view
|
||
<emphasis id="FA859D13DEEA5DC16DE39571424AFCA3" bold="true">D</emphasis>
|
||
female terminal sternum, ventral view.
|
||
<emphasis id="FE615E386915DF0AF4F36CA5D37E8FD2" bold="true">F</emphasis>
|
||
, Male sternum VIII, ventral view
|
||
<emphasis id="560878409B25AE8BBDA4ABF5FCFDD5D6" bold="true">H</emphasis>
|
||
Male tergum VIII, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F3DA8F026C685EA19DD0ECCC7756905B" ID-DOI="10.3897/zookeys.1204.114932.figure6" ID-arpha="69AA6233-2436-565D-9B9A-7605C94A21C6" httpUri="https://binary.pensoft.net/fig/1063780" startId="F6">
|
||
<paragraph id="D5C6B0103673E0F6CED826C7F899042A">
|
||
<label id="B9C7CF55282E7A2732896DBEE35A7A0B">Figure 6.</label>
|
||
</paragraph>
|
||
<paragraph id="A9355439E747429E5A26741689672CDA">
|
||
<emphasis id="17C80186F8940B24DC2BC46D7743C486" bold="true">A – G</emphasis>
|
||
, Male genitalia
|
||
<emphasis id="0F1C3B3A53B2F4317F85B15C7477BB02" bold="true">A, B</emphasis>
|
||
<taxonomicName id="32C547111976EAB72765C0D1280FF15F" authority="(Gorham, 1884)" baseAuthorityName="Gorham" baseAuthorityYear="1884" class="Insecta" family="Lycidae" genus="Cladocalon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="chiriquense">
|
||
<emphasis id="4C1A2828D60EF5AD6EA6AF9DF906EA40" italics="true">Cladocalon chiriquense</emphasis>
|
||
(
|
||
<bibRefCitation id="45D18022FF0C0E1E0C53543CD8D72C2A" author="Gorham HS" journalOrPublisher="Coleoptera" pagination="225-272" refId="B12" refString="Gorham HS (1884) Malacodermata. Suplement. Fam. Lycidae. Biologia Centrali-Americana. Coleoptera 3: 225 - 272." title="Malacodermata. Suplement. Fam. Lycidae. Biologia Centrali-Americana." volume="3" year="1884">Gorham, 1884</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="FD35C8AC08504A308CEDA0D2CFEABF43" bold="true">C – E</emphasis>
|
||
<taxonomicName id="D86AD25C7E024112B082FA468E54288C" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bidentatum" status="sp. nov.">
|
||
<emphasis id="D8AB60373965D33C2CEFD207E3EA827F" italics="true">Gorhamium bidentatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="4A9EE3C573BF25B8B89AD06DFDD231EC" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="585DD5273C9EC35A7E3B1A01098BEEA5" bold="true">F – H</emphasis>
|
||
<taxonomicName id="C0B9DE8E6A3E7A2594216A47288E1C83" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="unidentatum" status="sp. nov.">
|
||
<emphasis id="E38E4CBA7AA69DD859D0AC61C285E70A" italics="true">Gorhamium unidentatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="415935123A42448055736CDDF9D701D6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="F230E3A3832FBE02D5F10C5B89D9F15F" bold="true">H</emphasis>
|
||
female genitalia of
|
||
<taxonomicName id="8333188137CA4B5C46D33007228E855F" authority="Nascimento & Bocakova, 2024" authorityName="Nascimento & Bocakova" authorityYear="2024" class="Insecta" family="Lycidae" genus="Gorhamium" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="bidentatum" status="sp. nov.">
|
||
<emphasis id="031C86646FCCBD271E6798138275FA9B" italics="true">Gorhamium bidentatum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E4B961B7A2450CB82C76784836494B8C" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ventral view.
|
||
<emphasis id="75BF9E8662571E8A5F47959D8EB7B48E" bold="true">A, C, F</emphasis>
|
||
ventral view
|
||
<emphasis id="9CF1BC7246F495DB8E7D3B21D3CF3C72" bold="true">B, D, G</emphasis>
|
||
lateral view
|
||
<emphasis id="65A857E679071B3F30F282C1C2A3487A" bold="true">E</emphasis>
|
||
ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="197FADA728A1D204B467F78F4FDE7758">
|
||
<emphasis id="9458F1F445C3A448864C7535F4D9C395" bold="true">Female.</emphasis>
|
||
Eyes small, interocular distance 1.3 × longer than eye diameter, antennae serrate (Fig.
|
||
<figureCitation id="10F957B473E59638188F03D6DFD3F2A9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus, dorsal view A, B Gorhamium bidentatum sp. nov. A male B female C Gorhamium unidentatum sp. nov., male." figureDoi="10.3897/zookeys.1204.114932.figure2" httpUri="https://binary.pensoft.net/fig/1063776">2 B</figureCitation>
|
||
). Terminal sternum (
|
||
<collectionCode id="27EF9D11A686B59D2A0D65749450B9E3">IX</collectionCode>
|
||
) simple (Fig.
|
||
<figureCitation id="A78BFF6839EE542931F24CAE3434BB72" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Terminal abdominal segments A – D Gorhamium bidentatum sp. nov. E – H Gorhamium unidentatum sp. nov., male G Cladocalon chiriquense (Gorham, 1884) A, B, E, G male terminalia (sternum IX and tergum IX – X), A – ventral view; B, E, G – dorsal view C Male sternum and tergum VIII, ventral view D female terminal sternum, ventral view. F, Male sternum VIII, ventral view H Male tergum VIII, dorsal view." figureDoi="10.3897/zookeys.1204.114932.figure5" httpUri="https://binary.pensoft.net/fig/1063779">5 D</figureCitation>
|
||
), spiculum gastrale rudimentary, triangular. Ovipositor with valvifers 1.3 × longer than coxites (Fig.
|
||
<figureCitation id="F7E1B9A39372349A2C224570087C787D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. A – G, Male genitalia A, B Cladocalon chiriquense (Gorham, 1884) C – E Gorhamium bidentatum sp. nov. F – H Gorhamium unidentatum sp. nov. H female genitalia of Gorhamium bidentatum sp. nov., ventral view. A, C, F ventral view B, D, G lateral view E ventrolateral view. Abbreviations: a 1 – pointed anchor-shaped base of phallus, a 2 – flat anchor-shaped base of phallus, b – phallic ventral opening, c – dorsal dent, d 1 – arcuate ventrobasal parameral process, d 2 – flattened ventrobasal parameral process, d 3 – a ring-like ventral bridge (ventrobasal processes medially fused), e – internal sac." figureDoi="10.3897/zookeys.1204.114932.figure6" httpUri="https://binary.pensoft.net/fig/1063780">6 H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ET1AG" type="etymology">
|
||
<paragraph id="58E8447D0EBC1982DF7E2ECBCF46800A">
|
||
<heading id="57F5C419414208C3321E530EC3670099" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="F79F5E49A758A5AFEC63B749BB257297">
|
||
The genus is named in honor of H. S. Gorham, the author of chapters on Malacodermata in Biologia-Centrali Americana (
|
||
<bibRefCitation id="2BCC48141C195E1F278B96E3850E516D" author="Gorham" firstAuthor="Gorham" journalOrPublisher="Coleoptera" pagination="1-24" refId="B10" refString="Gorham HS (1880) Malacodermata. Fam. Lycidae. Biologia Centrali-Americana. Coleoptera 3: 1–24." title="Malacodermata. Fam. Lycidae. Biologia Centrali-Americana." volume="3" year="1880">Gorham 1880</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="7ACB10029380981B8A5D46FA3D63ECF5" author="Gorham" firstAuthor="Gorham" journalOrPublisher="Coleoptera" pagination="25-29" refId="B11" refString="Gorham HS (1881) Malacodermata. Fam. Lycidae. Biologia Centrali-Americana. Coleoptera 3: 25–29." title="Malacodermata. Fam. Lycidae. Biologia Centrali-Americana." volume="3" year="1881">1881</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="8ED8761E4BC85D1ADD8C94AA0ED09E62" author="Gorham" firstAuthor="Gorham" journalOrPublisher="Coleoptera" pagination="225-272" refId="B12" refString="Gorham HS (1884) Malacodermata. Suplement. Fam. Lycidae. Biologia Centrali-Americana. Coleoptera 3: 225–272." title="Malacodermata. Suplement. Fam. Lycidae. Biologia Centrali-Americana." volume="3" year="1884">1884</bibRefCitation>
|
||
), where he described many genera and species of
|
||
<taxonomicName id="F8A80FAFE3A933A50EF376F90FD42C0E" class="Insecta" family="Lycidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="tribe" tribe="Eurrhacini">Eurrhacini</taxonomicName>
|
||
and
|
||
<taxonomicName id="6E4C35E3EDD82D3A1754BC6D997BAEDF" class="Insecta" family="Lycidae" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="tribe" tribe="Calopterini">Calopterini</taxonomicName>
|
||
. The gender is neuter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EP2AG" type="distribution">
|
||
<paragraph id="7B3C8BC832D98DA11B3DE751355C6337">
|
||
<heading id="C5AD44B0453354C37E50429CCA255C28" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="759152BDF99B765D7634ACCEEBE6882D">
|
||
<collectingCountry id="76DA3EB6E053209A48E45039135E8D2B" name="Panama">Panama</collectingCountry>
|
||
,
|
||
<collectingCountry id="4FCB005C5D11633C94CC7B4FC5130078" name="Ecuador">Ecuador</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |